Pentanchidae
Pentanchidae | ||||||||||||
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Spotted shark ( Galeus melastomus ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Pentanchidae | ||||||||||||
Smith , 1912 |
The Pentanchidae are a species-rich family of small sharks , currently about 90 described species, of the order of the ground sharks (Carcharhiniformes). The family was established in 1912 by the American ichthyologist Hugh McCormick Smith with the first description of the single-fin cat shark ( Pentanchus profundicolus ) and assigned to the Hexanchiformes because of the lack of the first dorsal fin . His British colleague Charles Tate Regan assigned Pentanchus profundicolus to the cat sharks (Scyliorhinidae) in the same year , making the Pentanchidae family no longer valid.
In 1988 the shark expert Leonard Compagno divided the cat sharks into two subfamilies, the Pentanchinae and the Scyliorhininae, based on the skull morphology, but abandoned this subdivision in later publications. However, both morphological and studies based on DNA analyzes showed that the cat sharks are not a monophylum. In 2005 the Pentanchidae were therefore revalidated again by Iglésias, Lecointre and Sellos.
features
The species of the Pentanchidae are slender sharks which, depending on the species, can reach a length of 16 cm ( Parmaturus campechiensis ) to 90 cm ( Apristurus exsanguis , Asymbolus analis and Galeus melastomus ). Most species are about half a meter long. Like cat sharks, they have two dorsal fins (with the exception of Pentanchus profundicolus ) and two anal fins. In the Pentanchidae and the cat sharks, however, in contrast to all other shark families, it is far behind, at the level of the pelvic fins or just behind. A spray hole as well as a spiral casing are available. The only morphological feature that distinguishes the Pentanchidae from the cat sharks is the lack of a supraorbital ridge, a cartilage ridge above the eyes.
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Genera and species
There are 8 genera and around 90 species:
- Genus: Ghost cat sharks ( Apristurus Garman, 1913 )
- Flat-nosed dogfish ( Apristurus acanutus Chu, Meng & Li, 1985 ) (possibly synonymous with Apristurus platyrhynchus )
- Whitish dogfish ( Apristurus albisoma Nakaya & Seret, 1999 )
- Rough-skin dogfish ( Apristurus ampliceps Sasahara, Sato & Nakaya, 2008 )
- Bright deep-water cat shark ( Apristurus aphyodes Nakaya & Stehmann, 1998 )
- Pinocchio dogfish ( Apristurus australis Sato, Nakaya & Yorozu, 2008 )
- Apristurus breviventralis Kawauchi, Weigmann & Nakaya, 2014
- Apristurus brunneus (Gilbert, 1892)
- Big head dogfish ( Apristurus bucephalus White, Last & Pogonoski, 2008 )
- Gray dogfish ( Apristurus canutus Springer & Heemstra, 1979 )
- Bleicher dogfish ( Apristurus exsanguis Sato, Nakaya & Stewart, 1999 )
- Dicker dogfish ( Apristurus fedorovi Dolganov, 1983 )
- Apristurus garricki Sato, Stewart & Nakaya, 2013
- Humpback cat shark ( Apristurus gibbosus Meng, Chu & Li, 1985 )
- Long-finned dogfish ( Apristurus herklotsi (Fowler, 1934) )
- Small-bellied dogfish ( Apristurus indicus (Brauer, 1906) )
- Apristurus internatus Deng, Xiong & Zhan, 1988
- Broad-nosed cat shark ( Apristurus investigatoris (Misra, 1962) )
- Japanese dogfish ( Apristurus japonicus Nakaya, 1975 )
- Long-nosed dogfish ( Apristurus kampae Taylor, 1972 )
