Boophis: Difference between revisions

Skeleton frogs
	Boophis ankaratra
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Subclass:	Lissamphibia
Order:	Anura
Suborder:	Neobatrachia
Family:	Mantellidae
Subfamily:	Boophinae; Vences & Glaw, 2001
Genus:	Boophis; Tschudi, 1838
Type species
	Boophis goudotii; Tschudi, 1838
Diversity
	Nearly 80 described species

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Boophis is the only genus in the mantellid frog subfamily Boophinae. They show typical 'tree frog' traits, and are a good example of convergent evolution with morphologically similar species in the families Hylidae and Rhacophoridae, among others. This genus can only be found on Madagascar and Mayotte Island (Comoros).^[1]

Taxonomy

The genus Boophis was described by Johann Jakob von Tschudi in 1838.^[2] It was originally considered a member of the African-Asian family Rhacophoridae, but was moved to the family Mantellidae in 2001,^[3] into its own subfamily, Boophinae Vences & Glaw, 2001.

Ecology

Many species of Boophis have almost translucent skin, allowing bones and internal organs to be observed as in the unrelated glass frogs (Centrolenidae) of the tropical Americas. This has led to the vernacular name skeleton frogs for the present genus.

Species

The genus has presently nearly 80 species; new ones are being described every few months on average. This list may not be exhaustive.

Boophis occidentalis tadpole

Boophis albilabris (Boulenger, 1888)
Boophis albipunctatus Glaw & Thiesmeier, 1993
Boophis andohahela Andreone, Nincheri & Piazza, 1995
Boophis andreonei Glaw & Vences, 1994
Boophis androngoloaka (Ahl, 1928)
Boophis anjanaharibeensis Andreone, 1996
Boophis ankaratra Andreone, 1993
Boophis ankarafensis Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014
Boophis arcanus Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis axelmeyeri Vences, Andreone & Veities, 2005
Boophis baetkei Köhler, Glaw & Vences, 2008
Boophis blommersae Glaw & Vences, 1994
Boophis boehmei Glaw & Vences, 1992
Boophis boppa Hutter & Lambert, 2015
Boophis bottae Vences & Glaw, 2002
Boophis brachychir Boettger, 1882
Boophis burgeri Glaw & Vences, 1994
Boophis calcaratus Vallan, Vences & Glaw, 2010
Boophis doulioti Angel, 1934
Boophis elenae Andreone, 1993
Boophis englaenderi Glaw & Vences, 1994
Boophis entingae Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis erythrodactylus (Guibé, 1953)
Boophis feonnyala Glaw, Vences, Andreone & Vallan, 2001
Boophis fayi Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
Boophis goudotii Tschudi, 1838
Boophis guibei (McCarthy, 1978)
Boophis haematopus Glaw, Vences, Andreone & Vallan, 2001
Boophis haingana Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis idae (Steindachner, 1867)
Boophis jaegeri Glaw & Vences, 1992 – Green Skeleton Frog
Boophis laurenti Guibé, 1947
Boophis liami Vallan, Vences & Glaw, 2003
Boophis lichenoides Vallan, Glaw, Andreone & Cadle, 1998
Boophis lilianae Köhler, Glaw & Vences, 2008
Boophis luciae Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis luteus (Boulenger, 1882)
Boophis madagascariensis (Peters, 1874)

Boophis majori (Boulenger, 1896)
Boophis mandraka Blommers-Schlösser, 1979
Boophis marojezensis Glaw & Vences, 1994
Boophis miadana Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis microtympanum (Boettger, 1881)
Boophis miniatus (Mocquard, 1902)
Boophis narinsi Vences, Gehara, Köhler & Glaw, 2012
Boophis obscurus (Boettger, 1913)
Boophis occidentalis Glaw & Vences, 1994
Boophis opisthodon (Boulenger, 1888)
Boophis pauliani (Guibé, 1953)
Boophis periegetes Cadle, 1995
Boophis picturatus Glaw, Vences, Andreone & Vallan, 2001
Boophis piperatus Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis popi Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
Boophis praedictus Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis pyrrhus Glaw, Vences, Andreone & Vallan, 2001
Boophis quasiboehmei Vences, Köhler, Crottini & Glaw, 2010
Boophis rappiodes (Ahl, 1928)
Boophis reticulatus Blommers-Schlösser, 1979
Boophis rhodoscelis (Boulenger, 1882)
Boophis roseipalmatus Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis rufioculis Glaw & Vences, 1997
Boophis sambirano Vences & Glaw, 2005
Boophis sandrae Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis schuboeae Glaw & Vences, 2002
Boophis septentrionalis Glaw & Vences, 1994
Boophis sibilans Glaw & Thiesmeier, 1993
Boophis solomaso Vallan, Vences & Glaw, 2003
Boophis spinophis Glaw, Köhler, de la Riva, Vieites & Vences, 2010
Boophis tampoka Köhler, Glaw & Vences, 2007
Boophis tasymena Vences & Glaw, 2002
Boophis tephraeomystax (Duméril, 1853)
Boophis tsilomaro Vences, Andreone, Glos & Glaw, 2010
Boophis ulftunni Wollenberg, Andreone, Glaw & Vences, 2008
Boophis viridis Blommers-Schlösser, 1979
Boophis vittatus Glaw, Vences, Andreone & Vallan, 2001
Boophis williamsi (Guibé, 1974)
Boophis xerophilus Glaw & Vences, 1997

