Hebius: Difference between revisions

Hebius
	Japanese keelback (Hebius vibakari)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Suborder:	Serpentes
Family:	Colubridae
Subfamily:	Natricinae
Genus:	Hebius; Thompson, 1913

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Latest revision as of 19:55, 27 February 2024

Hebius is a genus of snakes in the family Colubridae.^[1]

Geographic range[edit]

The genus Hebius is endemic to Asia.^[2]

Taxonomy[edit]

All of the member species of the genus Hebius were formerly placed in the genus Amphiesma, but in 2014 Guo et al. placed most species of Amphiesma in the genus Hebius. They placed other species in the genus Herpetoreas, leaving Amphiesma a monotypic taxon containing only the species Amphiesma stolatum.^[2]

Species[edit]

The following 48 species are recognized as being valid.^[1]

Hebius andreae (Ziegler & Quyet, 2006) – Andrea’s keelback
Hebius annamensis Bourret, 1934 – Annam keelback
Hebius arquus (David & G. Vogel, 2010)
Hebius atemporalis (Bourret, 1934) – Tonkin keelback
Hebius bitaeniatus (Wall, 1925) – Kutkai keelback
Hebius boulengeri (Gressitt, 1937) – Tai-yong keelback, Boulenger’s keelback
Hebius celebicus (W. Peters & Doria, 1878) – Sulawesi keelback
Hebius chapaensis Bourret, 1934 – Vietnam water snake
Hebius clerki (Wall, 1925) – Yunnan keelback
Hebius concelarus (Malnate, 1963)
Hebius craspedogaster (Boulenger, 1899) – Kuatun keelback
Hebius deschauenseei (Taylor, 1934) – northern keelback, Deschauensee’s keelback
Hebius flavifrons (Boulenger, 1887) – Sabah keelback
Hebius frenatus (Dunn, 1923) – bridled keelback
Hebius groundwateri (M.A. Smith, 1922) – Groundwater's keelback
Hebius igneus David, G. Vogel, T.Q. Nguyen, Orlov, Pauwels, Teynié & Ziegler, 2021
Hebius inas (Laidlaw, 1901) – Malayan mountain keelback
Hebius ishigakiensis (Malnate & Munsterman, 1960) – Yaeyama keelback
Hebius johannis (Boulenger, 1908) – Johann's keelback
Hebius kerinciensis (David & Das, 2003)
Hebius khasiensis (Boulenger, 1890) – Khasi Hills keelback, Khasi keelback
Hebius lacrima Purkayastha & David, 2019 – crying keelback
Hebius leucomystax (David, Bain, T.Q. Nguyen, Orlov, G. Vogel, Thanh & Ziegler, 2007) – white-lipped keelback
Hebius maximus Malnate,.1962 – western China keelback
Hebius metusia (Inger, E. Zhao, Shaffer & G. Wu, 1990) – Wa Shan keelback
Hebius miyajimae (Maki, 1931) – Maki's keelback
Hebius modestus (Günther, 1875) – modest keelback
Hebius nigriventer (Wall, 1925)
Hebius octolineatus (Boulenger, 1904) – eight-lined keelback
Hebius optatus (Hu & E. Zhao, 1966) – Mount Omei keelback
Hebius parallelus (Boulenger, 1890) – Yunnan keelback
Hebius petersii (Boulenger, 1893) – Peters's keelback
Hebius popei (Schmidt, 1925) – Pope's keelback
Hebius pryeri (Boulenger, 1887) – Pryer's keelback
Hebius sanguineus (Smedley, 1931) – Cameron Highlands keelback
Hebius sangzhiensis Zhou et al., 2019^[3] – Sangzhi keelback
Hebius sarasinorum (Boulenger, 1896) – Sarasin’s keelback
Hebius sarawacensis (Günther, 1872) – Sarawak keelback
Hebius sauteri (Boulenger, 1909) – Kosempo keelback
Hebius septemlineatus (Schmidt, 1925) – Tengchong keelback
Hebius taronensis (M.A. Smith, 1940) – Kachin keelback
Hebius terrakarenorum Hauser, Smits, & David, 2022
Hebius venningi (Wall, 1910) – Chin Hills keelback
Hebius vibakari (H. Boie, 1826) – Japanese keelback
Hebius viperinus (Schenkel, 1901)
Hebius weixiensis Hou, Yuan, Wei, G. Zhao, G. Liu, Y. Wu, Shen, Chen, Guo & Che, 2021 – Weixi keelback
Hebius yanbianensis Q. Liu, Zhong, Wang, Y. Liu & Guo, 2018) – Yanbian keelback
Hebius youjiangensis Yang, Xu, Wu, Gong, Huang & Huang, 2023

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Hebius.