- Icelandic deepwater cat shark ( Apristurus laurussonii (Saemundsson, 1922) )
- Long-headed dogfish ( Apristurus longicephalus Nakaya, 1975 )
- Flat-headed dogfish ( Apristurus macrorhynchus (Tanaka, 1909) )
- Breitmaulkatzenhai ( Apristurus macrostomus Chu, Meng Li & 1985 )
- Ghost deepwater cat shark ( Apristurus manis (Springer, 1966) )
- Meat nose dogfish ( Apristurus melanoasper Iglésias, Nakaya & Stehmann, 2004 )
- Small-eyed deepwater cat shark ( Apristurus microps (Gilchrist, 1922) )
- Small- backed cat shark ( Apristurus micropterygeus Meng, Chu & Li, 1986 )
- Apristurus nakayai Iglésias, 2013
- Big-nosed deepwater cat shark ( Apristurus nasutus de Buen, 1959 )
- Smallfin cat shark ( Apristurus parvipinnis Springer & Heemstra, 1979 )
- Fat dogfish ( Apristurus pinguis Deng, Xiong & Zhan, 1983 )
- Spatula-nosed cat shark ( Apristurus platyrhynchus (Tanaka, 1909) )
- Deepwater cat shark ( Apristurus profundorum (Goode & Bean, 1896) )
- Broad-gilled catshark ( Apristurus riveri Bigelow & Schroeder, 1944 )
- Saldanha dogfish ( Apristurus saldanha (Barnard, 1925) )
- Pale dogfish ( Apristurus sibogae (Weber, 1913) )
- Chinese dogfish ( Apristurus sinensis Chu & Hu, 1981 )
- Squarepants cat shark ( Apristurus spongiceps (Gilbert, 1905) )
- Panama ghost cat shark ( Apristurus stenseni (Springer, 1979) )
- Apristurus yangi White et al., 2017
- Genus: Australian spotted dogfish ( Asymbolus Whitley, 1939 )
- Australian spotted dogfish ( Asymbolus analis (Ogilby, 1885) )
- Spotted dogfish ( Asymbolus funebris Compagno, Stevens & Last, 1999 )
- Asymbolus galacticus Séret & Last, 2008
- Western spotted dogfish ( Asymbolus occiduus Last, Gomon & Gledhill, 1999 )
- Light-spotted dogfish ( Asymbolus pallidus Last, Gomon & Gledhill, 1999 )
- Asymbolus parvus Compagno, Stevens & Last, 1999
- Asymbolus rubiginosus Last, Gomon & Gledhill, 1999
- Modified dogfish ( Asymbolus submaculatus Compagno, Stevens & Last, 1999 )
- Gulf dogfish ( Asymbolus vincenti ( Zietz , 1908) )
- Genus: Tadpole cat sharks ( Cephalurus Bigelow & Schroeder, 1941 )
- Tadpole dogfish ( Cephalurus cephalus (Gilbert, 1892) )
- Southern tadpole dogfish ( Cephalurus sp.A ; not yet described)
- Genus: Sawtail cat sharks ( Galeus Rafinesque, 1810 )
- Antilles dogfish ( Galeus antillensis Springer, 1979 )
- Rough-tailed dogfish ( Galeus arae (Nichols, 1927) )
- Atlantic sawtail shark ( Galeus atlanticus (Vaillant, 1888) )
- Long-fin sawtail dogfish ( Galeus cadenati Springer, 1966 )
- Galeus corriganae White, Mana & Naylor, 2016
- Gecko dogfish ( Galeus eastmani (Jordan & Snyder, 1904) )
- Slender sawtail dogfish ( Galeus gracilis Compagno & Stevens, 1993 )
- Long-nosed catfish ( Galeus longirostris Tachikawa & Taniuchi, 1987 )
- Spotted shark ( Galeus melastomus Rafinesque, 1810 )
- Southern sawtail dogfish ( Galeus mincaronei Soto, 2001 )
- Mouse tailed shark ( Galeus murinus (Collett, 1904) )
- Broadfin sawtail dogfish ( Galeus nipponensis Nakaya, 1975 )
- Pepper dogfish ( Galeus piperatus Springer & Wagner, 1966 )
- African sawtail dogfish ( Galeus polli Cadenat, 1959 )
- Galeus priapus Séret & Last, 2008
- Blacktip sawtail dogfish ( Galeus sauteri (Jordan & Richardson, 1909) )
- Dwarf sawtail dogfish ( Galeus schultzi Springer, 1979 )
- Springer's sawtail dogfish ( Galeus springeri Konstantinou & Cozzi, 1998 )
- Genus: Tiger cat sharks ( Halaelurus Gill, 1862 )
