Systematics of the bumblebees
The systematics of the bumblebees includes the relationships and the taxonomy of the bumblebees ( bombus ). The bumblebee species occurring worldwide are listed. The list follows the current phylogenetic knowledge .
Within the bees, the bumblebees belong to the group of cup collectors , a monophyletic group to which the honeybees also belong. The phylogenetic structure within the group has not yet been satisfactorily clarified, so according to the current state of research it cannot be said which is the sister group of the bumblebees.
The bumblebees themselves are divided into several sub-genera, but it is difficult to differentiate them from one another because the bumblebees have a very uniform body structure. The cuckoo bumblebees of the subgenus Psithyrus have been split off as a separate genus by many authors . Phylogenetic studies indicate, however, that deviations in the body structure of the cuckoo bumblebees can only be interpreted as adaptations to the way of life.
Subgenus Alpigenobombus
- Bombus angustus
- Bombus breviceps
- Genalis bomb
- Bombus grahami
- Bombus kashmirensis
- Bombus nobilis
- Bombus wurflenii - mountain forest bumblebee
Subgenus Alpinobombus
- Bombus alpinus - Alpine bumblebee
- Bombus balteatus -
- Bombus hyperboreus
- Bombus neoboreus
- Bombus polaris
Subgenus bombias
Subgenus Bombus sensu stricto
- Bombus affinis
- Bombus cryptarum
- Bombus franklini (very rare or even extinct)
- Bombus hypocrita
- Bombus ignitus
- Bombus jacobsoni
- Bombus lantschouensis
- Bombus longipennis
- Bombus lucorum - light yellow bumblebee
- Bombus magnus - Large earth bumblebee
- Bombus minshanicus
- Bombus occidentalis
- Bombus patagiatus
- Bombus sporadicus
- Bombus terrestris - Dark Earth Bumblebee
- Bombus terricola
- Bombus tunicatus
Subgenus Cullumanobombus
- Bombus baeri
- Bombus brachycephalus
- Bombus coccineus
- Bombus crotchii
- Bombus cullumanus - Cullumanushummel
- Bombus ecuadorius
- Bombus fraternus
- Bombus funebris
- Bombus griseocollis
- Bombus handlirschi
- Bombus haueri
- Bombus hortulanus
- Bombus macgregori
- Bombus melaleucus
- Bombus morrisoni
- Bombus robustus
- Bombus rohweri
- Bombus rubicundus
- Bombus rufocinctus
- Bombus semenoviellus
- Bombus tucumanus
- Bombus unicus
- Bombus vogti
Subgenus callobombus
- Bombus soroeensis - thistle bumblebee
Subgenus Megabombus
- Bombus argillaceus - clay bumblebee
- Bombus bicoloratus
- Bombus consobrinus
- Bombus czerskii
- Bombus diversus
- Bombus gerstaeckeri
- Bombus hortorum - garden bumblebee
- Bombus irisanensis
- Bombus koreanus
- Bombus longipes
- Bombus melanopoda (very rare and possibly extinct)
- Bombus portchinsky
- Bombus religiosus
- Bombus ruderatus - field bumblebee
- Bombus saltuarius
- Bombus securus
- Bombus senex
- Bombus supremus
- Bombus sushkini
- Bombus tichenkoi
- Bombus trifasciatus
- Bombus ussurensis
Subgenus melanobombus
- Bombus eximius
- Bombus festivus
- Bombus formosellus
- Bombus friseanus
- Bombus incertus
- Bombus keriensis
- Bombus ladakhensis
- Bombus lapidarius - stone bumblebee
- Bombus miniatus
- Bombus pyrosoma
- Bombus richardsiellus
- Bombus rufipes
- Bombus rufofasciatus
- Bombus semenovianus
- Bombus sichelii
- Bombus simillimus
- Bombus tanguticus
Subgenus Mendacibombus
- Bombus avinoviellus
- Bombus convexus
- Bombus handlirschianus
- Bombus himalayanus
- Bombus makarjini
- Bombus margreiteri
- Bombus marussinus
- Bombus mendax - Trughummel
- Bombus superbus
- Bombus turkestanicus
- Bombus waltoni
Subgenus Orientalibombus
Subgenus Psithyrus - cuckoo bumblebees
For a long time, the cuckoo bumblebee was viewed by most authors as an independent genus. Phylogenetic studies in the 1990s showed, however, that cuckoo bumblebees and bumblebees are not in a sister group relationship to one another, the genus Bombus is paraphyletic after splitting off from Psithyrus . Differences in the body structure of the cuckoo bumblebees are largely just an adaptation to the different way of life, but not an indication of a different origin. One solution would be to split the genus Bombus into further genres, which most experts reject because of the pronounced uniformity of the group. According to the currently accepted status of the system, Psithyrus is therefore only regarded as a subgenus of Bombus .