References[edit]

^ ^a ^b Genus Hebius at The Reptile Database.
^ ^a ^b Guo et al.
^ Zhou, Z.; Sun, Z.; Qi, S.; Lu, Y.; Lyu, Z.; Wang, Y.; Li, P.; Ma, J. (2019). "A new species of the genus Hebius (Squamata: Colubridae: Natricinae) from Hunan Province, China". Zootaxa. 4674 (1): 68–82. doi:10.11646/zootaxa.4674.1.3. PMID 31716020. S2CID 204136072.

@@ Line 1: / Line 1: @@
 {{Short description|Genus of snakes}}
 {{Automatic taxobox
-| image             = Amphiesma beddomei headtop.jpg
+| image = Hebius vibakari 04.jpg
-| image_caption     = [[Nilgiri keelback]] (''Hebius beddomei'')
+| image_caption = [[Japanese keelback]] (''Hebius vibakari'')
-| taxon             = Hebius
+| taxon = Hebius
-| authority         =  [[Joseph Cheesman Thompson|Thompson]], 1913
+| authority =  [[Joseph Cheesman Thompson|Thompson]], 1913
 | subdivision_ranks = Species
 }}
@@ Line 14: / Line 14: @@
 ==Taxonomy==
-All of the member [[species]] of the genus ''Hebius'' were formerly placed in the genus ''[[Amphiesma]]'', but in 2014 [[Peng Guo (herpetologist)|Guo]] et al. placed most species of ''Amphiesma'' in the genus ''Hebius''. They placed other species in the genus ''[[Herpetoreas]]'', leaving ''Amphiesma'' a [[monotypic taxon]] containing only the species ''[[Amphiesma stolatum]]''.<ref name=Guo/>
+All of the member [[species]] of the genus ''Hebius'' were formerly placed in the genus ''[[Amphiesma]]'', but in 2014 [[species:Peng Guo|Guo]] et al. placed most species of ''Amphiesma'' in the genus ''Hebius''. They placed other species in the genus ''[[Herpetoreas]]'', leaving ''Amphiesma'' a [[monotypic taxon]] containing only the species ''[[Amphiesma stolatum]]''.<ref name=Guo/>
 ==Species==
-The following 46 species are recognized as being valid.<ref name=RDB/>
+The following 48 species are recognized as being valid.<ref name=RDB/>
-*''[[Hebius andreae]]'' <small>([[:fr:Thomas Ziegler (zoologiste)|Ziegler]] & [[Le Khac Quyet|Quyet]], 2006)</small> – Andrea’s keelback
+*''[[Hebius andreae]]'' <small>([[:fr:Thomas Ziegler (zoologiste)|Ziegler]] & [[species:Le Khac Quyet|Quyet]], 2006)</small> – Andrea’s keelback
-*''[[Hebius annamensis]]'' <small>Bourret, 1934</small> – Annam keelback
+*''[[Hebius annamensis]]'' <small>[[René Léon Bourret|Bourret]], 1934</small> – Annam keelback
-*''[[Hebius arquus]]'' <small>([[Patrick David|David]] & [[Gernot Vogel|G. Vogel]], 2010)</small>
+*''[[Hebius arquus]]'' <small>([[species:Patrick David|David]] & [[species:Gernot Vogel|G. Vogel]], 2010)</small>
-*''[[Hebius atemporalis]]'' <small>([[René Léon Bourret|Bourret]], 1934)</small> – Tonkin keelback
+*''[[Hebius atemporalis]]'' <small>(Bourret, 1934)</small> – Tonkin keelback
-*''[[Hebius beddomei]]'' <small>([[Albert Günther|Günther]], 1864)</small> – Nilgiri keelback, Beddome’s keelback
 *''[[Hebius bitaeniatus]]'' <small>([[Frank Wall (herpetologist)|Wall]], 1925)</small> – Kutkai keelback