References

^ Glaw, Frank; Vences, Miguel (2007). A Field Guide to the Amphibians and Reptiles of Madagascar (3rd ed.). Cologne, Germany: Vences & Glaw Verlags. ISBN 978-3929449037.
^ Tschudi, J. J. v. (1838). Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Neuchâtel: Petitpierre.
^ Vences, Miguel; Glaw, Frank (2001). "When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs". Spixiana. 24 (1): 85–92.

[GnV07-1] Glaw, Frank; Vences, Miguel (2007). A Field Guide to the Amphibians and Reptiles of Madagascar (3rd ed.). Cologne, Germany: Vences & Glaw Verlags. ISBN 978-3929449037.

[Tschudi38-2] Tschudi, J. J. v. (1838). Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Neuchâtel: Petitpierre.

[VG01-3] Vences, Miguel; Glaw, Frank (2001). "When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs". Spixiana. 24 (1): 85–92.

[1]

[2]

[3]

@@ Line 12: / Line 12: @@
 | familia = [[Mantellidae]]
 | subfamilia = '''Boophinae'''
-| subfamilia_authority = Vences & Glaw, 2001
+| subfamilia_authority = [[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2001
 | genus = '''''Boophis'''''
 | genus_authority = [[Johann Jakob von Tschudi|Tschudi]], 1838
@@ Line 24: / Line 24: @@
 ==Taxonomy==
-The genus ''Boophis'' was described by [[Johann Jakob von Tschudi]] in 1838.<ref name="Tschudi38">{{cite book|last1=Tschudi|first1=J. J. v.|title=Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien|date=1838|publisher=Petitpierre|location=Neuchâtel}}</ref> It was originally considered a member of the African-Asian [[Family (biology)|family]] [[Rhacophoridae]], but was moved to the family [[Mantellidae]] in 2001,<ref name="VG01">{{cite journal|last1=Vences|first1=Miguel|last2=Glaw|first2=Frank|title=When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs|journal=Spixiana|date=2001|volume=24|issue=1|pages=85–92}}</ref> into its own subfamily, '''Boophinae''' <small>Vences & Glaw, 2001</small>.
+The genus ''Boophis'' was described by [[Johann Jakob von Tschudi]] in 1838.<ref name="Tschudi38">{{cite book|last1=Tschudi|first1=J. J. v.|title=Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien|date=1838|publisher=Petitpierre|location=Neuchâtel}}</ref> It was originally considered a member of the African-Asian [[Family (biology)|family]] [[Rhacophoridae]], but was moved to the family [[Mantellidae]] in 2001,<ref name="VG01">{{cite journal|last1=Vences|first1=Miguel|last2=Glaw|first2=Frank|title=When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs|journal=Spixiana|date=2001|volume=24|issue=1|pages=85–92}}</ref> into its own subfamily, '''Boophinae''' <small>[[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2001</small>.
 ==Ecology==
@@ Line 38: / Line 38: @@
 * ''[[Boophis albipunctatus]]'' <small>[[Frank Glaw|Glaw]] & Thiesmeier, 1993</small>
 * ''[[Boophis andohahela]]'' <small>Andreone, Nincheri & Piazza, 1995</small>
-* ''[[Boophis andreonei]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis andreonei]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
 * ''[[Boophis androngoloaka]]'' <small>(Ahl, 1928)</small>
 * ''[[Boophis anjanaharibeensis]]'' <small>Andreone, 1996</small>
 * ''[[Boophis ankaratra]]'' <small>Andreone, 1993</small>
 * ''[[Boophis ankarafensis]]'' <small>Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014</small>
-* ''[[Boophis arcanus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis arcanus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
-* ''[[Boophis axelmeyeri]]'' <small>Vences, Andreone & Veities, 2005</small>
+* ''[[Boophis axelmeyeri]]'' <small>[[Miguel Vences|Vences]], Andreone & Veities, 2005</small>
-* ''[[Boophis baetkei]]'' <small>Köhler, [[Frank Glaw|Glaw]] & Vences, 2008</small><!--  Zootaxa1814:37. -->
+* ''[[Boophis baetkei]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small><!--  Zootaxa1814:37. -->
-* ''[[Boophis blommersae]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis blommersae]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
