Sclerosomatidae: Difference between revisions

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{{Short description|Family of harvestmen/daddy longlegs}}
{{Taxobox
{{Taxobox
| fossil_range = {{Fossil range|Jurassic|present}}
| image = Opiliones harvestman.jpg
| image = Opiliones harvestman.jpg
| image_caption = ''[[Hadrobunus grandis]]''
| image_caption = ''[[Hadrobunus grandis]]''
| image_width = 250px
| image_width = 250px
| regnum = [[Animal]]ia
| regnum = [[Animal]]ia
| phylum = [[Arthropoda]]
| phylum = [[Arthropod]]a
| classis = [[Arachnida]]
| classis = [[Arachnid]]a
| ordo = [[Opiliones]]
| ordo = [[Opiliones]]
| subordo = [[Eupnoi]]
| subordo = [[Eupnoi]]
Line 11: Line 13:
| familia = '''Sclerosomatidae'''
| familia = '''Sclerosomatidae'''
| familia_authority = [[Eugène Simon|Simon]], 1879
| familia_authority = [[Eugène Simon|Simon]], 1879
| diversity_link = Sclerosomatidae
| diversity_link =
| diversity = 4 subfamilies, ca. 150 genera
| diversity = 4 subfamilies, ''c.'' 150 genera
| subdivision_ranks = Subfamilies
| subdivision_ranks = Subfamilies
| subdivision =
| subdivision =
[[Gagrellinae]] (117 genera)<br />
Gagrellinae (117 genera)<br />
[[Gyantinae]] (8 genera)<br />
Gyantinae (3 genera)<br />
[[Leiobuninae]] (13 genera)<br />
Leiobuninae (13 genera)<br />
[[Sclerosomatinae]] (10 genera)
Sclerosomatinae (10 genera)
}}
}}


The '''Sclerosomatidae''' are a [[family (taxonomy)|family]] of [[Opiliones|harvestmen]] with about 1,300 known species.<ref>{{citation|title=Sclerosomatidae Species List at Joel Hallan's Biology Catalog. Texas A&M University|url=http://insects.tamu.edu/research/collection/hallan/Acari/Family/Sclerosomatidae.txt|archive-url=https://web.archive.org/web/20141013064709/https://insects.tamu.edu/research/collection/hallan/Acari/Family/Sclerosomatidae.txt|access-date=30 August 2015|archive-date=13 October 2014|url-status=dead}}</ref> One former subfamily has been recently removed to form a new family, [[Globipedidae]].<ref>Kury, A.B. & J.C. Cokendolpher (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 (Kury et al., ed.): 52-54.</ref>
The '''Sclerosomatidae''' are a family of [[harvestmen]] with about 1,300 known species.


==Name==
==Name==
The name of the type genus is combined from [[Ancient Greek]] ''skleros'' "hard" and ''soma'' "body".<ref name=pinto-127ff>{{aut|Tourinho, Ana Lúcia}} (2007): Sclerosomatidae <small>Simon, 1879</small>. In: Pinto-da-Rocha ''et al.'' 2007: 127ff</ref>
The name is combined from [[Ancient Greek]] ''skleros'' "hard" and ''soma'' "body".<ref name=pinto-127ff>{{aut|Tourinho, Ana Lúcia}} (2007): Sclerosomatidae. Simon, 1879. In: Pinto-da-Rocha ''et al.'' 2007: 127ff</ref>


