Cat sharks
Cat sharks | ||||||||||||
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Great spotted dogfish ( Scyliorhinus stellaris ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Scyliorhinidae | ||||||||||||
Gill , 1862 |
The cat sharks (Scyliorhinidae) are a large family of the ground sharks (Carcharhiniformes), it consists of ten genera with 70 species that occur worldwide in the cool to warm seas. They are only missing in the polar seas. They also occur in all water depths from shallow water to deep sea.
They owe their German name to large eyes , which with their elongated pupils are reminiscent of the eyes of cats and have a nictitating skin (eyes are closed with the lower eyelid). What all species have in common is that they lay eggs .
features
Cat sharks have a clearly elongated body and have two stingless dorsal fins ( fins ) and two anal fins . In contrast to all other shark families, with the exception of the Pentanchidae , a shark family that was recently split off from the cat sharks, the first dorsal fin is at the level of the pelvic fins or just behind it. The last of the five gill slits lies above the base of the pectoral fin. A spray hole is available, as is a spiral casing . Cat sharks are 21 centimeters to 1.70 meters long, depending on the species. The only morphological feature that distinguishes the cat sharks from the Pentanchidae is the supraorbital ridge, a cartilage ridge above the eyes that the Pentanchidae lack.
A number of cat sharks are strikingly colored. The coral dogfish ( Atelomycterus marmoratus ) is brown in color with darker areas and white spots. Others have dark vertical stripes, like the pajama shark ( Poroderma africanum ), or a chain-like, dark pattern on the lighter skin, like the chain cat shark ( Scyliorhinus retifer ). Many cat sharks also show biofluorescence excited by incident light , similar to bioluminescence . There are current film recordings that show biofluorescence (distributed over the whole body of a swelling shark ), detectable only with special light.
Cat sharks feed on larger invertebrates and small fish.
Systematics
Within the basic sharks (Carcharhiniformes) the cat sharks occupy a basal position as sister group of all other families. With originally well over 100 species, they were the most species-rich family of sharks. After the separation of the Pentanchidae , ten genera and around 70 species are still counted among the cat sharks:
The following cladogram shows the basal position of the cat sharks within the basic sharks.
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- Subfamily Cephaloscylliinae Fowler 1947
- Genus: Swell sharks ( Cephaloscyllium Gill, 1862 )
- Cephaloscyllium albipinnum Last, Motomura & White, 2008
- Spotted swell shark ( Cephaloscyllium circulopullum Yano, Ahmad & Gambang, 2005) (possibly synonym of Cephaloscyllium sarawakensis )
- Cephaloscyllium cooki Last, Séret & White, 2008
- Net swell shark ( Cephaloscyllium fasciatum Chan, 1966 )
- Cephaloscyllium hiscosellum White & Ebert, 2008
- Lady shark ( Cephaloscyllium isabellum (Bonnaterre, 1788) )
- Australian swell shark ( Cephaloscyllium laticeps (Duméril, 1853) )
- Cephaloscyllium parvum Inoue & Nakaya, 2006 (possibly synonymous with Cephaloscyllium sarawakensis )
- Cephaloscyllium pictum Last, Séret & White, 2008
- Sarawak swell shark ( Cephaloscyllium sarawakensis Yano & Gambang, 2005 )
- Cephaloscyllium signourum Last, Séret & White, 2008
- Indian swell shark ( Cephaloscyllium silasi (Talwar, 1974) )
- Cephaloscyllium speccum Last, Séret & White, 2008
- Cephaloscyllium stevensi Clark & Randall, 2011
- Ballonhai ( Cephaloscyllium sufflans (Regan, 1921) )
- Spotted swell shark ( Cephaloscyllium umbratile Jordan & Fowler, 1903 )
- Cephaloscyllium variegatum Last & White, 2008
- Swell shark ( Cephaloscyllium ventriosum (Garman, 1880) )
- Cephaloscyllium zebrum Last & White, 2008
- Genus: Swell sharks ( Cephaloscyllium Gill, 1862 )
- Subfamily Schroederichthyinae Compagno, 1988