 *''[[Hebius boulengeri]]'' <small>([[J. Linsley Gressitt|Gressitt]], 1937)</small> – Tai-yong keelback, Boulenger’s keelback
@@ Line 28: / Line 27: @@
 *''[[Hebius chapaensis]]'' <small>Bourret, 1934</small> – Vietnam water snake
 *''[[Hebius clerki]]'' <small>(Wall, 1925)</small> – Yunnan keelback
-*''[[Hebius concelarus]]'' <small>([[Edmond Malnate|Malnate]], 1963)</small>
+*''[[Hebius concelarus]]'' <small>([[species:Edmond V. Malnate|Malnate]], 1963)</small>
 *''[[Hebius craspedogaster]]'' <small>([[George Albert Boulenger|Boulenger]], 1899)</small> – Kuatun keelback
 *''[[Hebius deschauenseei]]'' <small>([[Edward Harrison Taylor|Taylor]], 1934)</small> – northern keelback, Deschauensee’s keelback
@@ Line 34: / Line 33: @@
 *''[[Hebius frenatus]]'' <small>([[Emmett Reid Dunn|Dunn]], 1923)</small> – bridled keelback
 *''[[Hebius groundwateri]]'' <small>([[Malcolm Arthur Smith|M.A. Smith]], 1922)</small> – Groundwater's keelback
-*''[[Hebius igneus]]'' <small>David, Vogel, Nguyen, Orlov, Pauwels, Teynie, & Ziegler, 2021</small>
+*''[[Hebius igneus]]'' <small>David, G. Vogel, [[species:Truong Quang Nguyen|T.Q. Nguyen]], [[species:Nikolai Liutsianovich Orlov|Orlov]], [[species:Olivier Sylvain Gérard Pauwels|Pauwels]], [[species:Alexandre Teynié|Teynié]] & [[species:Thomas Ziegler|Ziegler]], 2021</small>
 *''[[Hebius inas]]'' <small>([[Frank Fortescue Laidlaw|Laidlaw]], 1901)</small> – Malayan mountain keelback
-*''[[Hebius ishigakiensis]]'' <small>(Malnate & [[Harold E. Munsterman|Munsterman]], 1960)</small> – Yaeyama keelback
+*''[[Hebius ishigakiensis]]'' <small>(Malnate & [[species:Harold E. Munsterman|Munsterman]], 1960)</small> – Yaeyama keelback
 *''[[Hebius johannis]]'' <small>(Boulenger, 1908)</small> – Johann's keelback
 *''[[Hebius kerinciensis]]'' <small>(David & [[:fr:Indraneil Das|Das]], 2003)</small>
 *''[[Hebius khasiensis]]'' <small>(Boulenger, 1890)</small> – Khasi Hills keelback, Khasi keelback
-*''[[Hebius lacrima]]'' <small>Purkayastha & David, 2019</small> – crying keelback
+*''[[Hebius lacrima]]'' <small>[[species:Jayaditya Purkayastha|Purkayastha]] & David, 2019</small> – crying keelback
-*''[[Hebius leucomystax]]'' <small>(David, [[Raoul H. Bain|Bain]], [[Nguyen Quang Truong|Truong]], [[:fr:Nikolaï Orlov|Orlov]], G. Vogel, [[Vu Ngoc Thanh|Thanh]] & Ziegler, 2007)</small> – white-lipped keelback
+*''[[Hebius leucomystax]]'' <small>(David, [[species:Raoul H. Bain|Bain]], T.Q. Nguyen, [[:fr:Nikolaï Orlov|Orlov]], G. Vogel, [[species:Vu Ngoc Thanh|Thanh]] & Ziegler, 2007)</small> – white-lipped keelback
-*''[[Hebius metusia]]'' <small>([[Robert F. Inger|Inger]], [[Er-mi Zhao|Zhao]], [[Howard Bradley Shaffer|Shaffer]] & [[Guanfu Wu|Wu]], 1990)</small> – Wa Shan keelback
+*''[[Hebius maximus]]'' <small>Malnate,.1962</small> – western China keelback
+*''[[Hebius metusia]]'' <small>([[Robert F. Inger|Inger]], [[Er-mi Zhao|E. Zhao]], [[species:H. Bradley Shaffer|Shaffer]] & [[species:Guan-fu Wu|G. Wu]], 1990)</small> – Wa Shan keelback