- Speckled dogfish ( Halaelurus boesemani Springer & D'Aubrey, 1972 )
- Black-spotted dogfish ( Halaelurus buergeri (Müller & Henle, 1838) )
- Banded dogfish ( Halaelurus lineatus Bass, D'Aubrey & Kistnasamy, 1975 )
- Halaelurus maculosus White, Last & Stevens, 2007
- Tiger dogfish ( Halaelurus natalensis (Regan, 1904) )
- Quagga dogfish ( Halaelurus quagga (Alcock, 1899) )
- Rust- colored dogfish ( Halaelurus sellus White, Last & Stevens, 2007 )
- Genus: Izak cat sharks ( Holohalaelurus Fowler, 1934 )
- Holohalaelurus favus Human, 2006
- Holohalaelurus grennian Human, 2006
- Tropical izak dogfish ( Holohalaelurus melanostigma (Norman, 1939) )
- African spotted dogfish ( Holohalaelurus punctatus (Gilchrist, 1914) )
- Izak dogfish ( Holohalaelurus regani (Gilchrist, 1922) )
- Genus: Hedge-tailed cat sharks ( Parmaturus Fowler, 1934 )
- Whitetip dogfish ( Parmaturus albimarginatus Séret & Last, 2007 )
- White penis dogfish ( Parmaturus albipenis Séret & Last, 2007 )
- Parmaturus angelae Araújo Soares et al., 2019
- Beige dogfish ( Parmaturus bigus Séret & Last, 2007 )
- Campeche dogfish ( Parmaturus campechiensis Springer, 1979 )
- Velvet dogfish ( Parmaturus lanatus Séret & Last, 2007 )
- McMillan's dogfish ( Parmaturus macmillani Hardy, 1985 )
- Black- gill dogfish ( Parmaturus melanobranchus (Chan, 1966) )
- Parmaturus nigripalatum Fahmi & Ebert, 2018
- Salamander dogfish ( Parmaturus pilosus Garman, 1906 )
- Hedge- tailed dogfish ( Parmaturus xaniurus (Gilbert, 1892) )
- Genus: Pentanchus Smith & Radcliffe in Smith, 1912 ( type genus )
- Single fin dogfish ( Pentanchus profundicolus Smith & Radcliffe, 1912 )
literature
- Samuel P. Iglésias, Guillaume Lecointre, Daniel Y. Sellos: Extensive paraphylies within sharks of the order Carcharhiniformes inferred from nuclear and mitochondrial genes. Molecular Phylogenetics and Evolution 34 (2005) 569-583, PDF
Individual evidence
- ^ Catalog of Fishes: Species by Family / Subfamily
- ^ HM Smith, 1912: Description of a new notidanoid shark from the Philippine Islands representing a new family. Scientific results of the Philippine cruise of the Fisheries steamer "Albatross," 1907-10 .-- No. 14.]. Proceedings of the United States National Museum, 41 (1872): 489-491, Pl. 42. online
- ↑ JG Maisey, 1984: Higher elasmobranch phylogeny and biostratigraphy. Zool. J. Linn. Soc. 82, 33-54
- ↑ CJ Winchell, AP Martin, J. Mallatt, 2004: Phylogeny of elasmobranchs based on LSU and SSU ribosomal RNA genes. Molecular Phylogenetics and Evolution 31 (2004), pp. 214–224, PDF ( Memento of the original from September 24, 2015 in the Internet Archive ) Info: The archive link was inserted automatically and has not yet been checked. Please check the original and archive link according to the instructions and then remove this notice.
- ↑ a b c d S. P. Iglésias, G. Lecointre & DY Sellos, 2005: Extensive paraphylies within sharks of the order Carcharhiniformes inferred from nuclear and mitochondrial genes. Mol. Phylogenet. Evol., 34: 569-583.
- ↑ Parmaturus campechiensis on Fishbase.org (English)
- ↑ Apristurus exsanguis on Fishbase.org (English)
- ↑ Asymbolus analis on Fishbase.org (English)
- ↑ Galeus melastomus on Fishbase.org (English)
- ^ Joseph S. Nelson : Fishes of the World. 4th edition. John Wiley & Sons , Hoboken NJ 2006, ISBN 0-471-25031-7 .
Web links
- Family Pentanchidae on Fishbase.org (English)