- Genre : Akheilos White, Fahmi & Weigmann, 2019
- Akheilos suwartanai White, Fahmi & Weigmann, 2019
- Genus: narrow-tailed cat sharks ( Schroederichthys Springer, 1966 )
- Narrow-mouth dogfish ( Schroederichthys bivius (Müller & Henle, 1838) )
- Red-spotted dogfish ( Schroederichthys chilensis (Guichenot, 1848) )
- Narrow-tailed dogfish ( Schroederichthys maculatus Springer, 1966 )
- Lizard dogfish ( Schroederichthys saurisqualus Soto, 2001 )
- Slender dogfish ( Schroederichthys tenuis Springer, 1966 )
- Genre : Akheilos White, Fahmi & Weigmann, 2019
- Subfamily Scyliorhininae Gill, 1862
- Genus: Coral cat sharks ( Atelomycterus Garman, 1913 )
- Bali dogfish ( Atelomycterus baliensis (White, Last & Dharmadi, 2005) )
- Atelomycterus erdmanni Fahmi & White, 2015
- Striped sand dogfish ( Atelomycterus fasciatus (Compagno & Stevens, 1993) )
- Australian marble dogfish ( Atelomycterus macleayi (Whitley, 1939) )
- Coral dogfish ( Atelomycterus marmoratus (Anonymous [Bennett], 1830) )
- Eastern striped dogfish ( Atelomycterus marnkalha (Jacobsen & Bennett, 2007) )
- Genus: Black-spotted cat sharks ( Aulohalaelurus Fowler, 1934 )
- New Caledonian dogfish ( Aulohalaelurus kanakorum (Séret, 1990) )
- Australian black-spotted dogfish ( Aulohalaelurus labiosus (Waite, 1905) )
- Genus: Bythaelurus Compagno, 1988
- Arab dogfish ( Bythaelurus alcocki (Garman, 1913) )
- Bythaelurus bachi Weigmann, Ebert, Clerkin, Stehmann & Naylor, 2016
- Black dogfish ( Bythaelurus canescens (Günther, 1878) )
- Wide head dogfish ( Bythaelurus clevai (SERET, 1987) )
- New Zealand dogfish ( Bythaelurus dawsoni (Springer, 1971) )
- Bythaelurus giddingsi McCosker, Long & Baldwin, 2012
- Spiny dogfish ( Bythaelurus hispidus (Alcock, 1891) )
- Spotless dogfish ( Bythaelurus immaculatus (Chu & Meng, 1982) )
- Bythaelurus incanus Last & Stevens, 2008
- Morass dogfish ( Bythaelurus lutarius (Springer & D'Aubrey, 1972) )
- Bythaelurus naylori Ebert & Clerkin, 2015
- Bythaelurus stewarti Weigmann, Kaschner & Thiel, 2018
- Bythaelurus tenuicephalus Kaschner, Weigmann & Thiel, 2015
- Bythaelurus vivaldii Weigmann & Kaschner, 2017
- Genus: Figaro Whitley, 1928
- Australian Sägeschwanz catshark ( Figaro boardmani (Whitley, 1928) )
- Northern Sawtail shark ( Figaro striatus Gledhill, Last & White, 2008 )
- Genus: shy sharks ( Haploblepharus Garman, 1913 )
- Puff adder dogfish ( Haploblepharus edwardsii (Schinz, 1822) )
- Brown dogfish ( Haploblepharus fuscus Smith, 1950 )
- Natal dogfish ( Haploblepharus kistnasamyi Human & Compagno, 2006 )
- Dark dogfish ( Haploblepharus pictus (Müller & Henle, 1838) )
- Genus: Bartel cat sharks ( Poroderma Smith, 1838 )
- Pajama shark ( Poroderma africanum (Gmelin, 1789) )
- Leopard dogfish ( Poroderma pantherinum (Müller & Henle, 1838) )
- Genus: Spotted cat sharks ( Scyliorhinus Blainville, 1816 )
- Boa dogfish ( Scyliorhinus boa (Goode & Bean, 1896) )
- Scyliorhinus cabofriensis Soares, Gomes & De Carvalho, 2016
- Small-spotted dogfish ( Scyliorhinus canicula (Linnaeus, 1758) )
- Yellow-spotted dogfish ( Scyliorhinus capensis (Müller & Henle, 1838) )
- West African dogfish ( Scyliorhinus cervigoni (Maurin & Bonnet, 1970) )
- Scyliorhinus comoroensis Compagno, 1988
- Scyliorhinus duhamelii (Garman, 1913)
- Brown-spotted dogfish ( Scyliorhinus garmani (Fowler, 1934) )
- Spotted dogfish ( Scyliorhinus haeckelii (Miranda Ribeiro, 1907) )
- White-backed dogfish ( Scyliorhinus hesperius (Springer, 1966) )
- Stained dogfish ( Scyliorhinus meadi (Springer, 1966) )
- Chain cat shark ( Scyliorhinus retifer (Garman, 1881) )
- Great-spotted dogfish ( Scyliorhinus stellaris (Linnaeus, 1758) )
- Scyliorhinus torazame (Tanaka, 1908)
- Dwarf dogfish ( Scyliorhinus torrei (Fowler, 1941) )
- Scyliorhinus ugoi Soares, Gomes & Gadig, 2015
- Genus: Coral cat sharks ( Atelomycterus Garman, 1913 )
literature
- Alfred Kaestner: Textbook of special zoology. Volume 2: Dietrich Starck: Vertebrates. Part 2: Kurt Fiedler: Fish. Gustav Fischer, Jena 1991, ISBN 3-334-00338-8 .
