Systematics of the snakes
The system of snakes presented here lists all taxa of snakes (Serpentes) down to the level of the genus . The systematics of snakes is still the subject of research and changes frequently according to more recent findings. Here the system is reproduced according to the Reptile Database , which also includes recent genetic studies. The following cladogram shows a possible relationship between superfamilies and families within the snakes:
Phylogenetic systematics of snakes according to Pyron et al. 2013 | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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As of July 1, 2018, the Reptile Database classifies a total of 3709 individual species into the following taxa .
Superfamily Acrochordoidea
Acrochordidae family
- Wart snakes ( Acrochordus )
Superfamily Boaiformes (Booidea or Henophidia)
Boas family (Boidae)
Subfamily Boaschlangen (Boinae)
- Boas ( boa )
- Chilabothrus
- Dog's head boas ( Corallus )
- Slim boas ( Epicrates )
- Anacondas ( Eunectes )
Subfamily earth pythons (Calabariinae)
- Earth python ( Calabaria )
Subfamily Candoiinae
- Pacific boas ( Candoia )
Subfamily dwarf boas (Charininae)
- Charina
- Exiliboa
- Lichanura
- Banana Boas ( Ungaliophis )
Subfamily Erycinae
- Sand boas ( Eryx )
Subfamily Madagascar boas ( Sanziniinae )
- Madagascar boas ( Acrantophis )
- Sanzinia
Superfamily adder-like and viper-like (Colubroidea)
Adder family (Colubridae)
Subfamily dwarf snakes (Calamariinae)
- Real dwarf snakes ( Calamaria )
- Calamorhabdium
- Collorhabdium
- Etheridgeum
- Mountain dwarf snakes ( Macrocalamus )
- Pseudorabdion
- Rabdion
Subfamily real snakes (Colubrinae)
- Aeluroglena
- Whip Snakes ( Ahaetulla )
- Aprosdoketophis
- Archelaphs
- Argyrogena
- Arizona snakes ( Arizona )
- Bamanophis
- North American rat snakes ( Bogertophis )
- Night tree snakes ( Boiga )
- Cemophora
- Chapinophis
- Sand snakes ( Chilomeniscus )
- Shovel nose snakes ( Chionactis )
- Chironius
- Ornamental tree snakes ( Chrysopelea )
- Coelognathus
- Angry Snakes ( Coluber )
- Colubroelaps
- Conopsis
- Smooth snakes ( Coronella )
- Crotaphopeltis
- Cyclophiops
- African egg snakes ( Dasypeltis )
- Bronze snakes ( Dendrelaphis )
- Wood snakes ( dendrophidion )
- Dipsadoboa
- Dispholidus
- Dolichophis
- Drymarchon
- Racing snakes ( Drymobius )
- Drymoluber
- Dryocalamus
- Dryophiops
- Eirenis
- Elaphe
- Euprepiophis
- Hooknose adders ( Ficimia )
- Geagras
- False climbing snakes ( Gonyosoma )
- Mexican hook- nosed snakes ( Gyalopion )
- Hapsidophrys
- Hemerophis
- Hemorrhois
- Hierophis
- King snakes ( Lampropeltis )
- Leptodrymus
- Thin snakes ( Leptophis )
- Lepturophis
- Liopeltis
- Wolf tooth snakes ( Lycodon )
- Snout snakes ( Lytorhynchus )
- Macroprotodon
- Masticophis
- Mastigodryas
- Meizodon
- Mopanveldophis
- Muhtarophis
- Kukrin Snakes ( Oligodon )
- Oocatochus
- Grass snakes ( Opheodrys )
- Oreocryptophis
- Orientocoluber
- Pointed Snakes ( Oxybelis )
- American climbing snakes ( Pantherophis )
- Philothamnus
- Phrynonax
- Phyllorhynchus
- Gopher, pine and bull snakes ( Pituophis )
- Platyceps
- Pliocercus
- Pseudelaphe
- Pseudoficimia
- Pseustes
- Asian racing snakes ( Ptyas )
- Rhamnophis
- Rhinechis
- Rhinobothryum
