Sclerosomatidae

Sclerosomatidae; Temporal range: Jurassic–present PreꞒ Ꞓ O S D C P T J K Pg N
	Hadrobunus grandis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Arachnida
Order:	Opiliones
Suborder:	Eupnoi
Superfamily:	Phalangioidea
Family:	Sclerosomatidae; Simon, 1879
Subfamilies
	Gagrellinae (117 genera); Gyantinae (3 genera); Leiobuninae (13 genera); Sclerosomatinae (10 genera)
Diversity
	4 subfamilies, c. 150 genera

The printable version is no longer supported and may have rendering errors. Please update your browser bookmarks and please use the default browser print function instead.

The Sclerosomatidae are a family of harvestmen with about 1,300 known species.^[1] One former subfamily has been recently removed to form a new family, Globipedidae.^[2]

Name

The name is combined from Ancient Greek skleros "hard" and soma "body".^[3]

Genera

Gagrellinae Thorell, 1889

Abaetetuba Tourinho-Davis, 2004 (5 species)
Adungrella Roewer, 1955 (3 species)
Aguilaia González-Sponga, 2003 (1 species; Venezuela)
Akalpia Roewer, 1915 (2 species; India and Japan)
Altobunus Roewer, 1910 (3 species; Celebes and Philippines)
Amazonesia Soares, 1970 (2 species)
Antigrella Roewer, 1954 (1 species)
Aurivilliola Roewer, 1910 (17 species; South and Southeast Asia)
Azucarella Roewer, 1959 (1 species)
Bakerinulus Roewer, 1955 (1 species)
Bastia Roewer, 1910 (3 species; South and Southeast Asia)
Bastioides Mello-Leitão, 1931 (1 species; South America)
Baturitia Roewer, 1931 (1 species; Sunda Islands)
Bellonia González-Sponga, 2003 (1 species; Venezuela)
Biceropsis Roewer, 1935 (1 species; Burma)
Bonthainia Roewer, 1913 (5 species; South and Southeast Asia)
Bullobunus Roewer, 1910 (7 species; Philippines)
Caluga Roewer, 1959 (1 species)
Cardenalia Kury, 2017 (1 species; Venezuela)
Carinobius Roewer, 1955 (1 species)
Carmenia Roewer, 1915 (1 species; Colombia)
Carmichaelus Roewer, 1929 (1 species; India)
Ceratobunellus Roewer, 1911 (3 species; India)
Ceratobunoides Roewer, 1923 (2 species; Sumatra)
Ceratobunus Thorell, 1889 (13 species)
Cervibunus Roewer, 1912 (3 Species; South and Southeast Asia)
Chasenella Roewer, 1955 (2 species; Borneo)
Chebabius Roewer, 1935 (1 species; Burma)
Coonoora Roewer, 1929 (1 species; India)
Dentobunus Roewer, 1910 (33 species; Southeast Asia)
Diangathia Roewer, 1955 (1 species)
Echinobunus Roewer, 1912 (1 species; Java)
Euceratobunus Roewer, 1923 (1 species; India)
Eugagrella Roewer, 1910 (25 species; South and Southeast Asia)
Euzaleptus Roewer, 1911 (4 species; South and Southeast Asia)
Fesa Roewer, 1953 (1 species)
Gagrella Stoliczka, 1869 (226 species; Asia)
Gagrellenna Roewer, 1929 (1 species; India)
Gagrellina Roewer, 1913 (1 species; Celebes)
Gagrellissa Roewer, 1931 (1 species; Sunda Islands)
Gagrellopsis Sato & Suzuki, 1939 (1 species; Japan)
Gagrellula Roewer, 1910 (67 species; Asia)
Garleppa Roewer, 1912 (4 species; South America)
Geaya Roewer, 1910 (82 species; Latin America)
Globulosoma Martens, 1987 (2 species; Nepal)
Guaranobunus Ringuelet, 1954 (1 species; Argentina)
Hamitergum Crawford, 1992 (1 species; Russia)
Harmanda Roewer, 1910 (15 species; Asia)
Harmandina Schenkel, 1954 (1 species)
Hehoa Roewer, 1929 (1 species; India)
Heterogagrella Roewer, 1954 (2 species; Southeast Asia)
Hexazaleptus Suzuki, 1966 (1 species)
Hexomma Thorell, 1876 (9 species; Southeast Asia)
Himaldroma Martens, 1987 (2 species; Nepal)
Himalzaleptus Martens, 1987 (1 species; Nepal)
Holcobunus Roewer, 1910 (26 or 27 species; Latin America)