==Genera==
==Genera==
{{commons category|Sclerosomatidae}}
* [[Gagrellinae]] <small>Thorell, 1889</small>
:* ''[[Abaetetuba (harvestman)|Abaetetuba]]'' <small>Tourinho-Davis, 2004</small>
* '''[[Gagrellinae]]''' <small>Thorell, 1889</small>
:* ''[[Adungrella]]'' <small>Roewer, 1955</small>
:* ''[[Abaetetuba (harvestman)|Abaetetuba]]'' <small>Tourinho-Davis, 2004</small> (5 species)
:* ''[[Akalpia]]'' <small>Roewer, 1915</small>
:* ''[[Adungrella]]'' <small>Roewer, 1955</small> (3 species)
:* ''[[Altobunus]]'' <small>Roewer, 1910</small>
:* ''[[Aguilaia]]'' <small>González-Sponga, 2003</small> (1 species; Venezuela)
:* ''[[Amazonesia]]'' <small>Soares, 1970</small>
:* ''[[Akalpia]]'' <small>Roewer, 1915</small> (2 species; India and Japan)
:* ''[[Antigrella]]'' <small>Roewer, 1954</small>
:* ''[[Altobunus]]'' <small>Roewer, 1910</small> (3 species; Celebes and Philippines)
:* ''[[Aurivilliola]]'' <small>Roewer, 1910</small>
:* ''[[Amazonesia]]'' <small>Soares, 1970</small> (2 species)
:* ''[[Azucarella]]'' <small>Roewer, 1959</small>
:* ''[[Antigrella]]'' <small>Roewer, 1954</small> (1 species)
:* ''[[Bakerinulus]]'' <small>Roewer, 1955</small>
:* ''[[Aurivilliola]]'' <small>Roewer, 1910</small> (17 species; South and Southeast Asia)
:* ''[[Bastia (harvestman)|Bastia]]'' <small>Roewer, 1910</small>
:* ''[[Azucarella]]'' <small>Roewer, 1959</small> (1 species)
:* ''[[Bastioides]]'' <small>Mello-Leitão, 1931</small>
:* ''[[Bakerinulus]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Baturitia]]'' <small>Roewer, 1931</small>
:* ''[[Bastia (harvestman)|Bastia]]'' <small>Roewer, 1910</small> (3 species; South and Southeast Asia)
:* ''[[Biceropsis]]'' <small>Roewer, 1935</small>
:* ''[[Bastioides]]'' <small>Mello-Leitão, 1931</small> (1 species; South America)
:* ''[[Bonthainia]]'' <small>Roewer, 1913</small>
:* ''[[Baturitia]]'' <small>Roewer, 1931</small> (1 species; Sunda Islands)
:* ''[[Bullobunus]]'' <small>Roewer, 1910</small>
:* ''[[Bellonia (Opiliones)|Bellonia]]'' <small>González-Sponga, 2003</small> (1 species; Venezuela)
:* ''[[Caiza]]'' <small>Roewer, 1925</small>
:* ''[[Biceropsis]]'' <small>Roewer, 1935</small> (1 species; Burma)
:* ''[[Caluga]]'' <small>Roewer, 1959</small>
:* ''[[Bonthainia]]'' <small>Roewer, 1913</small> (5 species; South and Southeast Asia)
:* ''[[Carinobius]]'' <small>Roewer, 1955</small>
:* ''[[Bullobunus]]'' <small>Roewer, 1910</small> (7 species; Philippines)
:* ''[[Carmenia]]'' <small>Roewer, 1915</small>
:* ''[[Caluga]]'' <small>Roewer, 1959</small> (1 species)
:* ''[[Carmichaelus]]'' <small>Roewer, 1929</small>
:* ''[[Cardenalia]]'' <small>Kury, 2017</small> (1 species; Venezuela)
:* ''[[Ceratobunellus]]'' <small>Roewer, 1911</small>
:* ''[[Carinobius]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Ceratobunoides]]'' <small>Roewer, 1923</small>
:* ''[[Carmenia]]'' <small>Roewer, 1915</small> (1 species; Colombia)
:* ''[[Cervibunus]]'' <small>Roewer, 1912</small>
:* ''[[Carmichaelus]]'' <small>Roewer, 1929</small> (1 species; India)
:* ''[[Chasenella]]'' <small>Roewer, 1955</small>
:* ''[[Ceratobunellus]]'' <small>Roewer, 1911</small> (3 species; India)
:* ''[[Chebabius]]'' <small>Roewer, 1935</small>
:* ''[[Ceratobunoides]]'' <small>Roewer, 1923</small> (2 species; Sumatra)
:* ''[[Coonoora]]'' <small>Roewer, 1929</small>
:* ''[[Ceratobunus]]'' <small>Thorell, 1889</small> (13 species)
:* ''[[Dentobunus]]'' <small>Roewer, 1910</small>
:* ''[[Cervibunus]]'' <small>Roewer, 1912</small> (3 Species; South and Southeast Asia)
:* ''[[Diangathia]]'' <small>Roewer, 1955</small>
:* ''[[Chasenella]]'' <small>Roewer, 1955</small> (2 species; Borneo)
:* ''[[Echinobunus]]'' <small>Roewer, 1912</small>
:* ''[[Chebabius]]'' <small>Roewer, 1935</small> (1 species; Burma)
:* ''[[Euceratobunus]]'' <small>Roewer, 1923</small>
:* ''[[Coonoora]]'' <small>Roewer, 1929</small> (1 species; India)
:* ''[[Eugagrella]]'' <small>Roewer, 1910</small>
:* ''[[Dentobunus]]'' <small>Roewer, 1910</small> (33 species; Southeast Asia)
:* ''[[Euzaleptus]]'' <small>Roewer, 1911</small>