- Rhinocheilus
- Rhynchocalamus
- Salvadora
- Scaphiophis
- Scolecophis
- Senticolis
- Simophis
- North American ground snakes ( Sonora )
- Spalerosophis
- Spilotes
- Stegonotus
- Stenorrhina
- Stichophanes
- Symphony
- Sympholis
- Black-headed Snakes ( Tantilla )
- Tantillita
- Cat Snakes ( Telescopus )
- Bird snakes ( Thelotornis )
- Black tree snakes ( Thrasops )
- Toxicodryas
- Lyra snakes ( Trimorphodon )
- Wallaceophis
- Xenelaphis
- Xyelodontophis
- Zamenis
Subfamily Dipsadinae
- Adelphicos
- Alsophis
- Amastridium
- Amnesteophis
- Apostolepis
- Arrhyton
- Atractus
- Boiruna
- Borikophis
- Caaeteboia
- Calamodontophis
- Caraiba
- Worm snakes ( Carphophis )
- Cercophis
- Chersodromus
- Mussuranas ( Clelia )
- Coniophanes
- Conophis
- Contia
- Coronelapse
- Crisantophis
- Cryophis
- Cubophis
- Ringhals snakes or collar adders (Diadophis)
- Diaphorolepis
- Thick-headed snakes or snail suckers ( dipsas )
- Ditaxodon
- Drepanoides
- Echinanthera
- Elapomorphus
- Emmochliophis
- Enuliophis
- Enulius
- False coral otters ( Erythrolamprus )
- Eutrachelophis
- Farancia
- Geophis
- Gomesophis
- Haitiophis
- Helicops
- Hook-nosed snakes ( heterodon )
- Hydrodynastes
- Hydromorphus
- Hydrops
- Night snakes ( Hypsiglena )
- Hypsirhynchus
- Ialtris
- Belt snakes or giant snakes ( Imantodes )
- Cat's-eye snakes ( Leptodeira )
- Lioheterophis
- Lygophis
- Magliophis
- Manolepis
- Mussurana
- Coffee snakes ( ninia )
- Nothopsis
- Omoadiphas
- Oxyrhopus
- Paraphimophis
- Phalotris
- Philodryas
- Phimophis
- Plesiodipsas
- Pseudalsophis
- Pseudoboa
- Pseudoeryx
- Pseudoleptodeira
- Pseudotomodon
- Psomophis
- Ptychophis
- Rhachidelus
- Rhadinaea
- Rhadinella
- Rhadinophanes
- Rodriguesophis
- Saphenophis
- South American snail snakes ( Sibon )
- Sibynomorphus
- Siphlophis
- Sordellina
- Synophis
- Tachymenis
- Taeniophallus
- Tantalophis
- Thamnodynastes
- Thermophis
- Tomodon
- Tretanorhinus
- Trimetopon
- Tropidodipsas
- Tropidodryas
- Uromacer
- Uromacerina
- Urotheca
- Hood adders ( Xenodon )
- Xenopholis
Subfamily Grayiinae
Subfamily water snakes (Natricinae)
- Adelophis
- Afronatrix
- Amphiesma
- Amphiesmoides
- Anoplohydrus
- Aspidura
- Atretium
- Balanophis
- Clonophis
- Haldea
- Hebius
- Helophis
- Herpetoreas
- Hydrablabes
- Hydraethiops
- Iguanognathus
- Isanophis
- Limnophis
- Liodytes
- Lycognathophis
- Macropisthodon
- Natriciteres
- European water snakes ( Natrix )
- American swimming snakes ( Nerodia )
- Opisthotropis
- Parahelicops
- Pararhabdophis
- Paratapinophis
- Crab Snakes ( Regina )
- Rhabdophis
- Rhabdops
- Sinonatrix
- Storeria
- Garter Snakes ( Thamnophis )
- Trachishium
- Tropidoclonion
- Australian quills ( Tropidonophis )
- Virginia
- Xenochrophis
Subfamily Pseudoxenodontinae
Subfamily Sibynophiinae
Currently not assigned to any subfamily
Snake family (Elapidae)
Subfamily Elapinae
- Mock cobras ( Aspidelaps )
- Kraits ( Bungarus )
- Calliophis
- Mambas ( Dendroaspis )
- African garter otters ( Elapsoidea )
- Hemachatus
- Hemibungarus
- Micruroides
- Coral otters ( Micrurus )
- Naja ( Well )
- Ophiophagus
- Forest Cobras ( Pseudohaje )
- Sinomicrurus
- Desert cobras ( Walterinnesia )
Terrestrial poisonous snakes of Australasia are usually assigned to the subfamily of the Hydrophiinae.
