Snails

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Cochlicellidae
Slender snail (Cochlicella acuta)

Slender snail ( Cochlicella acuta )

Systematics
Subclass : Orthogastropoda
Superordinate : Heterobranchia
Order : Lung snails (pulmonata)
Subordination : Land snails (Stylommatophora)
Superfamily : Helicoidea
Family : Cochlicellidae
Scientific name
Cochlicellidae
Schileyko , 1972

The tip snails (Cochlicellidae) are a relatively small family from the suborder of the land snails (Stylommatophora). The family includes about 25 species. Some authors question the independence of the family. In MolluscaBase they are only accepted as tribe Cochlicellini .

features

The housings are flattened-conical to highly conical. These are relatively small cases with a diameter of up to 12 mm and a height of up to 9 mm. The mouth is usually simple. The navel is narrow to closed. With the exception of the spindle area, the mouth is mostly simple, neither thickened nor turned over, the sculpting small. Exceptions are some types of Obelus with a heavily modified mouth and strong sculptures. In the genital system, the penis is relatively short and the epiphallus is usually much longer. The boundary between the penis and epiphallus is marked by the insertion of the penile retractor muscle. The penile papilla is perforated and z. T. with an edge. The flagellum is always present, but it can be of different lengths. The atrium or vagina has an appendix that is likely to be interpreted as a modified love arrow sack. There are no love arrows . One to three tubular, simple or branching mucous glands open in the apex of the appendix.

Occurrence, way of life and distribution

The species of the family live in southern Europe and North Africa, on the Atlantic coast of France, Belgium, Great Britain and Ireland, and on the Canary Islands . Some species have now also been introduced to other regions of the world (e.g. to Australia).

Systematics

The family Cochlicellidae is no longer recognized by all authors as an independent family. According to Schileyko (2004), with additions by Bank et al. (2002) the family contains the following genera and species:

  • Schileyko family Cochlicellidae, 1973

Cochlicella

  • Cochlicella acuta acuta (subadult)

  • Cochlicella acuta acuta (adult)

  • Cochlicella acuta acuta f. gigantea

  • Cochlicella acuta raphidia
    (pale form)

  • Cochlicella acuta raphidia
    (colored form)

  • Cochlicella barbara (slim shape)

  • Cochlicella barbara (plump shape)


Monilearia

  • Monilearia caementitia

  • Monilearia granostriata

  • Monilearia monilifera (fossil)

  • Monilearia monilifera

  • Monilearia oleacea

  • Monilearia persimilis

  • Monilearia phalerata

  • Monilearia praeposita


Obelus

  • Obelus despreauxii (fossil)

  • Obelus discogranulatus

  • Obelus moderatus

  • Obelus moratus


Phylogeny

According to the cladistic analysis by Steinke, Albrecht & Pfenninger (2004) based on molecular data , the genus Cochlicella is very closely related to Sphincterochila . Both genera are within the other subfamilies or genera of the leaf snails (Hygromiidae). Cochlicella is also systematized within the Hygromiidae in the cladogram by Koene & Schulenburg (2005) . If the findings are confirmed, the Cochlicellidae and Spincterochilidae can certainly no longer be maintained in the rank of families (cf.).



  Hygromiidae  




 Helicellinae 


   

 Cochlicella / Sphincterochila 





   

 Hygromiinae 



  Helicidae  


 Helicinae 


   

 Ariantinae 



   

 Helicodontidae 




   

 Bradybaenidae 



Template: Klade / Maintenance / Style

MolluscaBase devalues ​​the group to tribe and places them in the subfamily Geometrinae in the family Geometridae.

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literature

  • Ruud A. Bank, Klaus Groh and Theo EJ Ripken: Catalog and Bibliography of the non-marine Mollusca of Macaronesia. Collectanea Malacologica (Festschrift for Gerhard Falkner), pp. 89–235, ConchBooks, Hackenheim 2002 ISBN 3-925919-61-9
  • Philippe Bouchet & Jean-Pierre Rocroi: Part 2. Working classification of the Gastropoda . Malacologia, 47: 239-283, Ann Arbor 2005 ISSN  0076-2997
  • Joris M. Koene and Hinrich Schulenburg: Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails. BMC Evolutionary Biology, 5:25, 13 pages, 2005 doi : 10.1186 / 1471-2148-5-25
  • Hartmut Nordsieck: Higher classification of the Helicoidea and the molecular analyzes of their phylogeny . 2006 PDF (server currently not accessible)
  • Anatolij A. Schileyko: Treatise on Recent terrestrial pulmonate molluscs, Part 12 Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica, Supplement 2 (12): 1627-1763, Moscow 2005 ISSN  0136-0027
  • Dirk Steinke, Christian Albrecht and Markus Pfenninger: Molecular phylogeny and character evolution in the Western Palaearctic Helicidae sl (Gastropoda: Stylommatophora). Molecular Phylogenetics and Evolution, 32: 724-734, San Diego 2004 doi : 10.1016 / j.ympev.2004.03.004 .

Individual evidence

  1. ↑ Geoff H. Baker: Helicidae and Hygromiidae as Pests in Cereal Cropsand Pastures in Southern Australia. In: Gary M. Barker (Ed.): Molluscs as crop pests , pp. 193-215, Wallingford, 2002 ISBN 0-85199-320-6
  2. ↑ Miguel Ibáñez, Klaus Groh, María Alonso, Carolina Castillo and Yurena Yanes: The subgenus Monilearia (Lyrula) Wollaston, 1878 (Gastropoda: Helicoidea: Cochlicellidae) from Lanzarote and Fuerteventura (Canary Islands), with the description of Monilearia (Lyrula) tubaeformis sp. nov. Zootaxa, 1320: 29-41, Auckland 2006 Abstract (PDF; 14 kB)
  3. ↑ Miguel Ibáñez, María R. Alonso, Klaus Groh and Rainer Hutterer : The Genus Obelus Hartmann, 1842 (Gastropoda, Pulmonata, Helicoidea) and its Phylogenetic Relationships. Zoologischer Anzeiger, 242: 157-167, Amsterdam & Jena 2003 ISSN  0044-5231
  4. ↑ according to Bank et al. (2002)
  5. ↑ Cochlicella in AnimalBase - here already in Hygromiidae
  6. ↑ MolluscaBase: Cochlicellini Shileyko, 1972

Web links

Commons : Cochlicellidae  - Collection of images, videos, and audio files