:* ''[[Diangathia]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Fesa]]'' <small>Roewer, 1953</small>
:* ''[[Echinobunus]]'' <small>Roewer, 1912</small> (1 species; Java)
:* ''[[Gagrella]]'' <small>Stoliczka, 1869</small>
:* ''[[Euceratobunus]]'' <small>Roewer, 1923</small> (1 species; India)
:* ''[[Gagrellenna]]'' <small>Roewer, 1929</small>
:* ''[[Eugagrella]]'' <small>Roewer, 1910</small> (25 species; South and Southeast Asia)
:* ''[[Gagrellina]]'' <small>Roewer, 1913</small>
:* ''[[Euzaleptus]]'' <small>Roewer, 1911</small> (4 species; South and Southeast Asia)
:* ''[[Gagrellissa]]'' <small>Roewer, 1931</small>
:* ''[[Fesa tricolor|Fesa]]'' <small>Roewer, 1953</small> (1 species)
:* ''[[Gagrellopsis]]'' <small>Sato & Suzuki, 1939</small>
:* ''[[Gagrella]]'' <small>Stoliczka, 1869</small> (226 species; Asia)
:* ''[[Gagrellula]]'' <small>Roewer, 1910</small>
:* ''[[Gagrellenna]]'' <small>Roewer, 1929</small> (1 species; India)
:* ''[[Geaya]]'' <small>Roewer, 1910</small>
:* ''[[Gagrellina]]'' <small>Roewer, 1913</small> (1 species; Celebes)
:* ''[[Globulosoma]]'' <small>Martens, 1987</small>
:* ''[[Gagrellissa]]'' <small>Roewer, 1931</small> (1 species; Sunda Islands)
:* ''[[Guaranobunus]]'' <small>Ringuelet, 1954</small>
:* ''[[Gagrellopsis]]'' <small>Sato & Suzuki, 1939</small> (1 species; Japan)
:* ''[[Hamitergum]]'' <small>Crawford, 1992</small>
:* ''[[Gagrellula]]'' <small>Roewer, 1910</small> (67 species; Asia)
:* ''[[Harmanda]]'' <small>Roewer, 1910</small>
:* ''[[Garleppa]]'' <small>Roewer, 1912</small> (4 species; South America)
:* ''[[Harmandina]]'' <small>Schenkel, 1954</small>
:* ''[[Geaya]]'' <small>Roewer, 1910</small> (82 species; Latin America)
:* ''[[Hehoa]]'' <small>Roewer, 1929</small>
:* ''[[Globulosoma]]'' <small>Martens, 1987</small> (2 species; Nepal)
:* ''[[Heterogagrella]]'' <small>Roewer, 1954</small>
:* ''[[Guaranobunus]]'' <small>Ringuelet, 1954</small> (1 species; Argentina)
:* ''[[Hexazaleptus]]'' <small>Suzuki, 1966</small>
:* ''[[Hamitergum]]'' <small>Crawford, 1992</small> (1 species; Russia)
:* ''[[Himaldroma]]'' <small>Martens, 1987</small>
:* ''[[Harmanda]]'' <small>Roewer, 1910</small> (15 species; Asia)
:* ''[[Himalzaleptus]]'' <small>Martens, 1987</small>
:* ''[[Harmandina]]'' <small>Schenkel, 1954</small> (1 species)
:* ''[[Holcobunus]]'' <small>Roewer, 1910</small>
:* ''[[Hehoa]]'' <small>Roewer, 1929</small> (1 species; India)
:* ''[[Paratamboicus]]'' <small>Mello-Leitão, 1940</small>
:* ''[[Heterogagrella]]'' <small>Roewer, 1954</small> (2 species; Southeast Asia)
:* ''[[Holmbergiana]]'' <small>Mello-Leitão, 1931</small>
:* ''[[Hexazaleptus]]'' <small>Suzuki, 1966</small> (1 species)
:* ''[[Hologagrella]]'' <small>Roewer, 1910</small>
:* ''[[Hexomma]]'' <small>Thorell, 1876</small> (9 species; Southeast Asia)
:* ''[[Hypogrella]]'' <small>Roewer, 1955</small>
:* ''[[Himaldroma]]'' <small>Martens, 1987</small> (2 species; Nepal)
:* ''[[Jussara (harvestman)|Jussara]]'' <small>Mello-Leitão, 1935</small>
:* ''[[Himalzaleptus]]'' <small>Martens, 1987</small> (1 species; Nepal)
:* ''[[Koyamaia]]'' <small>Suzuki, 1972</small>
:* ''[[Holcobunus]]'' <small>Roewer, 1910</small> (26 or 27 species; Latin America)
:* ''[[Krusa (genus)|Krusa]]'' <small>Goodnight & Goodnight, 1947</small>
:* ''[[Holmbergiana]]'' <small>Mello-Leitão, 1931</small> (3 species; South America)
:* ''[[Liopagus]]'' <small>Chamberlin, 1916</small>
:* ''[[Hologagrella]]'' <small>Roewer, 1910</small> (7 species; Southeast Asia)
:* ''[[Marthana]]'' <small>Thorell, 1891</small>
:* ''[[Hypogrella]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Melanopella]]'' <small>Roewer, 1931</small>
:* ''[[Hypsibunus]]'' <small>Thorell, 1891</small> (11 species; India & Southeast Asia)
:* ''[[Melanopula]]'' <small>Roewer, 1929</small>
:* ''[[Jussara (harvestman)|Jussara]]'' <small>Mello-Leitão, 1935</small> (10 species; South America)
:* ''[[Metadentobunus]]'' <small>Roewer, 1915</small>
:* ''[[Koyamaia]]'' <small>Suzuki, 1972</small> (1 species)
:* ''[[Metahehoa]]'' <small>Suzuki, 1985</small>
:* ''[[Krusa (harvestman)|Krusa]]'' <small>Goodnight & Goodnight, 1947</small> (10 species; Mexico)
