Boophis: Difference between revisions

Bright-eyed frogs
	Boophis ankaratra
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Family:	Mantellidae
Subfamily:	Boophinae; Vences & Glaw, 2001
Genus:	Boophis; Tschudi, 1838
Type species
	Boophis goudotii; Tschudi, 1838
Synonyms
	Buccinator Gistel, 1848;

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Boophis is the only genus in the mantellid frog subfamily Boophinae. They are commonly known as bright-eyed or skeleton frogs. They show typical 'tree frog' traits, and are a good example of convergent evolution with morphologically similar species in the families Hylidae and Rhacophoridae, among others. This genus can only be found on Madagascar and Mayotte Island (Comoros).^[1]

Taxonomy[edit]

The genus Boophis was described by Johann Jakob von Tschudi in 1838.^[2] It was originally considered a member of the African-Asian family Rhacophoridae, but was moved to the family Mantellidae in 2001,^[3] into its own subfamily, Boophinae Vences & Glaw, 2001.

Morphology[edit]

Boophis are arboreal frogs, exhibiting many traits that define 'tree frogs', such as expanded toe discs, long hindlimbs, and large eyes.^[1] Boophis are especially characterised by bright colouration of the iris, which is typically intricately patterned towards its inside, and often green or blue, but occasionally also red, purple, or yellow in the outer iris area.^[1] This has led to the vernacular name of bright-eyed frogs for the genus. Many species of Boophis have almost translucent skin, allowing bones and internal organs to be observed as in the unrelated glass frogs (Centrolenidae) of the tropical Americas.^[4] This has led to the vernacular name skeleton frogs for some members of the genus.

Species[edit]

The genus has nearly 80 species; new ones are being described every few months on average. This list may not be exhaustive.

Boophis occidentalis tadpole

Boophis albilabris (Boulenger, 1888)
Boophis albipunctatus Glaw & Thiesmeier, 1993
Boophis andohahela Andreone, Nincheri & Piazza, 1995
Boophis andrangoloaka (Ahl, 1928)
Boophis andreonei Glaw & Vences, 1994
Boophis anjanaharibeensis Andreone, 1996
Boophis ankarafensis Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014
Boophis ankaratra Andreone, 1993
Boophis arcanus Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis axelmeyeri Vences, Andreone & Veities, 2005
Boophis baetkei J. Köhler, Glaw & Vences, 2008
Boophis blommersae Glaw & Vences, 1994
Boophis boehmei Glaw & Vences, 1992
Boophis boppa Hutter, Lambert, Andriampenomanana & Vences, 2015
Boophis bottae Vences & Glaw, 2002
Boophis brachychir Boettger, 1882
Boophis burgeri Glaw & Vences, 1994
Boophis calcaratus Vallan, Vences & Glaw, 2010
Boophis doulioti Angel, 1934
Boophis elenae Andreone, 1993
Boophis englaenderi Glaw & Vences, 1994
Boophis entingae Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis erythrodactylus (Guibé, 1953)
Boophis fayi J. Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
Boophis feonnyala Glaw, Vences, Andreone & Vallan, 2001
Boophis goudotii Tschudi, 1838
Boophis guibei (McCarthy, 1978)
Boophis haematopus Glaw, Vences, Andreone & Vallan, 2001
Boophis haingana Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis idae (Steindachner, 1867)
Boophis jaegeri Glaw & Vences, 1992 – Green Skeleton Frog
Boophis laurenti Guibé, 1947
Boophis liami Vallan, Vences & Glaw, 2003
Boophis lichenoides Vallan, Glaw, Andreone & Cadle, 1998
Boophis lilianae J. Köhler, Glaw & Vences, 2008
Boophis luciae Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis luteus (Boulenger, 1882)
Boophis madagascariensis (W. Peters, 1874)
Boophis majori (Boulenger, 1896)
Boophis mandraka Blommers-Schlösser, 1979
Boophis marojezensis Glaw & Vences, 1994
Boophis masoala Glaw, Scherz, Prötzel & Vences, 2018
Boophis miadana Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis microtympanum (Boettger, 1881)
Boophis miniatus (Mocquard, 1902)
Boophis narinsi Vences, Gehara, J. Köhler & Glaw, 2012
Boophis nauticus Glaw, Hawlitschek, K. Glaw & Vences, 2019
Boophis obscurus (Boettger, 1913)
Boophis occidentalis Glaw & Vences, 1994
Boophis opisthodon (Boulenger, 1888)
Boophis pauliani (Guibé, 1953)
Boophis periegetes Cadle, 1995
Boophis picturatus Glaw, Vences, Andreone & Vallan, 2001
Boophis piperatus Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis popi J. Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
Boophis praedictus Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis pyrrhus Glaw, Vences, Andreone & Vallan, 2001
Boophis quasiboehmei Vences, J. Köhler, Crottini & Glaw, 2010
Boophis rappiodes (Ahl, 1928)
Boophis reticulatus Blommers-Schlösser, 1979
Boophis rhodoscelis (Boulenger, 1882)
Boophis roseipalmatus Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis rufioculis Glaw & Vences, 1997
Boophis sambirano Vences & Glaw, 2005
Boophis sandrae Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis schuboeae Glaw & Vences, 2002
Boophis septentrionalis Glaw & Vences, 1994
Boophis sibilans Glaw & Thiesmeier, 1993
Boophis solomaso Vallan, Vences & Glaw, 2003
Boophis spinophis Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010
Boophis tampoka J. Köhler, Glaw & Vences, 2007
Boophis tasymena Vences & Glaw, 2002
Boophis tephraeomystax (A.H.A. Duméril, 1853)
Boophis tsilomaro Vences, Andreone, Glos & Glaw, 2010
Boophis ulftunni Wollenberg, Andreone, Glaw & Vences, 2008
Boophis viridis Blommers-Schlösser, 1979
Boophis vittatus Glaw, Vences, Andreone & Vallan, 2001
Boophis williamsi (Guibé, 1974)
Boophis xerophilus Glaw & Vences, 1997