-* ''[[Boophis boehmei]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1992</small>
+* ''[[Boophis boehmei]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1992</small>
 * ''[[Boophis boppa]]'' <small>Hutter & Lambert, 2015</small>
-* ''[[Boophis bottae]]'' <small>Vences & [[Frank Glaw|Glaw]], 2002</small>
+* ''[[Boophis bottae]]'' <small>[[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2002</small>
 * ''[[Boophis brachychir]]'' <small>Boettger, 1882</small>
-* ''[[Boophis burgeri]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis burgeri]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
-* ''[[Boophis calcaratus]]'' <small>Vallan, Vences & Glaw, 2010</small>
+* ''[[Boophis calcaratus]]'' <small>Vallan, [[Miguel Vences|Vences]] & Glaw, 2010</small>
 * ''[[Boophis doulioti]]'' <small>Angel, 1934</small>
 * ''[[Boophis elenae]]'' <small>Andreone, 1993</small>
-* ''[[Boophis englaenderi]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis englaenderi]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
-* ''[[Boophis entingae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis entingae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
 * ''[[Boophis erythrodactylus]]'' <small>(Guibé, 1953)</small>
-* ''[[Boophis feonnyala]]'' <small>[[Frank Glaw|Glaw]], Vences, Andreone & Vallan, 2001</small>
+* ''[[Boophis feonnyala]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
-* ''[[Boophis fayi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & Vences, 2011</small>
+* ''[[Boophis fayi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & [[Miguel Vences|Vences]], 2011</small>
 * ''[[Boophis goudotii]]'' <small>Tschudi, 1838</small>
 * ''[[Boophis guibei]]'' <small>(McCarthy, 1978)</small>
-* ''[[Boophis haematopus]]'' <small>[[Frank Glaw|Glaw]], Vences, Andreone & Vallan, 2001</small>
+* ''[[Boophis haematopus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
-* ''[[Boophis haingana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis haingana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
 * ''[[Boophis idae]]'' <small>(Steindachner, 1867)</small>
-* ''[[Boophis jaegeri]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1992</small> &ndash; [[Green Skeleton Frog]]
+* ''[[Boophis jaegeri]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1992</small> &ndash; [[Green Skeleton Frog]]
 * ''[[Boophis laurenti]]'' <small>Guibé, 1947</small>
-* ''[[Boophis liami]]'' <small>Vallan, Vences & [[Frank Glaw|Glaw]], 2003</small>
+* ''[[Boophis liami]]'' <small>Vallan, [[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2003</small>
 * ''[[Boophis lichenoides]]'' <small>Vallan, [[Frank Glaw|Glaw]], Andreone & Cadle, 1998</small>
-* ''[[Boophis lilianae]]'' <small>Köhler, [[Frank Glaw|Glaw]] & Vences, 2008</small><!--  Zootaxa1814:37. -->
+* ''[[Boophis lilianae]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small><!--  Zootaxa1814:37. -->
-* ''[[Boophis luciae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis luciae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
 * ''[[Boophis luteus]]'' <small>(Boulenger, 1882)</small>
 * ''[[Boophis madagascariensis]]'' <small>(Peters, 1874)</small>
@@ Line 76: / Line 76: @@
 * ''[[Boophis majori]]'' <small>(Boulenger, 1896)</small>
 * ''[[Boophis mandraka]]'' <small>Blommers-Schlösser, 1979</small>
-* ''[[Boophis marojezensis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis marojezensis]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
-* ''[[Boophis miadana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis miadana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
 * ''[[Boophis microtympanum]]'' <small>(Boettger, 1881)</small>
 * ''[[Boophis miniatus]]'' <small>(Mocquard, 1902)</small>
-* ''[[Boophis narinsi]]'' <small>Vences, Gehara, Köhler & [[Frank Glaw|Glaw]], 2012</small>
+* ''[[Boophis narinsi]]'' <small>[[Miguel Vences|Vences]], Gehara, Köhler & [[Frank Glaw|Glaw]], 2012</small>