:* ''[[Metasyleus]]'' <small>Roewer, 1955</small>
:* ''[[Krusella]]'' <small>Roewer, 1953</small> (1 species; Venezuela)
:* ''[[Metaverpulus]]'' <small>Roewer, 1912</small>
:* ''[[Liopagus]]'' <small>Chamberlin, 1916</small> (1 species)
:* ''[[Metazaleptus]]'' <small>Roewer, 1912</small>
:* ''[[Marthana]]'' <small>Thorell, 1891</small> (24 species; Southeast Asia)
:* ''[[Microzaleptus]]'' <small>Roewer, 1955</small>
:* ''[[Melanopa]]'' <small>Thorell, 1889</small> (30 species; India, Southeast Asia)
:* ''[[Munequita]]'' <small>Mello-Leitão, 1941</small>
:* ''[[Melanopella]]'' <small>Roewer, 1931</small> (3 species; Sunda Islands)
:* ''[[Neogagrella]]'' <small>Roewer, 1913</small>
:* ''[[Melanopula]]'' <small>Roewer, 1929</small> (5 species; South and Southeast Asia)
:* ''[[Metadentobunus]]'' <small>Roewer, 1915</small> (3 species; Taiwan)<ref name=Chen>{{cite journal|last1=Chen|first1=S. L.|last2=Shih|first2=H. T.|year=2018|title=A review of the harvestman genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with a description of a new species from Taiwan|url=http://zoolstud.sinica.edu.tw/Journals/57/57-57.html|journal=Zoological Studies|volume=57}}</ref>
:* ''[[Nepalgrella]]'' <small>Martens, 1987</small>
:* ''[[Nepalkanchia]]'' <small>Martens, 1990</small>
:* ''[[Metahehoa]]'' <small>Suzuki, 1985</small> (1 species)
:* ''[[Obigrella]]'' <small>Roewer, 1955</small>
:* ''[[Metasyleus]]'' <small>Roewer, 1955</small> (3 species)
:* ''[[Octozaleptus]]'' <small>Suzuki, 1966</small>
:* ''[[Metaverpulus]]'' <small>Roewer, 1912</small> (7 species; South Asia)
:* ''[[Onostemma]]'' <small>Mello-Leitão, 1938</small>
:* ''[[Metazaleptus]]'' <small>Roewer, 1912</small> (8 species; South and Southeast Asia)
:* ''[[Oobunus]]'' <small>Kishida, 1930</small>
:* ''[[Microzaleptus]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Orissula]]'' <small>Roewer, 1955</small>
:* ''[[Munequita]]'' <small>Mello-Leitão, 1941</small> (1 species; Brazil)
:* ''[[Padangrella]]'' <small>Roewer, 1954</small>
:* ''[[Neogagrella]]'' <small>Roewer, 1913</small> (3 species; South and Southeast Asia)
:* ''[[Palniella]]'' <small>Roewer, 1929</small>
:* ''[[Nepalgrella]]'' <small>Martens, 1987</small> (2 species; Nepal)
:* ''[[Paradentobunus]]'' <small>Roewer, 1915</small>
:* ''[[Nepalkanchia]]'' <small>Martens, 1990</small> (2 species; Nepal)
:* ''[[Paragagrella]]'' <small>Roewer, 1912</small>
:* ''[[Obigrella]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Paragagrellina]]'' <small>Schenkel, 1963</small>
:* ''[[Octozaleptus]]'' <small>Suzuki, 1966</small> (1 species)
:* ''[[Parageaya]]'' <small>Mello-Leitão, 1933</small>
:* ''[[Onostemma]]'' <small>Mello-Leitão, 1938</small> (2 species; Brazil)
:* ''[[Paraumbogrella]]'' <small>Suzuki, 1963</small>
:* ''[[Oobunus]]'' <small>Kishida, 1930</small> (1 species)
:* ''[[Paruleptes]]'' <small>Soares, 1970</small>
:* ''[[Orissula]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Pectenobunus]]'' <small>Roewer, 1910</small>
:* ''[[Padangrella]]'' <small>Roewer, 1954</small> (1 species)
:* ''[[Pergagrella]]'' <small>Roewer, 1954</small>
:* ''[[Palniella]]'' <small>Roewer, 1929</small> (1 species; India)
:* ''[[Pokhara (harvestman)|Pokhara]]'' <small>Suzuki, 1970</small>
:* ''[[Paradentobunus]]'' <small>Roewer, 1915</small> (1 species; India)
:* ''[[Prionostemma]]'' <small>Pocock, 1903</small>
:* ''[[Paragagrella]]'' <small>Roewer, 1912</small> (5 species; South and Southeast Asia)
:* ''[[Prodentobunus]]'' <small>Roewer, 1923</small>
:* ''[[Paragagrellina]]'' <small>Schenkel, 1963</small> (1 species)
:* ''[[Psammogeaya]]'' <small>Mello-Leitão, 1946</small>
:* ''[[Parageaya]]'' <small>Mello-Leitão, 1933</small> (6 species; Latin America)
:* ''[[Psathyropus]]'' <small>L. Koch, 1878</small>
:* ''[[Paratamboicus]]'' <small>Mello-Leitão, 1940</small> (30 species; South America)
:* ''[[Pseudarthromerus]]'' <small>Karsch, 1892</small>
:* ''[[Paraumbogrella]]'' <small>Suzuki, 1963</small> (1 species)
:* ''[[Pseudogagrella]]'' <small>Redikortsev, 1936</small>