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than boophis.

References[edit]

^ ^a ^b ^c Glaw, Frank; Vences, Miguel (2007). A Field Guide to the Amphibians and Reptiles of Madagascar (3rd ed.). Cologne, Germany: Vences & Glaw Verlags. ISBN 978-3929449037.
^ Tschudi, J. J. v. (1838). Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Neuchâtel: Petitpierre.
^ Vences, Miguel; Glaw, Frank (2001). "When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs". Spixiana. 24 (1): 85–92.
^ Hutter, Carl; Lambert, Shea M.; Andriampenomanana, Zo F.; Glaw, Frank; Vences, Miguel (2018-10-01). "Molecular phylogeny and diversification of Malagasy bright-eyed tree frogs (Mantellidae: Boophis)". Molecular Phylogenetics and Evolution. 127: 568–578. doi:10.1016/j.ympev.2018.05.027. ISSN 1055-7903. PMID 29894731.

[GnV07-1] Glaw, Frank; Vences, Miguel (2007). A Field Guide to the Amphibians and Reptiles of Madagascar (3rd ed.). Cologne, Germany: Vences & Glaw Verlags. ISBN 978-3929449037.

[Tschudi38-2] Tschudi, J. J. v. (1838). Classification der Batrachier mit Berücksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Neuchâtel: Petitpierre.

[VG01-3] Vences, Miguel; Glaw, Frank (2001). "When molecules claim for taxonomic changes: New proposals on the classification of Old World treefrogs". Spixiana. 24 (1): 85–92.

[4] Hutter, Carl; Lambert, Shea M.; Andriampenomanana, Zo F.; Glaw, Frank; Vences, Miguel (2018-10-01). "Molecular phylogeny and diversification of Malagasy bright-eyed tree frogs (Mantellidae: Boophis)". Molecular Phylogenetics and Evolution. 127: 568–578. doi:10.1016/j.ympev.2018.05.027. ISSN 1055-7903. PMID 29894731.