Subfamily sea snakes (Hydrophiinae)
Terrestrial Hydrophiinae of Australasia:
- Death adder ( Acanthophis )
- Antaioserpens
- Aspidomorphus
- Australian copper heads ( Austrelaps )
- Brachyurophis
- Cacophis
- Cryptophis
- Demansia
- Denisonia
- Drysdalia
- Echiopsis
- Elapognathus
- Furina
- Hemiaspis
- Hoplocephalus
- Loverid gelaps
- Micropechis
- Neelaps
- Tiger otters ( Notechis )
- Ogmodon
- Taipane ( Oxyuranus )
- Parapistocalamus
- Parasuta
- Paroplocephalus
- Black otters ( pseudechis )
- Brown snakes ( pseudonaja )
- Rhinoplocephalus
- Solomon rape
- Simoselaps
- Suta
- New Guinea wood adder ( Toxicocalamus )
- Tropidechis
- Vermicella
Subfamily Laticaudinae
Family snakes (Homalopsidae)
- Bitia
- Brachyorrhosis
- Calamophis
- Cantoria
- Cerberus
- Dieurostus
- Djokoiskandarus
- Enhydris
- Erpeton
- Ferania
- Fordonia
- Gerarda
- Gyiophis
- Heurnia
- Homalopsis
- Hypsiscopus
- Carnsophis
- Kualatahan
- Mintonophis
- Miralia
- Myron
- Myrrophis
- Phytolopsis
- Pseudoferania
- Raclitia
- Subsessor
- Sumatranus
Family Lamprophiidae
Subfamily Aparallactinae
- Amblyodipsas
- Aparallactus
- Brachyophis
- Chilorhinophis
- Hypoptophis
- Macrelaps
- Micrelaps
- Polemon
- Xenocalamus
Subfamily Erdvipern (Atractaspidinae)
Subfamily Lamprophiinae
- Boaedon
- Bothrolycus
- Bothrophthalmus
- Chamaelycus
- Dendrolycus
- Gonionotophis
- Hormonal activity
- Inyoka
- House snakes ( Lamprophis )
- Lycodonomorphus
- Wolf snakes ( Lycophidion )
- Pseudoboodon
Subfamily Psammophiinae
- Dwarf-beaked snake ( Dipsina )
- Bark snakes ( Hemirhagerrhis )
- Lizard snakes ( Malpolon )
- Mimophis
- Sand snakes ( Psammophis )
- Sheep stick ( psammophylax )
- Moila snake ( Rhagerhis )
- Beak- nosed snakes ( Rhamphiophis )
Subfamily Prosymninae
Subfamily Pseudaspidinae
- Mole snake ( pseudaspis )
- Western keel snake ( Pythonodipsas )
Subfamily Pseudoxyrhophiinae
- Alluaudina
- Amplorhinus
- Brygophis
- Compsophis
- Ditypophis
- Dromicodryas
- African snail eater ( Duberria )
- Elapotinus
- Heteroliodone
- Ithycyphus
- Leaf-nosed snakes ( Langaha )
- Malagasy hook- nosed snakes ( Leioheterodon )
- Liophidium
- Liopholidophis
- Lycodryas
- Madagascarophis
- Micropisthodon
- Pararhadinaea
- Parastenophis
- Phisalixella
- Pseudoxyrhopus
- Thamnosophis
Subfamily Cyclocorinae
Currently not assigned to any subfamily
Family Pareidae
- Subfamily Pareidae
- Subfamily Xylophiinae
Viper family (Viperidae)
Subfamily Azemiopinae
- Fea vipers ( Azemiops )
Subfamily pit vipers (Crotalinae)
- Triangular-headed adder ( Agkistrodon )
- Leaping Lance Vipers ( Atropoides )
- Palm lance vipers ( bothriechis )
- Wood lance vipers ( Bothriopsis )
- Bothrocophias
- American lance vipers ( Bothrops )
- Malay moccasin otters ( Calloselasma )
- Mountain pit vipers ( Cerrophidion )
- Rattlesnakes ( Crotalus )
- Chinese nosed viper ( Deinagkistrodon )
- Garthius
- Gloydius
- Ceylon nasal vipers ( Hypnale )
- Bushmaster ( Lachesis )
- Mixcoatlus
- Mexican Horned Lance Vipers ( Ophryacus )
- Ovophis
- Hook-nosed lance-viper ( porthidium )
- Protobothrops
- Dwarf rattlesnakes ( Sistrurus )
- Bamboo otters ( Trimeresurus )
- Temple otters ( Tropidolaemus )
Subfamily Real Vipers (Viperinae)
- Bush vipers ( Atheris )
- Puff adder ( bitis )
- Toad vipers ( Causus )
- African horn vipers ( Cerastes )
- Oriental Vipers ( Daboia )
- Sand rattle otters ( Echis )
- MacMahon vipers ( Eristicophis )
- Great vipers ( Macroektivena )
- East African mountain otters ( Montatheris )
- Mountain otters ( Montivipera )
- Marsh vipers ( Proatheris )
- Trughorn vipers ( Pseudocerastes )
- Real otters ( Vipera )
Mute snakes family (Xenodermatidae)
Superfamily Pythonoidea s. l.