:* ''[[Paruleptes]]'' <small>Soares, 1970</small> (1 species)
:* ''[[Pseudomelanopa]]'' <small>Suzuki, 1974</small>
:* ''[[Pectenobunus]]'' <small>Roewer, 1910</small> (4 species; South America)
:* ''[[Pseudosystenocentrus]]'' <small>Suzuki, 1985</small>
:* ''[[Pergagrella]]'' <small>Roewer, 1954</small> (1 species)
:* ''[[Romerella]]'' <small>Goodnight & Goodnight, 1943</small>
:* ''[[Pokhara (harvestman)|Pokhara]]'' <small>Suzuki, 1970</small> (7 species; Nepal)
:* ''[[Sarasinia]]'' <small>Roewer, 1913</small>
:* ''[[Prionostemma]]'' <small>Pocock, 1903</small> (114 species; Mexico, Central America, and South America)
:* ''[[Sataria]]'' <small>Roewer, 1915</small>
:* ''[[Prodentobunus]]'' <small>Roewer, 1923</small> (4 species; Asia)
:* ''[[Scotomenia]]'' <small>Thorell, 1889</small>
:* ''[[Psammogeaya]]'' <small>Mello-Leitão, 1946</small> (1 species; Uruguay)
:* ''[[Sericicorpus]]'' <small>Martens, 1987</small>
:* ''[[Psathyropus]]'' <small>L. Koch, 1878</small> (40 species; Asia)
:* ''[[Sinadroma]]'' <small>Roewer, 1955</small>
:* ''[[Pseudarthromerus]]'' <small>Karsch, 1892</small> (1 species)
:* ''[[Syleus]]'' <small>Thorell, 1876</small>
:* ''[[Pseudogagrella]]'' <small>Redikortsev, 1936</small> (13 species; East Asia)
:* ''[[Syngagrella]]'' <small>Roewer, 1913</small>
:* ''[[Pseudomelanopa]]'' <small>Suzuki, 1974</small> (2 species; East Asia)
:* ''[[Systenocentrus]]'' <small>Simon, 1886</small>
:* ''[[Pseudosystenocentrus]]'' <small>Suzuki, 1985</small> (1 species)
:* ''[[Tamboicus]]'' <small>Roewer, 1912</small>
:* ''[[Romerella]]'' <small>Goodnight & Goodnight, 1943</small> (5 species; Mexico and South America)
:* ''[[Taperina]]'' <small>Roewer, 1953</small>
:* ''[[Sarasinia]]'' <small>Roewer, 1913</small> (1 species; Celebes)
:* ''[[Tetraceratobunus]]'' <small>Roewer, 1915</small>
:* ''[[Sataria]]'' <small>Roewer, 1915</small> (3 species; India)
:* ''[[Toragrella]]'' <small>Roewer, 1955</small>
:* ''[[Scotomenia]]'' <small>Thorell, 1889</small> (1 species; Burma)
:* ''[[Trachyrhinus]]'' <small>Weed, 1892</small>
:* ''[[Sericicorpus]]'' <small>Martens, 1987</small> (1 species; Nepal)
:* ''[[Umbogrella]]'' <small>Roewer, 1955</small>
:* ''[[Sinadroma]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Verpulus]]'' <small>Simon, 1901</small>
:* ''[[Syleus]]'' <small>Thorell, 1876</small> (2 species; India)
:* ''[[Verrucobunus]]'' <small>Roewer, 1931</small>
:* ''[[Syngagrella]]'' <small>Roewer, 1913</small> (1 species; Celebes)
:* ''[[Xerogrella]]'' <small>Martens, 1987</small>
:* ''[[Systenocentrus]]'' <small>Simon, 1886</small> (6 species; Southeast and East Asia)
:* ''[[Zaleptiolus]]'' <small>Roewer, 1955</small>
:* ''[[Tamboicus]]'' <small>Roewer, 1912</small> (3 species; South America)
:* ''[[Zaleptulus]]'' <small>Roewer, 1955</small>
:* ''[[Taperina]]'' <small>Roewer, 1953</small> (2 species)
:* ''[[Zaleptus]]'' <small>Thorell, 1876</small>
:* ''[[Tetraceratobunus]]'' <small>Roewer, 1915</small> (3 species; South Asia)
:* ''[[Pseudoarthromerus]]'' <small>Karsch, 1891</small>
:* ''[[Thunbergia (harvestman)|Thunbergia]]'' <small>Martens, 2020</small> (4 species; South America)
:* ''[[Toragrella]]'' <small>Roewer, 1955</small> (2 species)
:* ''[[Trachyrhinus]]'' <small>Weed, 1892</small> (7 species; North America: Mexico and United States)
:* ''[[Umbogrella]]'' <small>Roewer, 1955</small> (1 species)
:* ''[[Umbopilio]]'' <small>Roewer, 1956</small> (2 species)
:* ''[[Vaho]]'' <small>González-Sponga, 2003</small> (1 species)
:* ''[[Valle (harvestman)|Valle]]'' <small>González-Sponga, 2003</small> (3 species)
:* ''[[Varinodulia]]'' <small>Canals, 1935</small> (1 species)
:* ''[[Verpulus]]'' <small>Simon, 1901</small> (13 species; South and Southeast Asia)
:* ''[[Verrucobunus]]'' <small>Roewer, 1931</small> (3 species; East and Southeast Asia)
:* ''[[Xerogrella]]'' <small>Martens, 1987</small> (1 species; Nepal)
:* ''[[Zaleptiolus]]'' <small>Roewer, 1955</small> (4 species; Asia)
:* ''[[Zaleptulus]]'' <small>Roewer, 1955</small> (3 species)
:* ''[[Zaleptus]]'' <small>Thorell, 1876</small> (72 species; Asia and Australia)