- Joseph S. Nelson : Fishes of the World. 4th edition. John Wiley & Sons, Hoboken NJ 2006, ISBN 0-471-25031-7 .
- Brett A. Humana, E. Patricia Owena, Leonard JV Compagnob , Eric H. Harleya: Testing morphologically based phylogenetic theories within the cartilaginous fishes with molecular data, with special reference to the catshark family (Chondrichthyes; Scyliorhinidae) and the interrelationships within them. In: Molecular Phylogenetics and Evolution. Volume 39, Issue 2, May 2006, ISSN 1055-7903 , pages 384-391, doi : 10.1016 / j.ympev.2005.09.009 .
Individual evidence
- ↑ WoRMS taxon details Scyliorhinidae Gill, 1862
- ^ Catalog of Fishes: Species by Family / Subfamily
- ↑ a b c S. P. Iglésias, G. Lecointre & DY Sellos, 2005: Extensive paraphylies within sharks of the order Carcharhiniformes inferred from nuclear and mitochondrial genes. Mol. Phylogenet. Evol., 34: 569-583.
- ↑ David F. Gruber, Ellis R. Loew, Dimitri D. Deheyn, Derya Akkaynak, Jean P. Gaffney, W. Leo Smith, Matthew P. Davis, Jennifer H. Stern, Vincent A. Pieribone, John S. Sparks: Biofluorescence in catsharks (Scyliorhinidae): Fundamental description and relevance for Elasmobranch visual ecology. In: Sci. Rep. Volume 6, 2016, p. 24751, doi : 10.1038 / srep24751 .
- ↑ TV report on ServusTV on May 25, 2016.
- ^ PR Last, H. Motomura, WT White: Cephaloscyllium albipinnum sp. nov., a new swell shark (Carcharhihiformes: Scyliorhinidae) from Southeastern Australia. In: PR Last WT White & JJ Pogonoski (Eds.): Descriptions of New Australian Chondrichthyans. In: CSIRO Marine and Atmospheric Research Paper. Volume 22, 2008.
- ^ PR Last, B. Séret, WT White: New swellsharks (Cephaloscyllium: Scyliorhinidae) from the Indo-Australian region. In: PR Last WT White & JJ Pogonoski (Eds.): Descriptions of New Australian Chondrichthyans. In: CSIRO Marine and Atmospheric Research Paper. Volume 22, 2008.
- ^ William T. White, Fahmi Fahmi and Simon Weigmann. 2019. A New Genus and Species of Catshark (Carcharhiniformes: Scyliorhinidae) from eastern Indonesia. Zootaxa. 4691 (5); 444-460. DOI: 10.11646 / zootaxa.4691.5.2
- ↑ Karla DA Soares & Marcelo R. De Carvalho (2019): The catshark genus Scyliorhinus (Chondrichthyes: Carcharhiniformes: Scyliorhinidae): taxonomy, morphology and distribution. Zootaxa , 4601 (1): 1-147. DOI: 10.11646 / zootaxa.4601.1.1
- ↑ Karla DA Soares, Ulisses L. Gomes & Marcelo R. De Carvalho: Taxonomic review of catsharks of the Scyliorhinus haeckelii group, with the description of a new species (Chondrichthyes: Carcharhiniformes: Scyliorhinidae). Zootaxa, 4066, 5, 501-534, January 2016 doi : 10.11646 / zootaxa.4066.5.1
Web links
- Cat sharks on Fishbase.org (English)
- Weinheimer, M. 2004, Animal Diversity Web. Scyliorhinidae