[1]

[2]

[3]

[4]

@@ Line 1: / Line 1: @@
+{{Short description|Genus of amphibians}}
 {{Automatic taxobox
 | name = Bright-eyed frogs
@@ Line 8: / Line 9: @@
 | type_species = ''[[Boophis goudotii]]''
 | type_species_authority = [[Johann Jakob von Tschudi|Tschudi]], 1838
-| diversity_link = #Species
-| diversity = 79 species
 | synonyms =
 * ''Buccinator'' {{small|Gistel, 1848}}
@@ Line 25: / Line 24: @@
 ==Species==
-The genus has presently nearly 80  [[species]]; new ones are being described every few months on average. This list may not be exhaustive.
+The genus has nearly 80 [[species]]; new ones are being described every few months on average. This list may not be exhaustive.
 [[File:Boophis occidentalis03.jpg|thumb|right|''[[Boophis occidentalis]]'' [[tadpole]]]]
+{{div col|colwidth=35em}}
-<div float="left">
+* ''[[Boophis albilabris]]'' <small>([[George Albert Boulenger|Boulenger]], 1888)</small>
-{{col-begin|width=60%}}
+* ''[[Boophis albipunctatus]]'' <small>[[Frank Glaw|Glaw]] & [[species:Burkhard Thiesmeier|Thiesmeier]], 1993</small>
-{{col-1-of-2}}
-* ''[[Boophis albilabris]]'' <small>(Boulenger, 1888)</small>
+* ''[[Boophis andohahela]]'' <small>[[species:Franco Andreone|Andreone]], [[species:Riccardo Nincheri|Nincheri]] & [[species:Roberto Piazza|Piazza]], 1995</small>
-* ''[[Boophis albipunctatus]]'' <small>[[Frank Glaw|Glaw]] & Thiesmeier, 1993</small>
+* ''[[Boophis andrangoloaka]]'' <small>([[Ernst Ahl|Ahl]], 1928)</small>
-* ''[[Boophis andohahela]]'' <small>Andreone, Nincheri & Piazza, 1995</small>
+* ''[[Boophis andreonei]]'' <small>Glaw & [[Miguel Vences|Vences]], 1994</small>
-* ''[[Boophis andrangoloaka]]'' <small>(Ahl, 1928)</small>
-* ''[[Boophis andreonei]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
 * ''[[Boophis anjanaharibeensis]]'' <small>Andreone, 1996</small>
-* ''[[Boophis ankarafensis]]'' <small>Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014</small>
+* ''[[Boophis ankarafensis]]'' <small>[[species:Samuel G. Penny|Penny]], Andreone, [[species:Angelica Crottini|Crottini]], [[species:Marc W. Holderied|Holderied]], [[species:Lovasoa Manuelle Sylviane Rakotozafy|Rakotozafy]], [[species:Christoph Schwitzer|Schwitzer]] & [[species:Gonçalo M. Rosa|Rosa]], 2014</small>
 * ''[[Boophis ankaratra]]'' <small>Andreone, 1993</small>
-* ''[[Boophis arcanus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis arcanus]]'' <small>Glaw, [[Jörn Köhler|J. Köhler]], [[species:Ignacio J. De la Riva|De la Riva]], [[species:David R. Vieites|Vieites]] & Vences, 2010</small>
-* ''[[Boophis axelmeyeri]]'' <small>[[Miguel Vences|Vences]], Andreone & Veities, 2005</small>
+* ''[[Boophis axelmeyeri]]'' <small>Vences, Andreone & Veities, 2005</small>
-* ''[[Boophis baetkei]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small><!--  Zootaxa1814:37. -->
+* ''[[Boophis baetkei]]'' <small>J. Köhler, Glaw & Vences, 2008</small><!--  Zootaxa1814:37. -->
-* ''[[Boophis blommersae]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
+* ''[[Boophis blommersae]]'' <small>Glaw & Vences, 1994</small>
-* ''[[Boophis boehmei]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1992</small>
+* ''[[Boophis boehmei]]'' <small>Glaw & Vences, 1992</small>
-* ''[[Boophis boppa]]'' <small>Hutter & Lambert, 2015</small>
+* ''[[Boophis boppa]]'' <small>[[species:Carl R. Hutter|Hutter]], [[species:Shea M. Lambert|Lambert]], [[species:Zo Faniry Andriampenomanana|Andriampenomanana]] & Vences, 2015</small>