* [[Gyantinae]] <small>Silhavý, 1946</small>
* '''Gyantinae''' <small>Silhavý, 1946</small>
:* ''[[Gyas (harvestman)|Gyas]]'' <small>Simon, 1879</small>
:* ''[[Gyas (harvestman)|Gyas]]'' <small>Simon, 1879</small> (1 species; Alps)
:* ''[[Gyoides]]'' <small>Martens, 1982</small>
:* ''[[Gyoides]]'' <small>Martens, 1982</small> (6 species; Nepal)
:* ''[[Rongsharia]]'' <small>Roewer, 1957</small>
:* ''[[Rongsharia]]'' <small>Roewer, 1957</small> (3 species; Nepal)
:* ''[[Dalquestia]]'' <small>Cokendolpher, 1984</small>
:* ''[[Diguetinus]]'' <small>Roewer, 1912</small>
:* ''[[Eurybunus]]'' <small>Banks, 1893</small>
:* ''[[Globipes]]'' <small>Banks, 1893</small>
:* ''[[Metopilio]]'' <small>Roewer, 1911</small>


* [[Leiobuninae]] <small>Banks, 1893</small>
* '''Leiobuninae''' <small>Banks, 1893</small>
:* † ''[[Amauropilio]]'' <small>Mello-Leitão, 1937 (fossil)</small>
:* †''[[Amauropilio]]'' <small>Mello-Leitão, 1937 (fossil)</small> (2 species; fossils found in Colorado)
:* ''[[Cosmobunus]]'' <small>Simon, 1879</small>
:* ''[[Cosmobunus]]'' <small>Simon, 1879</small> (5 species; Spain and Mexico)
:* ''[[Dilophiocara]]'' <small>Redikorzev, 1931</small>
:* ''[[Dilophiocara]]'' <small>Redikorzev, 1931</small> (2 species)
:* ''[[Eumesosoma]]'' <small>Cokendolpher, 1980</small>
:* ''[[Eumesosoma]]'' <small>Cokendolpher, 1980</small> (6 species; United States)
:* ''[[Eusclera]]'' <small>Roewer, 1910</small>
:* ''[[Eusclera]]'' <small>Roewer, 1910</small> (2 species; China and India)
:* ''[[Hadrobunus]]'' <small>Banks, 1900</small>
:* ''[[Hadrobunus]]'' <small>Banks, 1900</small> (2 species; United States)
:* ''[[Leiobunum]]'' <small>C.L.Koch, 1839</small>
:* ''[[Leiobunum]]'' <small>C.L.Koch, 1839</small> (126 species - 2 extinct fossils; worldwide)
:* ''[[Leuronychus]]'' <small>Banks, 1900</small>
:* ''[[Leuronychus]]'' <small>Banks, 1900</small> (2 species; North America)
:* ''[[Microliobunum]]'' <small>Roewer, 1912</small>
:* ''[[Microliobunum]]'' <small>Roewer, 1912</small> (2 species; Middle East)
:* ''[[Micronelima]]'' <small>Schenkel, 1938</small>
:* ''[[Micronelima]]'' <small>Schenkel, 1938</small> (1 species; Spain)
:* ''[[Nelima]]'' <small>Roewer, 1910</small>
:* ''[[Nelima]]'' <small>Roewer, 1910</small> (44 species; worldwide)
:* ''[[Paranelima]]'' <small>Caporiacco, 1938</small>
:* ''[[Paranelima]]'' <small>Caporiacco, 1938</small> (6 species; Mexico and Central America)
:* ''[[Schenkeliobunum]]'' <small>Starega, 1964</small>
:* ''[[Schenkeliobunum]]'' <small>Starega, 1964</small> (1 species)
:* ''[[Togwoteeus]]'' <small>Roewer, 1952</small>
:* ''[[Togwoteeus]]'' <small>Roewer, 1952</small> (1 species; United States)