Holmbergiana Mello-Leitão, 1931 (3 species; South America)
Hologagrella Roewer, 1910 (7 species; Southeast Asia)
Hypogrella Roewer, 1955 (1 species)
Hypsibunus Thorell, 1891 (11 species; India & Southeast Asia)
Jussara Mello-Leitão, 1935 (10 species; South America)
Koyamaia Suzuki, 1972 (1 species)
Krusa Goodnight & Goodnight, 1947 (10 species; Mexico)
Krusella Roewer, 1953 (1 species; Venezuela)
Liopagus Chamberlin, 1916 (1 species)
Marthana Thorell, 1891 (24 species; Southeast Asia)
Melanopa Thorell, 1889 (30 species; India, Southeast Asia)
Melanopella Roewer, 1931 (3 species; Sunda Islands)
Melanopula Roewer, 1929 (5 species; South and Southeast Asia)
Metadentobunus Roewer, 1915 (3 species; Taiwan)^[4]
Metahehoa Suzuki, 1985 (1 species)
Metasyleus Roewer, 1955 (3 species)
Metaverpulus Roewer, 1912 (7 species; South Asia)
Metazaleptus Roewer, 1912 (8 species; South and Southeast Asia)
Microzaleptus Roewer, 1955 (1 species)
Munequita Mello-Leitão, 1941 (1 species; Brazil)
Neogagrella Roewer, 1913 (3 species; South and Southeast Asia)
Nepalgrella Martens, 1987 (2 species; Nepal)
Nepalkanchia Martens, 1990 (2 species; Nepal)
Obigrella Roewer, 1955 (1 species)
Octozaleptus Suzuki, 1966 (1 species)
Onostemma Mello-Leitão, 1938 (2 species; Brazil)
Oobunus Kishida, 1930 (1 species)
Orissula Roewer, 1955 (1 species)
Padangrella Roewer, 1954 (1 species)
Palniella Roewer, 1929 (1 species; India)
Paradentobunus Roewer, 1915 (1 species; India)
Paragagrella Roewer, 1912 (5 species; South and Southeast Asia)
Paragagrellina Schenkel, 1963 (1 species)
Parageaya Mello-Leitão, 1933 (6 species; Latin America)
Paratamboicus Mello-Leitão, 1940 (30 species; South America)
Paraumbogrella Suzuki, 1963 (1 species)
Paruleptes Soares, 1970 (1 species)
Pectenobunus Roewer, 1910 (4 species; South America)
Pergagrella Roewer, 1954 (1 species)
Pokhara Suzuki, 1970 (7 species; Nepal)
Prionostemma Pocock, 1903 (114 species; Mexico, Central America, and South America)
Prodentobunus Roewer, 1923 (4 species; Asia)
Psammogeaya Mello-Leitão, 1946 (1 species; Uruguay)
Psathyropus L. Koch, 1878 (40 species; Asia)
Pseudarthromerus Karsch, 1892 (1 species)
Pseudogagrella Redikortsev, 1936 (13 species; East Asia)
Pseudomelanopa Suzuki, 1974 (2 species; East Asia)
Pseudosystenocentrus Suzuki, 1985 (1 species)
Romerella Goodnight & Goodnight, 1943 (5 species; Mexico and South America)
Sarasinia Roewer, 1913 (1 species; Celebes)
Sataria Roewer, 1915 (3 species; India)
Scotomenia Thorell, 1889 (1 species; Burma)
Sericicorpus Martens, 1987 (1 species; Nepal)
Sinadroma Roewer, 1955 (1 species)
Syleus Thorell, 1876 (2 species; India)
Syngagrella Roewer, 1913 (1 species; Celebes)
Systenocentrus Simon, 1886 (6 species; Southeast and East Asia)
Tamboicus Roewer, 1912 (3 species; South America)
Taperina Roewer, 1953 (2 species)
Tetraceratobunus Roewer, 1915 (3 species; South Asia)
Thunbergia Martens, 2020 (4 species; South America)
Toragrella Roewer, 1955 (2 species)
Trachyrhinus Weed, 1892 (7 species; North America: Mexico and United States)
Umbogrella Roewer, 1955 (1 species)
Umbopilio Roewer, 1956 (2 species)
Vaho González-Sponga, 2003 (1 species)
Valle González-Sponga, 2003 (3 species)
Varinodulia Canals, 1935 (1 species)
Verpulus Simon, 1901 (13 species; South and Southeast Asia)
Verrucobunus Roewer, 1931 (3 species; East and Southeast Asia)
Xerogrella Martens, 1987 (1 species; Nepal)
Zaleptiolus Roewer, 1955 (4 species; Asia)
Zaleptulus Roewer, 1955 (3 species)
Zaleptus Thorell, 1876 (72 species; Asia and Australia)

