Azecidae
Azecidae | ||||||||||||
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Notched smooth snail ( Azeca goodalli ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Azecidae | ||||||||||||
Watson , 1920 |
The Azecidae are a family of snails belonging to the land snails (Stylommatophora). The type genus Azeca Folin & Bérillon, 1877 was until a few years ago counted by some authors to the Cochlicopidae. Other authors treat the Azecidae as a subfamily of the smooth snail (Cochlicopidae).
features
The housings are elongated-egg-shaped, spindle-shaped to almost cylindrical. The up to 8 turns are slightly arched. The shell is comparatively thick and firm, also thin, the surface smooth and shiny. The color varies from light horn-colored to cherry red. The mouth is comparatively small, egg-shaped and pointed at the top; the longitudinal axis of the mouth is inclined to the longitudinal axis of the housing. The mouth is provided with teeth and lamellae, especially a parietal callus. Protoconch and Teleoconch have very fine spiral stripes. The housing is not marked.
The male sexual apparatus is usually very simple without an epiphallus or with only a very short epiphallus. The spermatic duct opens apically or well before the apex into the moderately long penis. The retractor muscle attaches apically. In the female tract, the sperm duct (vas deferens) branches off relatively high up from the egg duct (spermoviduct). The free fallopian tube is therefore comparatively long. The spermathec is only moderately large, the stem comparatively short. The vagina has a thick muscular layer in which numerous pigment cells are embedded. The area shortly before or after the branch of the spermatic stalk from the free fallopian tube can be more or less thickened. The vagina is short to long.
Geographical distribution
Most of the species live in southern Europe (Balkan, Apennine and Iberian Peninsula), only one species also penetrates to Central and Western Europe. Two species are also found in North Africa (Algeria, Morocco).
Taxonomy
The taxon was established in 1920 by Hugh Watson as the subfamily Azecinae. This means that the name is also available as a tribe, family and superfamily name. The type genus is Azeca Fleming, 1828. Kennard & Woodward (1926) raised the taxon to the family level. As a result, the taxon was often considered a synonym for Cochlicopidae. Bouchet & Rocroi (2005) accepted the taxon, but downgraded it to subfamily within the Cochlicopoidea. Schileyko (1998) sees the taxon as a younger synonym of Cochlicopidae. According to recent molecular genetic studies by Medaira et al. (2010) Azecidae and Cochlicopidae are not closely related. Presumably they even have to be reassigned to a different superfamily. However, detailed investigations are still pending. It is very likely that the Azecidae can be removed from the superfamily Cochlicopoidea. Genera and species currently assigned to the Azecidae:
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Azeca Fleming, 1828
- Azeca goodalli (Férussac, 1821)
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Hypnophila Bourghuignat, 1858
- Hypnophila bisacchii Giusti, 1970
- Hypnophila boissii (Dupuy, 1850)
- Hypnophila cyclothyra (Boettger, 1885)
- Hypnophila cylindracea (Calcara, 1840)
- Hypnophila dohrni (Paulucci, 1882)
- Hypnophila emiliana (Bourguignat, 1858)
- Hypnophila girottii Esu, 1978
- Hypnophila incerta (Bourguignat, 1858)
- Hypnophila malagana Gittenberger & Menkhorst in Gittenberger, 1983
- Hypnophila maroccana (Mousson, 1873)
- Hypnophila polita (Porro, 1838)
- Hypnophila psathyrolena (Bourguignat, 1858)
- Hypnophila pupaeformis (Cantraine, 1835)
- Hypnophila remyi (Boettger, 1949)
- Hypnophila zacynthia (Roth, 1855)
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Cryptazeca Folin & Bérillon, 1877
- Cryptazeca elongata Gómez, 1990
- Cryptazeca kobelti Gittenberger, 1983
- Cryptazeca monodonta (de Folin & Bérillon, 1877)
- Cryptazeca spelaea Gómez, 1990
- Cryptazeca subcylindica Folin & Bérillon, 1877
- Cryptazeca vasconica Zilch, 1973
Schileyko (1998) still lists the genus Gomphroa Westerlund, 1902 , among the Cochlicopidae. Most authors rate this taxon as the younger synonym of Hypnophila Bourguignat, 1858. If the taxon were recognized as an independent taxon, it would have to be placed in the family Azecidae (type species: Zua boissii Dupuy, 1850).
supporting documents
literature
- Philippe Bouchet & Jean-Pierre Rocroi: Part 2. Working classification of the Gastropoda . Malacologia, 47: 239-283, Ann Arbor 2005 ISSN 0076-2997
- Anatolij A. Schileyko: Treatise on Recent terrestrial pulmonate molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. Ruthenica, Supplement 2 (1): 1–127, Moscow 1998 ISSN 0136-0027
- María J. Madeira, María A. Elejalde, Luis J. Chueca & Benjamín J. Gómez-Moliner 2010: Phylogenetic Position of the Genus Cryptazeca and the Family Azecidae within the System of the Stylommatophora. Malacologia, 52 (1): 163-168 .
- Edmund Gittenberger: On Iberian Cochlicopidae and the genus Cryptazeca Gastropoda, Pulmonata. Zoological Mededelingen (Leiden): 5723: 301-320, Leiden 1983 PDF .
Individual evidence
- ^ Hugh Watson: The affinities of Pyramidula, Patulastra, Acanthinula, and Vallonia . Proceedings of the Malacological Society of London, 14: 6-30, London 1920 Online at www.biodiversitylibrary.org (p. 24)
- ^ AS Kennard & BB Woodward: Synonymy of the British non-marine Mollusca. 1926 Online at www.biodiversitylibrary.org (p. 144).
- ↑ Fauna Europaea: Azecidae
- ↑ Gittenberger, pp. 301ff.
- ↑ Welter Schultes in AnimalBase