* [[Sclerosomatinae]] <small>Simon, 1879</small>
* '''Sclerosomatinae''' <small>Simon, 1879</small>
:* ''[[Astrobunus]]'' <small>Thorell, 1876</small>
:* ''[[Astrobunus]]'' <small>Thorell, 1876</small> (12 species; Europe)
:* ''[[Granulosoma]]'' <small>Martens, 1973</small>
:* ''[[Granulosoma]]'' <small>Martens, 1973</small> (1 species)
:* ''[[Homalenotus]]'' <small>C.L.Koch, 1839</small>
:* ''[[Homalenotus]]'' <small>C.L.Koch, 1839</small> (13 species; Southern Europe and North Africa)
:* ''[[Mastobunus]]'' <small>Simon, 1879</small>
:* ''[[Mastobunus]]'' <small>Simon, 1879</small> (2 species; Southern Europe and North Africa)
:* ''[[Metasclerosoma]]'' <small>Roewer, 1912</small>
:* ''[[Metasclerosoma]]'' <small>Roewer, 1912</small> (5 species; Italy)
:* ''[[Protolophus]]'' <small>Banks, 1893</small>
:* ''[[Pseudastrobunus]]'' <small>Martens, 1973</small> (1 species)
:* ''[[Pseudastrobunus]]'' <small>Martens, 1973</small>
:* ''[[Pseudohomalenotus]]'' <small>Caporiacco, 1935</small> (1 species; the Karakoram)
:* ''[[Pseudohomalenotus]]'' <small>Caporiacco, 1935</small>
:* ''[[Pygobunus]]'' <small>Roewer, 1957</small> (2 species; Japan)
:* ''[[Pygobunus]]'' <small>Roewer, 1957</small>
:* ''[[Umbopilio]]'' <small>Roewer, 1956</small> (1 species)
:* ''[[Umbopilio]]'' <small>Roewer, 1956</small>