 *''[[Hebius miyajimae]]'' <small>([[:fr:Moichirō Maki|Maki]], 1931)</small> – Maki's keelback
 *''[[Hebius modestus]]'' <small>(Günther, 1875)</small> – modest keelback
-*''[[Hebius monticola]]'' <small>([[Thomas C. Jerdon|Jerdon]], 1853)</small> – Wynad keelback
-*''[[Hebius nicobariensis]]'' <small>(Sclater, 1891)</small> – Nicobar Island keelback
 *''[[Hebius nigriventer]]'' <small>(Wall, 1925)</small>
 *''[[Hebius octolineatus]]'' <small>(Boulenger, 1904)</small> – eight-lined keelback
-*''[[Hebius optatus]]'' <small>(Hu & Zhao, 1966)</small> – Mount Omei keelback
+*''[[Hebius optatus]]'' <small>([[species:Shu-qin |Hu]] & E. Zhao, 1966)</small> – Mount Omei keelback
 *''[[Hebius parallelus]]'' <small>(Boulenger, 1890)</small> – Yunnan keelback
 *''[[Hebius petersii]]'' <small>(Boulenger, 1893)</small> – Peters's keelback
 *''[[Hebius popei]]'' <small>([[Karl Patterson Schmidt|Schmidt]], 1925)</small> – Pope's keelback
 *''[[Hebius pryeri]]'' <small>(Boulenger, 1887)</small> – Pryer's keelback
-*''[[Hebius sanguineus]]'' <small>([[Norman Smedley|Smedley]], 1931)</small> – Cameron Highlands keelback
+*''[[Hebius sanguineus]]'' <small>([[species:Norman Smedley|Smedley]], 1931)</small> – Cameron Highlands keelback
-*''[[Hebius sangzhiensis]]'' <small>Zhou et al., 2019</small><ref name="ZT2019">{{cite journal |last1=Zhou |first1=Z. |last2=Sun |first2=Z. |last3=Qi |first3=S. |last4=Lu |first4=Y. |last5=Lyu |first5=Z. |last6=Wang |first6=Y. |last7=Li |first7=P. |last8=Ma |first8=J. |title=A new species of the genus ''Hebius'' (Squamata: Colubridae: Natricinae) from Hunan Province, China |journal=Zootaxa |date=2019 |volume=4674 |issue=1 |pages=68–82 |doi=10.11646/zootaxa.4674.1.3}}</ref> – Sangzhi keelback
+*''[[Hebius sangzhiensis]]'' <small>[[species:Zheng-Yan Zhou|Zhou]] et al., 2019</small><ref name="ZT2019">{{cite journal |last1=Zhou |first1=Z. |author-link=species:Zheng-Yan Zhou |last2=Sun |first2=Z. |author2-link=species:Zhi-Yong Sun |last3=Qi |first3=S. |author3-link=species:Shuo Qi |last4=Lu |first4=Y. |author4-link=species:Yu-Yan Lu |last5=Lyu |first5=Z. |author5-link=species:Zhi-Tong Lyu |last6=Wang |first6=Y. |author6-link=species:Ying-Yong Wang |last7=Li |first7=P. |author7-link=species:Pi-Peng Li |last8=Ma |first8=J. |author8-link=species:Jian-Zhang Ma |title=A new species of the genus ''Hebius'' (Squamata: Colubridae: Natricinae) from Hunan Province, China |journal=Zootaxa |date=2019 |volume=4674 |issue=1 |pages=68–82 |doi=10.11646/zootaxa.4674.1.3|pmid=31716020 |s2cid=204136072 }}</ref> – Sangzhi keelback
 *''[[Hebius sarasinorum]]'' <small>(Boulenger, 1896)</small> – Sarasin’s keelback
 *''[[Hebius sarawacensis]]'' <small>(Günther, 1872)</small> – Sarawak keelback
@@ Line 61: / Line 59: @@
 *''[[Hebius septemlineatus]]'' <small>(Schmidt, 1925)</small> – Tengchong keelback
