from Wikipedia, the free encyclopedia
Beaded grain snail (Granaria frumentum)

Beaded grain snail ( Granaria frumentum )

Order : Lung snails (pulmonata)
Subordination : Land snails (Stylommatophora)
Superfamily : Pupilloidea
Family : Corn snails (Chondrinidae)
Subfamily : Granariinae
Genre : Granaria
Scientific name
Hero , 1838

Granaria is a genus of grain snails (Chondrinidae) from the suborder of land snails (Stylommatophora). The genus currently comprises 6 to 8 species, whereby the position of some taxa is still controversially discussed.


The conical-cylindrical to spindle-shaped housings are 7 to 10 mm high and 2.7 to 3.7 mm wide. They have 8 to 9 well-arched turns with a tightly rounded apex . The skin is rather thin and translucent. They are yellowish to light horn-colored. The embryonic convolutions have a characteristic ornamentation with fine grains. The postembryonic convolutions have fine, irregular stripes to more or less regular ribbing, which can disappear on the last convolutions. The mouth is rounded with a weakly pronounced parietal callus. The mouth edge is bent back and widened. Two lamellae, one angular and one parietal lamella, are formed on the parietal wall. There are two spindle lamellas. The palatal wall (palatal wall) has two to five plicae .

In the male part of the hermaphroditic sexual apparatus, the spermatic duct (vas deferens) is short and barely twisted and forms an epiphallus before it enters the penis . The moderately long penis has a comparatively very long blind sac (caecum or flagellum) that is about a third of the length of the penis. The penile retractor muscle splits into two broad ligaments that attach approximately in the middle of the penis and the epiphallus. The seminal vesicle (spermathec) reaches the albumin gland (protein gland), the stem of the seminal vesicle is usually very long and usually lies close to the prostate gland. The vagina is shorter than the penis, about the same length as the free fallopian tube (oviduct).

Geographical distribution and habitat

The species of the genus Granaria occur in Europe, Asia Minor, on the Arabian Peninsula (Yemen, Oman), in Iran and Pakistan.

They mostly live at the foot of rocks (mostly limestone rocks), not on the surface of the rocks, such as the species of the genus Chondrina Reichenbach, 1838.

Taxonomy and systematics

The taxon was set up by Friedrich Held in 1837 . The following species are currently assigned to the genus.

supporting documents


  • Edmund Gittenberger: Contributions to the knowledge of the pupillacea: III. Chondrininae. Zoologische Verhandelingen, 127 (1): 3-267, 1973 ISSN  0024-1652 PDF .
  • Michael P. Kerney, RAD Cameron & Jürgen H. Jungbluth: The land snails of Northern and Central Europe. 384 pp., Paul Parey, Hamburg & Berlin 1983 ISBN 3-490-17918-8
  • Bas Kokshoorn: "Resolving riddles and presenting new puzzles in Chondrinidae phylogenetics". Doctoral thesis, Leiden University , ISBN 978-90-71382-56-7 . Open Access
  • Bas Kokshoorn, Edmund Gittenberger: "Chondrinidae taxonomy revisited: new synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata)". Zootaxa, 2539: 1-62, 2010 Preview (PDF; 22 kB).
  • Anatolij A. Schileyko: Treatise on Recent terrestrial pulmonate molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. Ruthenica, Supplement 2 (1): 1-126, Moscow 1998 ISSN  0136-0027
  • Francisco W. Welter-Schultes: European non-marine molluscs, a guide for species identification = identification book for European land and freshwater mollusks. A1-A3 S., 679 S., Q1-Q78 S., Göttingen, Planet Poster Ed., 2012 ISBN 3-933922-75-5 , ISBN 978-3-933922-75-5

Individual evidence

  1. ^ Friedrich Held: Notes on the molluscs of Bavaria. (Continuation.). Isis , 1837 (12): 902-919, Leipzig 1838 Online at (p. 919).
  2. Fauna Europaea: Granaria Held 1838
  3. AnimalBase - Granaria Held, 1838

Web links

Commons : Granaria  - collection of images, videos and audio files