Gyantinae Silhavý, 1946

Gyas Simon, 1879 (1 species; Alps)
Gyoides Martens, 1982 (6 species; Nepal)
Rongsharia Roewer, 1957 (3 species; Nepal)

Leiobuninae Banks, 1893

†Amauropilio Mello-Leitão, 1937 (fossil) (2 species; fossils found in Colorado)
Cosmobunus Simon, 1879 (5 species; Spain and Mexico)
Dilophiocara Redikorzev, 1931 (2 species)
Eumesosoma Cokendolpher, 1980 (6 species; United States)
Eusclera Roewer, 1910 (2 species; China and India)
Hadrobunus Banks, 1900 (2 species; United States)
Leiobunum C.L.Koch, 1839 (126 species - 2 extinct fossils; worldwide)
Leuronychus Banks, 1900 (2 species; North America)
Microliobunum Roewer, 1912 (2 species; Middle East)
Micronelima Schenkel, 1938 (1 species; Spain)
Nelima Roewer, 1910 (44 species; worldwide)
Paranelima Caporiacco, 1938 (6 species; Mexico and Central America)
Schenkeliobunum Starega, 1964 (1 species)
Togwoteeus Roewer, 1952 (1 species; United States)

Sclerosomatinae Simon, 1879

Astrobunus Thorell, 1876 (12 species; Europe)
Granulosoma Martens, 1973 (1 species)
Homalenotus C.L.Koch, 1839 (13 species; Southern Europe and North Africa)
Mastobunus Simon, 1879 (2 species; Southern Europe and North Africa)
Metasclerosoma Roewer, 1912 (5 species; Italy)
Pseudastrobunus Martens, 1973 (1 species)
Pseudohomalenotus Caporiacco, 1935 (1 species; the Karakoram)
Pygobunus Roewer, 1957 (2 species; Japan)
Umbopilio Roewer, 1956 (1 species)

Removed

Caiza Roewer, 1925. Subjective synonym of Pectenobunus Roewer, 1910) in Tourinho, 2004a

Pseudoarthromerus Karsch, 1891 (1 species; Japan, nomen nudum, therefore non-valid)

References

^ Sclerosomatidae Species List at Joel Hallan's Biology Catalog. Texas A&M University, archived from the original on 13 October 2014, retrieved 30 August 2015
^ Kury, A.B. & J.C. Cokendolpher (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 (Kury et al., ed.): 52-54.
^ Tourinho, Ana Lúcia (2007): Sclerosomatidae. Simon, 1879. In: Pinto-da-Rocha et al. 2007: 127ff
^ Chen, S. L.; Shih, H. T. (2018). "A review of the harvestman genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with a description of a new species from Taiwan". Zoological Studies. 57.

External links

Media related to Sclerosomatidae at Wikimedia Commons
Data related to Sclerosomatidae at Wikispecies

[1] Sclerosomatidae Species List at Joel Hallan's Biology Catalog. Texas A&M University, archived from the original on 13 October 2014, retrieved 30 August 2015

[2] Kury, A.B. & J.C. Cokendolpher (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 (Kury et al., ed.): 52-54.

[pinto-127ff-3] Tourinho, Ana Lúcia (2007): Sclerosomatidae. Simon, 1879. In: Pinto-da-Rocha et al. 2007: 127ff

[Chen-4] Chen, S. L.; Shih, H. T. (2018). "A review of the harvestman genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with a description of a new species from Taiwan". Zoological Studies. 57.

[1]

[2]

[3]

[4]