 * ''[[Boophis obscurus]]'' <small>(Boettger, 1913)</small>
 * ''[[Boophis occidentalis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
@@ Line 86: / Line 86: @@
 * ''[[Boophis pauliani]]'' <small>(Guibé, 1953)</small>
 * ''[[Boophis periegetes]]'' <small>Cadle, 1995</small>
-* ''[[Boophis picturatus]]'' <small>[[Frank Glaw|Glaw]], Vences, Andreone & Vallan, 2001</small>
+* ''[[Boophis picturatus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
-* ''[[Boophis piperatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis piperatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
-* ''[[Boophis popi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & Vences, 2011</small>
+* ''[[Boophis popi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & [[Miguel Vences|Vences]], 2011</small>
 * ''[[Boophis praedictus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis pyrrhus]]'' <small>[[Frank Glaw|Glaw]], Vences, Andreone & Vallan, 2001</small>
+* ''[[Boophis pyrrhus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
-* ''[[Boophis quasiboehmei]]'' <small>vences, Köhler, Crottini & [[Frank Glaw|Glaw]], 2010</small>
+* ''[[Boophis quasiboehmei]]'' <small>[[Miguel Vences|Vences]], Köhler, Crottini & [[Frank Glaw|Glaw]], 2010</small>
 * ''[[Boophis rappiodes]]'' <small>(Ahl, 1928)</small>
 * ''[[Boophis reticulatus]]'' <small>Blommers-Schlösser, 1979</small>
 * ''[[Boophis rhodoscelis]]'' <small>(Boulenger, 1882)</small>
-* ''[[Boophis roseipalmatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis roseipalmatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
 * ''[[Boophis rufioculis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1997</small>
 * ''[[Boophis sambirano]]'' <small>Vences & [[Frank Glaw|Glaw]], 2005</small>
-* ''[[Boophis sandrae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis sandrae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
-* ''[[Boophis schuboeae]]'' <small>[[Frank Glaw|Glaw]] & Vences, 2002</small>
+* ''[[Boophis schuboeae]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2002</small>
-* ''[[Boophis septentrionalis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis septentrionalis]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
 * ''[[Boophis sibilans]]'' <small>[[Frank Glaw|Glaw]] & Thiesmeier, 1993</small>
-* ''[[Boophis solomaso]]'' <small>Vallan, Vences & [[Frank Glaw|Glaw]], 2003</small>
+* ''[[Boophis solomaso]]'' <small>Vallan, [[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2003</small>
-* ''[[Boophis spinophis]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis spinophis]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
-* ''[[Boophis tampoka]]'' <small>Köhler, [[Frank Glaw|Glaw]] & Vences, 2007</small>
+* ''[[Boophis tampoka]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2007</small>
 * ''[[Boophis tasymena]]'' <small>Vences & [[Frank Glaw|Glaw]], 2002</small>
 * ''[[Boophis tephraeomystax]]'' <small>(Duméril, 1853)</small>
-* ''[[Boophis tsilomaro]]'' <small>Vences, Andreone, Glos & [[Frank Glaw|Glaw]], 2010</small>
+* ''[[Boophis tsilomaro]]'' <small>[[Miguel Vences|Vences]], Andreone, Glos & [[Frank Glaw|Glaw]], 2010</small>
-* ''[[Boophis ulftunni]]'' <small>Wollenberg, Andreone, [[Frank Glaw|Glaw]] & Vences, 2008</small><!-- Zootaxa1684:58. -->
+* ''[[Boophis ulftunni]]'' <small>Wollenberg, Andreone, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small><!-- Zootaxa1684:58. -->
 * ''[[Boophis viridis]]'' <small>Blommers-Schlösser, 1979</small>
-* ''[[Boophis vittatus]]'' <small>[[Frank Glaw|Glaw]], Vences, Andreone & Vallan, 2001</small>
+* ''[[Boophis vittatus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
 * ''[[Boophis williamsi]]'' <small>(Guibé, 1974)</small>
-* ''[[Boophis xerophilus]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1997</small>
+* ''[[Boophis xerophilus]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1997</small>
 {{col-end}}
 </div>