-* ''[[Boophis bottae]]'' <small>[[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2002</small>
+* ''[[Boophis bottae]]'' <small>Vences & Glaw, 2002</small>
-* ''[[Boophis brachychir]]'' <small>Boettger, 1882</small>
+* ''[[Boophis brachychir]]'' <small>[[Oskar Boettger|Boettger]], 1882</small>
-* ''[[Boophis burgeri]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
+* ''[[Boophis burgeri]]'' <small>Glaw & Vences, 1994</small>
-* ''[[Boophis calcaratus]]'' <small>Vallan, [[Miguel Vences|Vences]] & Glaw, 2010</small>
+* ''[[Boophis calcaratus]]'' <small>[[species:Denis Vallan|Vallan]], Vences & Glaw, 2010</small>
-* ''[[Boophis doulioti]]'' <small>Angel, 1934</small>
+* ''[[Boophis doulioti]]'' <small>[[Fernand Angel|Angel]], 1934</small>
 * ''[[Boophis elenae]]'' <small>Andreone, 1993</small>
-* ''[[Boophis englaenderi]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
+* ''[[Boophis englaenderi]]'' <small>Glaw & Vences, 1994</small>
-* ''[[Boophis entingae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis entingae]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis erythrodactylus]]'' <small>(Guibé, 1953)</small>
+* ''[[Boophis erythrodactylus]]'' <small>([[Jean Marius René Guibé|Guibé]], 1953)</small>
-* ''[[Boophis fayi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & [[Miguel Vences|Vences]], 2011</small>
+* ''[[Boophis fayi]]'' <small>J. Köhler, Glaw, Rosa, [[species:Philip-Sebastian Gehring|Gehring]], [[species:Maciej Pabijan|Pabijan]], Andreone & Vences, 2011</small>
-* ''[[Boophis feonnyala]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
+* ''[[Boophis feonnyala]]'' <small>Glaw, Vences, Andreone & Vallan, 2001</small>
-* ''[[Boophis goudotii]]'' <small>Tschudi, 1838</small>
+* ''[[Boophis goudotii]]'' <small>[[Johann Jakob von Tschudi|Tschudi]], 1838</small>
-* ''[[Boophis guibei]]'' <small>(McCarthy, 1978)</small>
+* ''[[Boophis guibei]]'' <small>([[species:Colin John McCarthy|McCarthy]], 1978)</small>
-* ''[[Boophis haematopus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
+* ''[[Boophis haematopus]]'' <small>Glaw, Vences, Andreone & Vallan, 2001</small>
-* ''[[Boophis haingana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis haingana]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis idae]]'' <small>(Steindachner, 1867)</small>
+* ''[[Boophis idae]]'' <small>([[Franz Steindachner|Steindachner]], 1867)</small>
-* ''[[Boophis jaegeri]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1992</small> – [[Green Skeleton Frog]]
+* ''[[Boophis jaegeri]]'' <small>Glaw & Vences, 1992</small> – [[Green Skeleton Frog]]
 * ''[[Boophis laurenti]]'' <small>Guibé, 1947</small>
-* ''[[Boophis liami]]'' <small>Vallan, [[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2003</small>
+* ''[[Boophis liami]]'' <small>Vallan, Vences & Glaw, 2003</small>
-* ''[[Boophis lichenoides]]'' <small>Vallan, [[Frank Glaw|Glaw]], Andreone & Cadle, 1998</small>
+* ''[[Boophis lichenoides]]'' <small>Vallan, Glaw, Andreone & [[species:John E. Cadle|Cadle]], 1998</small>
-* ''[[Boophis lilianae]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small><!--  Zootaxa1814:37. -->
+* ''[[Boophis lilianae]]'' <small>J. Köhler, Glaw & Vences, 2008</small><!--  Zootaxa1814:37. -->