- Bombus ashtoni
- Bombus barbutellus - Bearded Cuckoo Bumblebee
- Bombus bellardii
- Bombus bohemicus
- Bombus branickii
- Bombus campestris - field cuckoo bumblebee
- Bombus chinensis
- Bombus citrinus
- Bombus coreanus
- Bombus cornutus
- Bombus expolitus
- Bombus ferganicus
- Bombus fernaldae
- Bombus flavidus
- Bombus insularis
- Bombus maxillosus
- Bombus monozonus
- Bombus morawitzianus
- Bombus norvegicus
- Bombus novus
- Bombus perezi
- Bombus quadricolor - four-colored cuckoo bumblebee
- Bombus rupestris - rock cuckoo bumblebee
- Bombus skorikovi
- Bombus suckleyi
- Bombus sylvestris - forest cuckoo bumblebee
- Bombus tibetanus
- Bombus turneri
- Bombus variabilis
- Bombus vestalis
Subgenus Pyrobombus
- Bombus abnormal
- Bombus ardens
- Bombus avanus
- Bombus beaticola
- Bombus bifarius
- Bombus bimaculatus
- Bombus biroi
- Bombus brodmannicus
- Bombus caliginosus
- Bombus centralis
- Bombus cingulatus
- Bombus cockerelli
- Bombus ephippiatus
- Bombus flavescens
- Bombus flavifrons
- Bombus frigidus
- Bombus haematurus
- Bombus huntii
- Bombus hypnorum - tree bumblebee
- Bombus impatiens
- Bombus infirmus
- Bombus infrequent
- Bombus jonellus - Heidehummel
- Bombus kotzschi
- Bombus lapponicus
- Bombus sylvicola
- Bombus lemniscatus
- Bombus lepidus
- Bombus luteipes
- Bombus melanopygus
- Bombus mirus
- Bombus mixtus
- Bombus modestus
- Bombus monticola - mountain bumblebee
- Bombus oceanicus
- Bombus parthenius
- Bombus perplexus
- Bombus picipes
- Bombus pratorum - meadow bumblebee
- Bombus pressus
- Bombus pyrenaeus
- Bombus rotundiceps
- Bombus sandersoni
- Bombus sitkensis
- Bombus sonani
- Bombus subtypicus
- Bombus ternarius
- Bombus vagans
- Bombus vandykei
- Bombus vosnesenskii
- Bombus wilmattae
Subgenus Sibiricobombus
- Bombus asiaticus
- Bombus morawitzi
- Bombus niveatus
- Bombus oberti
- Bombus obtusus
- Bombus sibiricus
- Bombus sulfureus
Sub-genus subterranean obombus
- Bombus amurensis
- Appositus bomb
- Bombus borealis
- Bombus distinguendus - Deichhummel
- Bombus fedtschenkoi
- Bombus fragrans
- Bombus melanurus
- Bombus mongolensis
- Bombus personatus
- Bombus subterraneus
Subgenus Thoracobombus
- Bombus anachoreta
- Bombus armeniacus - Armeniacushummel
- Bombus atratus
- Bombus atripes
- Bombus bellicosus
- Bombus brasiliensis
- Bombus brevivillus
- Bombus dahlbomii
- Bombus deuteronymus
- Bombus digressus
- Bombus diligens
- Bombus excellens
- Bombus exile
- Bombus fervidus
- Bombus filchnerae
- Bombus hedini
- Bombus honshuensis
- Bombus humilis - mutable bumblebee
- Bombus imitator
- Bombus impetuosus
- Bombus inexspectatus
- Bombus laesus
- Bombus medius
- Bombus mesomelas
- Bombus mexicanus
- Bombus mlokosievitzii
- Bombus morio
- Bombus mucidus - off- white bumblebee
- Bombus muscorum - Mooshummel
- Bombus opifex
- Bombus opulentus
- Bombus pascuorum - Bumblebee
- Bombus pensylvanicus
- Bombus persicus
- Bombus pomorum - fruit bumblebee
- Bombus pseudobaicalensis
- Bombus pullatus
- Bombus remotus
- Bombus rubriventris (probably extinct)
- Bombus ruderarius - grass bumblebee
- Bombus schrencki
- Bombus steindachneri
- Bombus sylvarum - wood bumblebee
- Transversalis bomb
- Bombus tricornis
- Bombus trinominatus
- Bombus velox
- Bombus veteranus
- Bombus white
- Bombus zonatus
literature
- PH Williams: Phylogenetic relationships among bumblebees. in: Systematic entomology. Blackwell, Oxford 19.1994, 327-344, ISSN 0307-6970
- PH Williams: An annotated checklist of bumblebees with an analysis of patterns of description. in: Bulletin of the Natural History Museum (Entomology). Intercept, Andover 67.1998, 79-152, ISSN 0968-0454
Individual evidence
- ^ PH Williams. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini) . in: Bulletin of The Natural History Museum (Entomology) 67: 79-152 [updated by http://www.nhm.ac.uk/research-curation/projects/bombus/ ].
- ^ SA Cameron, HM Hines, PH Williams. (2007) A comprehensive phylogeny of the bumble bees (Bombus) . in: Biological Journal of the Linnean Society 91, 161-188.
- ^ Williams, PH, Cameron SA, Hines HM, Cederberg B., Rasmont P. (2008) A simplified subgeneric classification of the bumblebees (genus Bombus) . Apidology 39, [1-29].
Web links
- bombus.de Hummel page by Alexander Schlecht