Shrub snails
| Shrub snails | ||||||||||||
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Nabeled shrub snail ( Fruticicola fruticum ) |
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| Scientific name | ||||||||||||
| Bradybaenidae | ||||||||||||
| Pilsbry , 1934 |
The shrub snails ( Bradybaenidae) are a family from the suborder of the land snails (Stylommatophora). There is only one type of family in Europe.
features
The housings are relatively large and usually rolled up in a spherical shape, less often lenticular or tower-shaped. The embryonic convolutions are smooth, slightly folded radially or provided with small pustules. The winding sections of the juvenile and adult housing usually have clear spiral pits that are rarely weak or even absent. Hair can also be formed on this section of the housing. The sole of the foot is smooth. The animals only have a single love arrow , with the exception of a single genus that has two love arrows. However, in some species the love arrows are completely missing.
Occurrence and distribution
The family of the shrub snail is found mainly in Central and East Asia. It penetrates into the tropical zones. Only one species is native to Western Asia and Europe.
Systematics
The family was revised by Schileyko in 2004; the following system follows this work.
- Bradybaenidae family Pilsbry, 1924
- Subfamily Aegistini Kuroda & Habe, 1949
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Aegista Albers, 1850
- Aegista tokyoensis Sorita, 1980
- Coelorus Pilsbry, 1900
- Dolicheulota Pilsbry, 1901
- Euhadra Pilsbry, 1890
- Guamampa Schileyko, 1996
- Landouria Godwin-Austin, 1918
- Lepidopisum Habe, 1957
- Mandarina Pilsbry, 1895
- Mastigeulota Pilsbry, 1895
- Miyakoia Minato, 1980
- Neoaegista Azuma, 1955
- Neochloritis Minato, 1982
- Nesiohelix Kuroda & Emura, 1943
- Nipponochloritis Habe, 1955
- Pancala Kuroda & Habe, 1949
- Plecteulota Moellendorf, 1892
- Plectotropis Martens in Albers, 1860
- Pseudaspasita Moelendorf, 1891
- Satsuma A. Adams, 1868 (with the subgenera Satsuma (Satsuma) A. Adams, 1868, Satsuma (Coniglobus) Pilsbry & Hirase, 1906 and Satsuma (Luchuhadra) Kuroda & Habe, 1949)
- Thaitropis Schileyko, 2004
- Torobaena Haas, 1935
- Tricheulota Pilsbry, 1895
- Trishoplita Jacobi, 1898
- Yakuchloritis Got, 1955
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Aegista Albers, 1850
- Bradybaeninae Pilsbry subfamily, 1934
- Acusta Martens, 1860
- Ainohelix Kuroda & Taki, 1933
- Apatetes Gude, 1914
- Armandiella Ancey, 1901
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Bradybaena Beck, 1837
- Bradybaena similaris (Férussac, 1821)
- Buliminidius Heude, 1890
- Cathaica Moellendorff, 1884 (with the subgenera Cathaica (Cathaica) Moellendorf, 1884, Cathaica (Pliocathaica) Andreae, 1900 and Cathaica (Xerocathaica) Andreae, 1900)
- Chalepotaxis Ancey, 1887
- Chosenelix Pilsbry, 1927
- Coccoglypta Pilsbry, 1895
- Ezohelix Kuroda & Emura, 1938
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Fruticicola hero, 1838
- Nabeled shrub snail ( Fruticicola fruticum )
- Grabauia Yen, 1935
- Karaftohelix Pilsbry, 1927
- Laeocathaica Moellendorff, 1899
- Metadontia Moellendorff, 1886
- Mikiria Godwin-Austen, 1918
- Neseulota Ehrmann, 1912
- Paraegista Kuroda & Azuma, 1951
- Phaeohelix Kuroda & Habe, 1949
- Platypetasus Pilsbry, 1895
- Ponsadenia Schileyko, 1978 (with the subgenera Ponsadenia (Ponsadenia) Schileyko, 1978, Pondadenia (Tarbagataja) Schileyko, 1978, Ponsadenia (Mesasiata) Schileyko, 1978 and Ponsadenia (Dzungaria) Schileyko, 2004)
- Pseudiberus Ancey, 1887
- Pseudobuliminus Gredler, 1887
- Rudens Heude, 1890
- Secusana Gredler, 1894
- Semibuliminus Moellendorff, 1898
- Stenogyropsis Moellendorff, 1899
- Stilpnodiscus Moellendorff, 1899
- Trichobradybaena Wu & Guo, 2003
- Trichocathaica Gude, 1919
- Subfamily Helicostylinae Ihering, 1909
- Anixia Pilsbry, 1894
- Canistrum Mörch, 1852
- Chloraea Albers, 1850
- Chrysallis Albers, 1850
- Cochlodryas Martens, 1860
- Cochlostyla Férussac, 1821
- Corasia Albers, 1850
- Dolichostyla Pilsbry, 1896
- Dryocochlias Moellendorff, 1898
- Helicobulinus Broderip, 1841
- Helicostyla Férussac, 1821 (with the subgenera Helicostyla (Helicostyla) Férussac, 1821, Helicostyla (Opalliostyla) Pilsbry, 1896, Helicostyla (Calocochlea) Hartmann, 1843 and Helicostyla (Chromatosphaera) Pilsbry, 1892)
- Hypselostyla Martens, 1865
- Leytia Pilsbry, 1892
- Mesanella Clench & Turner, 1952
- Orustia Mörch, 1852
- Pachysphaera Pilsbry, 1891
- Pfeifferia J. Gray, 1853
- Phengus Albers, 1850
- Phoenicobius Mörch, 1852
- Pyrochilus Pilsbry, 1893
- Rhymbocochlias Moellendorf, 1895
- Steatodryas Pilsbry, 1932
- Trachystyla Pilsbry, 1892
- Subfamily Aegistini Kuroda & Habe, 1949
Phylogeny
The systematic position of the family in the Helicoidea system has not yet been conclusively clarified. In the cladogram by Steinke, Albrecht & Pfenninger (2004), the Bradybaenidae are the sister group of a monophylum consisting of Helicidae and Hygromiidae.
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In the cladogram by Koene & Schulenburg (2005), however, the Bradybaenidae are the sister group of a monophylum consisting of Helicidae and Helminthoglyptidae. The Hygromiidae, on the other hand, form the outer group of Bradybaenidae, Helicidae and Helminthoglyptidae.
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literature
- Rosina Fechter and Gerhard Falkner: molluscs . (Steinbach Naturführer 10), 287 p.,
- Jürgen H. Jungbluth and Dietrich von Knore: Trivial names of land and freshwater mollusks in Germany (Gastropoda et Bivalvia). Mollusca, 26 (1): 105-156, Dresden 2008 ISSN 1864-5127 PDF
- Joris M. Koene and Hinrich Schulenburg: Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails. BMC Evolutionary Biology, 5:25, 13 pages, 2005 doi : 10.1186 / 1471-2148-5-25
- Anatolij A. Schileyko: Treatise on Recent terrestrial pulmonate molluscs, Part 12 Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica, Supplement 2 (12): 1627-1763, Moscow 2005 ISSN 0136-0027
- Dirk Steinke, Christian Albrecht and Markus Pfenninger: Molecular phylogeny and character evolution in the Western Palaearctic Helicidae sl (Gastropoda: Stylommatophora). Molecular Phylogenetics and Evolution, 32: 724-734, San Diego 2004 doi : 10.1016 / j.ympev.2004.03.004 .