Pointed-head pythons family (Loxocemidae)
- Pointed head python ( Loxocemus )
Python family (Pythonidae)
- Southern pythons ( Antaresia )
- Black-headed pythons ( Aspidites )
- Bothrochilus
- Water pythons ( liasis )
- Malayopython
- Diamond pythons ( Morelia )
- Actual Pythons ( Python )
- Amethyst pythons ( Simalia )
Family Xenopeltidae
Superfamily blind snakes (Scolecophidia)
American blind snakes family (Anomalepididae)
Family Gerrhopilidae
Slender blind snakes family (Leptotyphlopidae, Glauconiidae)
Subfamily Leptotyphlopinae
Subfamily Epictinae
Family blind snakes (Typhlopidae)
Subfamily Afrotyphlopinae
Subfamily Asiatyphlopinae
- Acutotyphlops
- Anilios
- Argyrophis
- Asiaticyphlops
- Cyclotyphlops
- Grypotyphlops
- Indotyphlops
- Malayotyphlops
- Ramphotyphlops
- Sundatyphlops
- Xerotyphlops
Subfamily Madatyphlopinae
Subfamily Typhlopinae
Family Xenotyphlopidae
Superfamily Uropeltoidea s. l.
Family Anomochilidae
- Burrowing snakes ( anomochilus )
Family Cylindrophiidae
- Roller snakes ( Cylindrophis )
Family tails (Uropeltidae)
Currently not part of any superfamily
Family Aniliidae
Family bolyeriidae (Bolyeriidae)
Erdboas family (Tropidophiidae)
Pine snakes family (Xenophidiidae)
literature
The system presented here is mainly based on the following publications:
- R. Alexander Pyron, Frank T. Burbrink, Guarino R. Colli, Adrian Nieto Montes de Oca, Laurie J. Vitt, Caitlin A. Kuczynski and John J. Wiens: The phylogeny of advanced snakes (Colubroidea), with discovery of a new subfamily and comparison of support methods for likelihood trees . In: Molecular Phylogenetics and Evolution . 2010, p. 329–342 , doi : 10.1016 / j.ympev.2010.11.006 .
- Michael SY Lee, Andrew F. Hugall, Robin Lawson and John D. Scanlon: Phylogeny of snakes (Serpentes): combining morphological and molecular data in likelihood, Bayesian and parsimony analyzes . In: Systematics and Biodiversity . tape 5 , 2007, p. 371-389 , doi : 10.1017 / S1477200007002290 .
- Nicolas Vidal, Anne-Sophie Delmas, Patrick David, Corinne Cruaud, Arnaud Couloux and S. Blair Hedges: The phylogeny and classification of caenophidian snakes inferred from seven nuclear protein-coding genes . In: Comptes Rendus Biologies . tape 330 , 2007, pp. 182-187 , doi : 10.1016 / j.crvi.2006.10.001 .
- Nicolas Vidal, Anne-Sophie Delmas and S. Blair Hedges: The higher-level relationships of alethinophidian snakes inferred from seven nuclear and mitochondrial genes . In: Biology of the Boas and Pythons . 2007, p. 27-33 .
Individual evidence
- ↑ Uetz, P. et al .: The Reptile Database. Retrieved March 13, 2012.
- ↑ a b The Reptile Database: Higher Taxa in Extant Reptiles - Ophidia (Serpentes) - Snakes.
- ↑ Robert Alexander Pyron, Frank T. Burbrink & John J. Wiens: A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 2013, 13:93 doi: 10.1186 / 1471-2148-13-93
- ^ The Reptile Database: Species Numbers.
See also
Web links
- The New Reptile Database: Phylogeny of snakes (english)
- Mikko's Phylogeny Archive: Serpentes (English)
- Integrated Taxonomic Information System: Serpentes (English)