-* ''[[Boophis luciae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis luciae]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
 * ''[[Boophis luteus]]'' <small>(Boulenger, 1882)</small>
-* ''[[Boophis madagascariensis]]'' <small>(Peters, 1874)</small>
+* ''[[Boophis madagascariensis]]'' <small>([[Wilhelm Peters|W. Peters]], 1874)</small>
-{{col-2-of-2}}
 * ''[[Boophis majori]]'' <small>(Boulenger, 1896)</small>
-* ''[[Boophis mandraka]]'' <small>Blommers-Schlösser, 1979</small>
+* ''[[Boophis mandraka]]'' <small>[[species:Rose M.A. Blommers-Schlösser|Blommers-Schlösser]], 1979</small>
-* ''[[Boophis marojezensis]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
+* ''[[Boophis marojezensis]]'' <small>Glaw & Vences, 1994</small>
-* ''[[Boophis masoala]]'' <small>[[Frank Glaw|Glaw]], Scherz, Prötzel & [[Miguel Vences|Vences]], 2018</small>
+* ''[[Boophis masoala]]'' <small>Glaw, [[species:Mark D. Scherz|Scherz]], [[species:David Prötzel|Prötzel]] & Vences, 2018</small>
-* ''[[Boophis miadana]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis miadana]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
 * ''[[Boophis microtympanum]]'' <small>(Boettger, 1881)</small>
-* ''[[Boophis miniatus]]'' <small>(Mocquard, 1902)</small>
+* ''[[Boophis miniatus]]'' <small>([[François Mocquard|Mocquard]], 1902)</small>
-* ''[[Boophis narinsi]]'' <small>[[Miguel Vences|Vences]], Gehara, Köhler & [[Frank Glaw|Glaw]], 2012</small>
+* ''[[Boophis narinsi]]'' <small>Vences, [[Marcelo Gehara|Gehara]], J. Köhler & Glaw, 2012</small>
-* ''[[Boophis nauticus]]'' <small>Glaw, Hawlitschek, Glaw, and Vences, 2019</small>
+* ''[[Boophis nauticus]]'' <small>Glaw, [[species:Oliver Hawlitschek|Hawlitschek]], [[species:Kathrin Glaw|K. Glaw]] & Vences, 2019</small>
 * ''[[Boophis obscurus]]'' <small>(Boettger, 1913)</small>
-* ''[[Boophis occidentalis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1994</small>
+* ''[[Boophis occidentalis]]'' <small>Glaw & Vences, 1994</small>
 * ''[[Boophis opisthodon]]'' <small>(Boulenger, 1888)</small>
 * ''[[Boophis pauliani]]'' <small>(Guibé, 1953)</small>
 * ''[[Boophis periegetes]]'' <small>Cadle, 1995</small>
-* ''[[Boophis picturatus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
+* ''[[Boophis picturatus]]'' <small>Glaw, Vences, Andreone & Vallan, 2001</small>
-* ''[[Boophis piperatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis piperatus]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis popi]]'' <small>Köhler, [[Frank Glaw|Glaw]], Rosa, Gehring, Pabijan, Andreone & [[Miguel Vences|Vences]], 2011</small>
+* ''[[Boophis popi]]'' <small>J. Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011</small>
-* ''[[Boophis praedictus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & Vences, 2010</small>
+* ''[[Boophis praedictus]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis pyrrhus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
+* ''[[Boophis pyrrhus]]'' <small>Glaw, Vences, Andreone & Vallan, 2001</small>
-* ''[[Boophis quasiboehmei]]'' <small>[[Miguel Vences|Vences]], Köhler, Crottini & [[Frank Glaw|Glaw]], 2010</small>
+* ''[[Boophis quasiboehmei]]'' <small>Vences, J. Köhler, Crottini & Glaw, 2010</small>
 * ''[[Boophis rappiodes]]'' <small>(Ahl, 1928)</small>
 * ''[[Boophis reticulatus]]'' <small>Blommers-Schlösser, 1979</small>
 * ''[[Boophis rhodoscelis]]'' <small>(Boulenger, 1882)</small>