 *''[[Hebius taronensis]]'' <small>(M.A. Smith, 1940)</small> – Kachin keelback
+*''[[Hebius terrakarenorum]]'' <small>Hauser, Smits, & David, 2022</small>
 *''[[Hebius venningi]]'' <small>(Wall, 1910)</small> – Chin Hills keelback
 *''[[Hebius vibakari]]'' <small>([[Heinrich Boie|H. Boie]], 1826)</small> – Japanese keelback
-*''[[Hebius viperinus]]'' <small>([[Ehrenfried Schenkel|Schenkel]], 1901)</small>
+*''[[Hebius viperinus]]'' <small>([[species:Ehrenfried Schenkel-Haas|Schenkel]], 1901)</small>
-*''[[Hebius weixiensis]]'' <small>Hou, Yuan, Wei, Zhao, Liu, Wu, Shen, Chen, Guo, & Che, 2021</small> – Weixi keelback
+*''[[Hebius weixiensis]]'' <small>[[species:Shao-Bing Hou|Hou]], [[species:Zhi-Yong Yuan|Yuan]], [[species:Ping-Fan Wei|Wei]], [[species:Gui-Gang Zhao|G. Zhao]], [[species:Gao-Hui Liu|G. Liu]], [[species:Yun-He Wu|Y. Wu]], [[species:Wen-Jing Shen|Shen]], [[species:Jin-Min Chen|Chen]], [[species:Peng Guo|Guo]] & [[species:Jing Che|Che]], 2021</small> – Weixi keelback
-*''[[Hebius yanbianensis]]'' <small>Liu, Zhong, Wang, Liu, & Guo, 2018)</small> – Yanbian keelback
+*''[[Hebius yanbianensis]]'' <small>[[species:Qin Liu|Q. Liu]], [[species:Guang-Hui Zhong|Zhong]], [[species:Ping Wang|Wang]], [[species:Yang Liu|Y. Liu]] & Guo, 2018)</small> – Yanbian keelback
+*''[[Hebius youjiangensis]]'' <small>Yang, Xu, Wu, Gong, Huang & Huang, 2023</small>
 ''[[Nota bene]]'': A [[Binomial nomenclature|binomial authority]] in parentheses indicates that the species was originally described in a genus other than ''Hebius''.
@@ Line 73: / Line 73: @@
 ==Further reading==
-*Guo, Peng; Zhu, Fei; Liu, Qin; Zhang, Liang; Li, Jian X.; Huang, Yu Y.; Pyron, R. Alexander (2014). "A taxonomic revision of the Asian keelback snakes, genus ''Amphiesma'' (Serpentes: Colubridae: Natricinae), with description of a new species". ''Zootaxa'' '''3873''' (4): 425–440.
+*[[species:Peng Guo|Guo, Peng]]; [[species:Fei Zhu|Zhu, Fei]]; [[species:Qin Liu|Liu, Qin]]; [[species:Liang Zhang|Zhang, Liang]]; Li, Jian X.; Huang, Yu Y.; [[species:Robert Alexander Pyron|Pyron, R. Alexander]] (2014). "A taxonomic revision of the Asian keelback snakes, genus ''Amphiesma'' (Serpentes: Colubridae: Natricinae), with description of a new species". ''Zootaxa'' '''3873''' (4): 425–440.
-*[[Joseph Cheesman Thompson|Thompson JC]] (1913). "Contributions to the Anatomy of the Ophidia". ''Proc. Zool. Soc. London'' '''1913''': 414–425. (''Hebius'', new genus, p.&nbsp;424).
+*[[Joseph Cheesman Thompson|Thompson JC]] (1913). "Contributions to the Anatomy of the Ophidia". ''Proceedings of the Zoological Society of London'' '''1913''': 414–425. (''Hebius'', new genus, p.&nbsp;424).
 {{Taxonbar|from=Q18616085}}
-[[Category:Hebius| ]]
+[[Category:Hebius]]
-[[Category:Colubrids|Hebius]]
 [[Category:Snake genera]]

Latest revision as of 19:55, 27 February 2024

Geographic range[edit]

Taxonomy[edit]

Species[edit]

References[edit]

Further reading[edit]