==Footnotes==
==Removed==
:* ''[[Caiza]]'' <small>Roewer, 1925</small>. Subjective synonym of ''Pectenobunus'' Roewer, 1910) in Tourinho, 2004a
{{reflist}}

:* ''[[Pseudoarthromerus]]'' <small>Karsch, 1891</small> (1 species; Japan, nomen nudum, therefore non-valid)


==References==
==References==
{{Reflist}}
* Joel Hallan's Biology Catalog: [http://insects.tamu.edu/research/collection/hallan/Acari/Family/Sclerosomatidae.txt Sclerosomatidae]


==External links==
[[Category:Harvestmen]]
* {{Commons category-inline}}
* {{Wikispecies-inline}}


{{Taxonbar|from=Q3179711}}
{{Opiliones-stub}}


[[Category:Harvestman families]]
[[es:Sclerosomatidae]]
[[fr:Sclerosomatidae]]
[[Category:Sclerosomatidae]]
[[no:Sclerosomatidae]]
[[pt:Sclerosomatidae]]

Latest revision as of 00:42, 26 April 2024

Sclerosomatidae
Temporal range: Jurassic–present
Hadrobunus grandis
Scientific classification
Kingdom:
Phylum:
Class:
Order:
Suborder:
Superfamily:
Family:
Sclerosomatidae

Simon, 1879
Subfamilies

Gagrellinae (117 genera)
Gyantinae (3 genera)
Leiobuninae (13 genera)
Sclerosomatinae (10 genera)

Diversity
4 subfamilies, c. 150 genera

The Sclerosomatidae are a family of harvestmen with about 1,300 known species.[1] One former subfamily has been recently removed to form a new family, Globipedidae.[2]

Name[edit]

The name is combined from Ancient Greek skleros "hard" and soma "body".[3]

Genera[edit]

  • Gyantinae Silhavý, 1946
  • Gyas Simon, 1879 (1 species; Alps)
  • Gyoides Martens, 1982 (6 species; Nepal)
  • Rongsharia Roewer, 1957 (3 species; Nepal)
  • Leiobuninae Banks, 1893
  • Amauropilio Mello-Leitão, 1937 (fossil) (2 species; fossils found in Colorado)
  • Cosmobunus Simon, 1879 (5 species; Spain and Mexico)
  • Dilophiocara Redikorzev, 1931 (2 species)
  • Eumesosoma Cokendolpher, 1980 (6 species; United States)
  • Eusclera Roewer, 1910 (2 species; China and India)
  • Hadrobunus Banks, 1900 (2 species; United States)
  • Leiobunum C.L.Koch, 1839 (126 species - 2 extinct fossils; worldwide)
  • Leuronychus Banks, 1900 (2 species; North America)
  • Microliobunum Roewer, 1912 (2 species; Middle East)
  • Micronelima Schenkel, 1938 (1 species; Spain)
  • Nelima Roewer, 1910 (44 species; worldwide)
  • Paranelima Caporiacco, 1938 (6 species; Mexico and Central America)
  • Schenkeliobunum Starega, 1964 (1 species)
  • Togwoteeus Roewer, 1952 (1 species; United States)
  • Sclerosomatinae Simon, 1879

Removed[edit]

  • Caiza Roewer, 1925. Subjective synonym of Pectenobunus Roewer, 1910) in Tourinho, 2004a

References[edit]

  1. ^ Sclerosomatidae Species List at Joel Hallan's Biology Catalog. Texas A&M University, archived from the original on 13 October 2014, retrieved 30 August 2015
  2. ^ Kury, A.B. & J.C. Cokendolpher (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 (Kury et al., ed.): 52-54.
  3. ^ Tourinho, Ana Lúcia (2007): Sclerosomatidae. Simon, 1879. In: Pinto-da-Rocha et al. 2007: 127ff
  4. ^ Chen, S. L.; Shih, H. T. (2018). "A review of the harvestman genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with a description of a new species from Taiwan". Zoological Studies. 57.

External links[edit]