-* ''[[Boophis roseipalmatus]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis roseipalmatus]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis rufioculis]]'' <small>[[Frank Glaw|Glaw]] & Vences, 1997</small>
+* ''[[Boophis rufioculis]]'' <small>Glaw & Vences, 1997</small>
-* ''[[Boophis sambirano]]'' <small>Vences & [[Frank Glaw|Glaw]], 2005</small>
+* ''[[Boophis sambirano]]'' <small>Vences & Glaw, 2005</small>
-* ''[[Boophis sandrae]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis sandrae]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis schuboeae]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2002</small>
+* ''[[Boophis schuboeae]]'' <small>Glaw & Vences, 2002</small>
-* ''[[Boophis septentrionalis]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1994</small>
+* ''[[Boophis septentrionalis]]'' <small>Glaw & Vences, 1994</small>
-* ''[[Boophis sibilans]]'' <small>[[Frank Glaw|Glaw]] & Thiesmeier, 1993</small>
+* ''[[Boophis sibilans]]'' <small>Glaw & Thiesmeier, 1993</small>
-* ''[[Boophis solomaso]]'' <small>Vallan, [[Miguel Vences|Vences]] & [[Frank Glaw|Glaw]], 2003</small>
+* ''[[Boophis solomaso]]'' <small>Vallan, Vences & Glaw, 2003</small>
-* ''[[Boophis spinophis]]'' <small>[[Frank Glaw|Glaw]], Köhler, de la Riva, Vieites & [[Miguel Vences|Vences]], 2010</small>
+* ''[[Boophis spinophis]]'' <small>Glaw, J. Köhler, De la Riva, Vieites & Vences, 2010</small>
-* ''[[Boophis tampoka]]'' <small>Köhler, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2007</small>
+* ''[[Boophis tampoka]]'' <small>J. Köhler, Glaw & Vences, 2007</small>
-* ''[[Boophis tasymena]]'' <small>Vences & [[Frank Glaw|Glaw]], 2002</small>
+* ''[[Boophis tasymena]]'' <small>Vences & Glaw, 2002</small>
-* ''[[Boophis tephraeomystax]]'' <small>(Duméril, 1853)</small>
+* ''[[Boophis tephraeomystax]]'' <small>([[Auguste Duméril|A.H.A. Duméril]], 1853)</small>
-* ''[[Boophis tsilomaro]]'' <small>[[Miguel Vences|Vences]], Andreone, Glos & [[Frank Glaw|Glaw]], 2010</small>
+* ''[[Boophis tsilomaro]]'' <small>Vences, Andreone, [[species:Julian Glos|Glos]] & Glaw, 2010</small>
-* ''[[Boophis ulftunni]]'' <small>Wollenberg, Andreone, [[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 2008</small>
+* ''[[Boophis ulftunni]]'' <small>[[species:Katharina C. Wollenberg|Wollenberg]], Andreone, Glaw & Vences, 2008</small>
 * ''[[Boophis viridis]]'' <small>Blommers-Schlösser, 1979</small>
-* ''[[Boophis vittatus]]'' <small>[[Frank Glaw|Glaw]], [[Miguel Vences|Vences]], Andreone & Vallan, 2001</small>
+* ''[[Boophis vittatus]]'' <small>Glaw, Vences, Andreone & Vallan, 2001</small>
 * ''[[Boophis williamsi]]'' <small>(Guibé, 1974)</small>
-* ''[[Boophis xerophilus]]'' <small>[[Frank Glaw|Glaw]] & [[Miguel Vences|Vences]], 1997</small>
+* ''[[Boophis xerophilus]]'' <small>Glaw & Vences, 1997</small>
-{{col-end}}
+{{div col end}}
-</div>
+''[[Nota bene]]'': A [[Binomial nomenclature|binomial authority]] in parentheses indicates that the species was originally described in a genus other than ''boophis''.
-==References==
-{{reflist}}
+==References==
 {{Commons category}}
+{{Reflist}}
 {{Taxonbar|from=Q2322420}}
+[[Category:Boophis| ]]
 [[Category:Amphibians of Sub-Saharan Africa]]
 [[Category:Endemic fauna of Madagascar]]
 [[Category:Amphibian genera]]
-[[Category:Boophis| ]]
 [[Category:Fauna of Mayotte]]
 [[Category:Taxa named by Johann Jakob von Tschudi]]