Panphagia

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Panphagia
Temporal range: Middle Triassic, 231.4 Ma
Reconstruction of the fossil specimen
Scientific classification Edit this classification
Missing taxonomy template (fix): Panphagia protos

Panphagia is a genus of sauropodomorph dinosaur described in 2009. It lived 231 million years ago, during the late Ladinian age of the mid Triassic period (according to the ICS;[1] alternately the early Carnian age of the Late Triassic in the system used by the Geological Society of America), in what is now northwestern Argentina.[2] The name Panphagia comes from the Greek words pan, meaning "all", and phagein, meaning "to eat", in reference to its inferred omnivorous diet. Panphagia is one of the earliest known dinosaurs, and is an important find which may mark the transition of diet in early sauropodomorph dinosaurs.

Discovery

Silhouette reconstruction of the skeleton of Panphagia protos
Restoration

Fossils of Panphagia were found in late 2006 by the Argentinean paleontologist Ricardo N. Martínez in rocks of the Ischigualasto Formation of Valle Pintado, Ischigualasto Provincial Park, San Juan Province, Argentina. The bones were found at approximately the same level as a 231.4 million year old ash layer, indicating it lived during the early Carnian of the Late Triassic.[2] Panphagia is currently known from holotype PVSJ 874, the disarticulated remains of one partially-grown individual of about 1.30 metres (4.3 ft) long. Portions of the skull, vertebrae, pectoral girdle, pelvic girdle, and hindlimb bones have been recovered. The russet-colored fossils were embedded in a greenish sandstone matrix and took several years to prepare and describe.[3]

Classification

 Saurischia 
    
    

Eoraptor

    

Theropoda

Sauropodomorpha

Panphagia

    

Saturnalia

Other sauropodomorphs

Basal saurischian phylogeny simplified after Martinez and Alcober, 2009.[3] This is only one of many proposed cladograms for basal saurischians.

Panphagia was described in 2009 by Ricardo N. Martínez and Oscar A. Alcober, both of the Museo de Ciencias Naturales, in San Juan, Argentina. They performed a phylogenetic analysis and found it to be the most basal known sauropodomorph dinosaur: the fossils shared similar features to those of Saturnalia, an early sauropodomorph, including similarities in the ischium, astralagus, and the scapular blade. Yet the fossils also exhibited similar features to those of Eoraptor, an early carnivorous saurischian, including hollow bones, sublanceolate teeth, and overall proportions. Based on analysis and comparison of the Panphagia fossils and those of its closest kin, Martínez and Alcober concluded that the evolution of saurischian dinosaurs likely began with small, cursorial animals similar to Panphagia, and that there is a "general similarity among all of these basal dinosaurs [suggesting] that few structural changes stand" between Panphagia, Eoraptor, and two basal theropods which have yet to be described.[3]

The type species of Panphagia is P. protos; the specific name, meaning "the first" in Greek, is a reference to its basal position.[3]

Diet

Preserved skull bones of Panphagia protos (PVSJ 874)

The teeth of Panphagia indicate a possible omnivorous diet, transitional in form between the mostly carnivorous theropods and the herbivorous sauropodomorphs. The teeth in the back of the jaw are shorter than those in the front, are leaf-shaped, and also have more marked serrations.[3]

See also

References

  1. ^ Gradstein, F.M., Ogg, J.G. and Smith, A.G. (2004). A Geologic Time Scale 2004, Cambridge University Press.
  2. ^ a b Alcober, Oscar A. (2010). "A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of northwestern Argentina". ZooKeys. 63: 55–81. doi:10.3897/zookeys.63.550. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)CS1 maint: unflagged free DOI (link)
  3. ^ a b c d e Martínez, Ricardo N. (2009). "A basal sauropodomorph (Dinosauria: Saurischia) from the Ischigualasto Formation (Triassic, Carnian) and the early evolution of Sauropodomorpha" (pdf). PLoS ONE. 4 (2): 1–12. doi:10.1371/journal.pone.0004397. PMC 2635939. PMID 19209223. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)CS1 maint: unflagged free DOI (link)