Sphaerocarpales

Sphaerocarpales
	Sphaerocarpos texanus
Systematics
without rank:	Streptophyta
Empire :	Plants (Plantae)
Department :	Liverworts (Marchantiophyta)
Class :	Marchantiopsida
Subclass :	Sphaerocarpidae
Order :	Sphaerocarpales
Scientific name of the subclass
	Sphaerocarpidae
	Stotler & Crand.-Stotl.
Scientific name of the order
	Sphaerocarpales
	Cavers

The Sphaerocarpales are an order of mosses from the Marchantiopsida class (which in turn belong to the liverworts ).

features

In terms of habit, the two families are very different: The Sphaerocarpaceae form round rosettes on moist soil, the Riellaceae form upright, winged stems and grow under water (submersed).

The common features are: Archegonia and sporogons are located in conspicuously pear-shaped, proliferating thallus tissue . Antheridia and archegonia have a single-layered shell. Oil bodies are missing. The Sporogon has no seta . The capsules are celistocarp and only release the spores , which are usually combined into tetrads, after the capsule has dissolved. No elaters are formed.

Systematics

The large systematic position of the Sphaerocarpidae was previously unclear, as their habitus differed greatly from the Marchantiidae. The lack of oil bodies in the normal lamina cells and the unicellular (leptosporangiate) sporogon wall justify their classification among the Marchantiopsida .

With only around 30 species in two sub-orders, the Sphaerocarpidae are one of the largest groups of liverworts with the fewest species:

Riellineae, monotypical
- Riella with 12 species, submerged in salt water.
Sphaerocarpineae with only two genera
- Sphaerocarpos with 12 species. In Germany
  - Sphaerocarpos michelii
  - Sphaerocarpos texanus
- Geothallus with only one species in California

literature

Jan-Peter Frahm, Wolfgang Frey, J. Döring: Moosflora . 4th, revised and expanded edition (UTB for Science, Volume 1250). Ulmer, Stuttgart 2004, ISBN 3-8001-2772-5 (Ulmer) & ISBN 3-8252-1250-5 (UTB)
Jan-Peter Frahm: Biology of Mosses . Spectrum Academic Publishing House, Heidelberg and Berlin 2001, ISBN 3-8274-0164-X
Photos of Naiadita lanceolata on page 87 (PDF-page 37)

Web links

Commons : Sphaerocarpales - collection of images, videos and audio files

Individual evidence

↑ Wolfgang Frey, Eberhard Fischer, Michael Stech: Bryophytes and seedless Vascular Plants . In: Wolfgang Frey (Ed.): Syllabus of Plant Families - A. Engler's Syllabus of Plant Families . 13th edition. tape 3 . Borntraeger, Berlin / Stuttgart 2009, ISBN 978-3-443-01063-8 , pp. 27-28 .

[1] Wolfgang Frey, Eberhard Fischer, Michael Stech: Bryophytes and seedless Vascular Plants . In: Wolfgang Frey (Ed.): Syllabus of Plant Families - A. Engler's Syllabus of Plant Families . 13th edition. tape 3 . Borntraeger, Berlin / Stuttgart 2009, ISBN 978-3-443-01063-8 , pp. 27-28 .