Hackelia

Hackelia
	Hackelia diffusa
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Boraginales
Family:	Boraginaceae
Subfamily:	Boraginoideae
Genus:	Hackelia; Opiz (1839)
Type species
	Hackelia deflexa; Opiz
Species
	About 54 (see text)
Synonyms
	Austrocynoglossum Popov ex R.Mill. (1989)

Hackelia (stickseeds) is a genus of plants in the borage family, Boraginaceae. They are found in North America and southeast Asia. Of the approximately 40 species described, 10 are endemic to California.

The genus was named after Josef Hackel, a Czech botanist. The common name, stickseed, refers to the tendency of the barbed nutlets to stick to animal fur.^[2]

Species

54 species are currently accepted:^[1]

Hackelia amethystina J.T.Howell - amethyst stickseed – California
Hackelia andicola (K.Krause) Brand – Peru
Hackelia bella (J.F.Macbr.) I.M.Johnst. - greater showy stickseed – northwestern California and southwestern Oregon
Hackelia besseyi (Rydb.) J.L. Gentry - Bessey's stickseed – Arizona, Colorado, New Mexico and Texas
Hackelia bhutanica R.R.Mill – eastern Nepal to northwestern Arunachal Pradesh
Hackelia brachytuba (Diels) I.M.Johnst. – Nepal to central China
Hackelia brevicula (Jeps.) J.L.Gentry - Poison Canyon stickseed – eastern California
Hackelia californica (A.Gray) I.M.Johnst. - California stickseed – northern California and west-central Oregon
Hackelia ciliata (Douglas ex Lehm.) I.M.Johnst. - British Columbia, Idaho, and Washington
Hackelia cinerea (Piper) I.M.Johnst. - gray stickseed – Washington, Idaho, and Montana
Hackelia cronquistii J.L.Gentry - Cronquist's stickseed – Oregon and Idaho
Hackelia cusickii (Piper) Brand - Cusick's stickseed – Oregon, northern California, and northwestern Nevada
Hackelia davisii Cronquist – Idaho
Hackelia deflexa (Wahlenb.) Opiz – northern Eurasia and northern North America
Hackelia difformis (Y.S.Lian & J.Q.Wang) Riedl – south-central China and Tibet
Hackelia diffusa (Douglas ex Lehm.) I.M.Johnst. - Spreading stickseed – British Columbia, Washington, Oregon
Hackelia floribunda (Lehm.) I.M.Johnst. - manyflower stickseed – western and central Canada and United States
Hackelia gracilenta I.M.Johnst. – Colorado
Hackelia heliocarpa (Brand) Brand – Mexico (Chihuahua)
Hackelia hendersonii (Piper) Brand – Washington, Oregon, and Montana
Hackelia hintoniorum (B.L.Turner) Sutorý – Mexico (Oaxaca)
Hackelia hirsuta (Wooton & Standl.) I.M.Johnst. – New Mexico
Hackelia hispida I.M.Johnst. - showy stickseed – Washington, Oregon, Idaho
Hackelia ibapensis L.M.Shultz & J.S.Shultz – Utah
Hackelia latifolia (R.Br.) Dimon & M.A.M.Renner – eastern Australia and Tasmania
Hackelia leonotis I.M.Johnst. – northeastern and central Mexico
Hackelia macrophylla (Brand) I.M.Johnst. – northern Pakistan and western Himalaya
Hackelia meeboldii Brand – western Himalaya
Hackelia mexicana (Schltdl. & Cham.) I.M.Johnst. – Mexico to Venezuela and Peru
Hackelia micrantha (Eastw.) J.L.Gentry - Jessica sticktight – western Canada and United States
Hackelia mundula (Jeps.) Ferris - pink stickseed – southwestern Oregon to central California
Hackelia murgabica Czukav. – Tajikistan
Hackelia nervosa (Kellogg) I.M.Johnst. - Sierra stickseed – California and northwestern Nevada
Hackelia obtusifolia R.R.Mill – Bhutan
Hackelia ophiobia R.L.Carr – southeastern Oregon, southwestern Idaho, and northern Nevada
Hackelia parviflora (K.Krause) Brand – Peru (Junín)
Hackelia patens (Nutt.) I.M.Johnst. - spotted stickseed – west-central United States
Hackelia pinetorum (Greene ex A.Gray) I.M.Johnst. – Arizona, New Mexico, Texas, and northern Mexico
Hackelia popovii Czukav. – Tajikistan
Hackelia rattanii (Brand) Brand – Mexico (Baja California)
Hackelia revoluta (Ruiz & Pav.) I.M.Johnst. – Colombia to northwestern Argentina
Hackelia sessilifructa (Y.S.Lian & J.Q.Wang) Ovczinnikova – Xinjiang
Hackelia setosa (Piper) I.M.Johnst. - bristly stickseed – northern California and southwestern Oregon
Hackelia sharsmithii I.M.Johnst. - Sharsmith's stickseed – central California and central Nevada
Hackelia skutchii I.M.Johnst. – Guatemala and Costa Rica
Hackelia stewartii I.M.Johnst. – northern Pakistan and western Himalaya
Hackelia stricta I.M.Johnst. – southwestern Mexico to Veracruz
Hackelia suaveolens (R.Br.) Dimon & M.A.M.Renner – eastern and southern Australia and Tasmania
Hackelia taylorii Harrod, Malmquist & R.L.Carr – Washington
Hackelia uncinata (Royle ex Benth.) C.E.C.Fisch. – northeastern Pakistan to Himalaya and southern China
Hackelia ursina (Greene ex A.Gray) I.M.Johnst. – southestern Arizona, southwestern New Mexico, and northern Mexico (Sonora)
Hackelia velutina (Piper) I.M.Johnst. - velvet stickseed – north-central and central California and northwestern Nevada
Hackelia venusta (Piper) H.St.John - lesser showy stickseed – central Washington
Hackelia virginiana (L.) I.M.Johnst. - beggar's lice – central and eastern United States and eastern Canada

References

^ ^a ^b Hackelia Opiz. Plants of the World Online. Retrieved 17 June 2023.
^ Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN 0-87842-280-3. OCLC 25708726.

External links

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[powo-1] Hackelia Opiz. Plants of the World Online. Retrieved 17 June 2023.

[2] Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN 0-87842-280-3. OCLC 25708726.

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