Hyaenodontidae

Hyaenodontidae Temporal range: early Eocene to early Miocene
Hyaenodon gigas and H. mongoliensis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Hyaenodonta
Superfamily:	†Hyaenodontoidea
Family:	†Hyaenodontidae Leidy, 1869
Type genus
†Hyaenodon Laizer & Parieu, 1838
Genera
[see text]
Synonyms
list of synonyms: Hyaenodontida (Haeckel, 1895) Hyaenodontoidea (Trouessart, 1885)

Hyaenodontidae ("hyena teeth") is a family of extinct predatory mammals from extinct superfamily Hyaenodontoidea within extinct order Hyaenodonta. Hyaenodontids arose during the early Eocene and persisted well into the early Miocene.

Classification and phylogeny

Taxonomy

• Family: †Hyaenodontidae (Leidy, 1869)
  • Genus: †Boritia (Solé, 2014)
    • †Boritia duffaudi (Solé, 2014)
  • Genus: †Preregidens (Solé, 2015)
    • †Preregidens langebadrae (Solé, 2015)
  • Genus: †Protoproviverra (Lemoine, 1891)
    • †Protoproviverra palaeonictides (Lemoine, 1880)
  • (unranked): †Cynohyaenodon/Quercytherium clade
    • Genus: †Cynohyaenodon ^{(paraphyletic genus)} (Filhol, 1873)
      • †Cynohyaenodon cayluxi (Filhol, 1873)
      • †Cynohyaenodon lautricensis (Lange-Badré, 1978)
      • †Cynohyaenodon ruetimeyeri (Van Valen, 1965)
      • †Cynohyaenodon smithae (Solé, 2021)
      • †Cynohyaenodon trux (Van Valen, 1965)
    • Genus: †Paracynohyaenodon (Martin, 1906)
      • †Paracynohyaenodon magnus (Crochet, 1988)
      • †Paracynohyaenodon schlosseri (Martin, 1906)
    • Genus: †Quercytherium (Filhol, 1880)
      • †Quercytherium simplicidens (Lange-Badré, 1975)
      • †Quercytherium tenebrosum (Filhol, 1880)
  • (unranked): †Eurotherium clade
    • Genus: †Alienetherium (Lange-Badré, 1981)
      • †Alienetherium buxwilleri (Lange-Badré, 1981)
    • Genus: †Cartierodon (Solé & Mennecart, 2019)
      • †Cartierodon egerkingensis (Solé & Mennecart, 2019)
    • Genus: †Eurotherium ^{(paraphyletic genus)} (Polly & Lange-Badré, 1993)
      • †Eurotherium mapplethorpei (Solé, 2021)
      • †Eurotherium matthesi (Lange-Badré & Haubold, 1990)
      • †Eurotherium theriodis (Van Valen, 1965)
    • Genus: †Paenoxyaenoides (Lange-Badre, 1979)
      • †Paenoxyaenoides liguritor (Lange-Badre, 1979)
    • Genus: †Prodissopsalis (Matthes, 1950)
      • †Prodissopsalis eocaenicus (Matthes, 1952)
  • (unranked): †Leonhardtina clade
    • Genus: †Leonhardtina (Matthes, 1952)
      • †Leonhardtina godinoti (Solé, 2014)
      • †Leonhardtina gracilis (Matthes, 1952)
      • †Leonhardtina meridianum (Solé, 2020)
  • (unranked): †Matthodon clade
    • Genus: †Matthodon (Lange-Badré & Haubold, 1990)
      • †Matthodon menui (Rich, 1971)
      • †Matthodon peignei (Solé, 2020)
      • †Matthodon tritens (Lange-Badré & Haubold, 1990)
  • (unranked): †Oxyaenoides clade
    • Subfamily: †Oxyaenoidinae (Lavrov, 1999) (syn. Francotheriini (Lavrov, 1999))
      • Genus: †Oxyaenoides (Matthes, 1967)
        • †Oxyaenoides aumelasiensis (Solé, 2020)
        • †Oxyaenoides bicuspidens (Matthes, 1967)
        • †Oxyaenoides lindgreni (Rich, 1971)
        • †Oxyaenoides schlosseri (Rütimeyer, 1891)
  • Subfamily: †Hyaenodontinae (Trouessart, 1885)
    • Genus: †Consobrinus (Lange-Badre, 1979)
      • †Consobrinus quercy (Lange-Badre, 1979)
    • Genus: †Propterodon ^{(paraphyletic genus)} (Martin, 1906)
      • †Propterodon morrisi (Matthew & Granger, 1924)
      • †Propterodon paganensis (de Bonis, 2018)
      • †Propterodon tongi (Liu & Huang, 2002)
      • †Propterodon witteri (Zack, 2019)
    • Tribe: †Epipterodontini (Lavrov, 1999)
      • Род: †Epipterodon (Lavrov, 1999)
        • †Epipterodon hyaenoides (Matthew & Granger, 1925)
      • Род: †Immanopterodon (Lavrov, 1999)
        • †Immanopterodon implacabilis (Lavrov, 1999)
    • Tribe: †Hyaenodontini (Leidy, 1869)
      • Genus: †Hyaenodon (Laizer & Parieu, 1838)

Phylogeny

The phylogenetic relationships of family Hyaenodontidae are shown in the following cladogram:

†Hyaenodontoidea	†Proviverridae File:Lesmesodon behnkeae juveline.jpg  †Hyaenodontidae   †Boritia  †Boritia duffaudi  †Quercytherium  †Quercytherium simplicidens †Quercytherium tenebrosum  †Paracynohyaenodon  †Paracynohyaenodon schlosseri †Paracynohyaenodon magnus †Cynohyaenodon lautricensis †Cynohyaenodon cayluxi †Cynohyaenodon smithae †Cynohyaenodon †Cynohyaenodon trux †Cynohyaenodon ruetimeyeri  †Protoproviverra  †Protoproviverra palaeonictides  ?   †Leonhardtina  †Leonhardtina gracilis †Leonhardtina godinoti †Leonhardtina meridianum  †Preregidens  †Preregidens langebadrae  †Prodissopsalis  †Prodissopsalis eocaenicus  †Cartierodon  †Cartierodon egerkingensis  †Paenoxyaenoides  †Paenoxyaenoides liguritor  ?  †Eurotherium theriodis †Eurotherium mapplethorpei †Eurotherium †Eurotherium matthesi  †Alienetherium  †Alienetherium buxwilleri    †Matthodon  †Matthodon peignei †Matthodon menui †Matthodon tritens    †Oxyaenoidinae   †Oxyaenoides  †Oxyaenoides bicuspidens †Oxyaenoides aumelasiensis †Oxyaenoides lindgreni †Oxyaenoides schlosseri    †Hyaenodontinae  †Propterodon morrisi †Propterodon sp. (ZIN no. 34494) †Propterodon paganensis †Propterodon tongi †Propterodon witteri  †Consobrinus  †Consobrinus quercy  ?   †Hyaenodontini  †Hyaenodon	†Cynohyaenodon/Quercytherium clade †Leonhardtina clade †Eurotherium clade †Matthodon clade †Oxyaenoides clade †Propterodon