Cochlespiridae

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Cochlespiridae
Aforia circinata from Taiwan

Aforia circinata from Taiwan

Systematics
Superordinate : Caenogastropoda
Order : Sorbeoconcha
Subordination : Hypsogastropoda
Partial order : Neogastropoda
Superfamily : Conoidea
Family : Cochlespiridae
Scientific name
Cochlespiridae
Powell , 1942
Cochlespira radiata from Barbados

The Cochlespiridae are a family of small to medium-sized marine shell snails . For a long time they were counted as the subfamily Cochlespirinae to the family of slug snails (Turridae) within the superfamily Conoidea (poisonous bugs). In 2011 they were raised to family rank mainly on the basis of molecular genetic findings. Outwardly, the various members of this family are very different from one another and the genetic relationships are hardly reflected in the shape of the housing. The delimitation of individual genera and species is difficult and requires revision.

features

The case has a pagoda-like shape with a high winding tip. This shell morphology was for a long time decisive for the position of the Cochlespiridae as a subfamily within the slug tower snails. The snail shell is usually 20-30 millimeters high, with individual species such as Nihonia maxima it can reach a height of 13 centimeters or even 15 centimeters as with Aforia magnifica . In contrast to most families of the conoid, the siphonal canal is often very long and narrow. The sculpting of the case shows hardly any axial ribs, if present they are very short. The protoconch is smooth, paucispiral, or multispiral. In some multispiral protoconches, the youngest turns are axially or spirally ribbed. The operculum has a peripheral core (nucleus).

The radula in this family is rachigloss, with the radula formula 1 + 0 + R + 0 + 1. The central teeth are broad, arched or almost rectangular, with a small jagged tip in the middle of the upper edge. This middle tooth formation is absent in some species of the genus Aforia . The marginal teeth are forkbone-shaped .

Systematics and taxonomy

Most genera of the Cochlespiridae were united in the subfamily Turriculinae within the slug snails (Turridae) until the second half of the 20th century. As early as 1972 Walter Oliver Cernohorsky pointed out that the name Turriculinae had already been used to subdivide Mitra snails , even if it was no longer valid in this group. The type genus Turricula was created in 1993 by Taylor et al. transferred to the subfamily Clavatulinae , so that a new name for the remaining species of the subfamily had to be found anyway. According to the rules of the ICZN , naming at subfamily, superfamily and family level is only possible according to a genus in this group. Therefore, as suggested by Cernohorsky, the name Cochlespirinae was chosen, and the genus Cochlespira , described in 1895, became the type genus of this subfamily. This also corresponded to the classification and naming of New Zealand sea snails by Arthur William Baden Powell in 1942. The survey as an independent family took place in 2011 in the work of Bouchet, Pouillandre et al.

Genera

There are currently 10 recent genera within this family.

An attempt was made to place Sibogasyrinx Powell, 1969 as a subgenus in the genus Leucosyrinx Dall, 1889 from the family Pseudomelatomidae . Although some species of the genus Sibogasyrinx are more in line with Leucosyrinx in terms of the housing shape , they were left in the Cochlespiridae family due to molecular-biological relationships.

Synonyms

  • Coronasyrinx Powell, 1944, Pagodasyrinx Shuto, 1969, and Tahusyrinx Powell, 1944 are synonyms of Cochlespira Conrad, 1865.
  • Danilacarina Bozzetti, 1997 and Irenosyrinx Dall, 1908, were synonymous with Aforia Dall, 1889.

Individual evidence

  1. ^ AWB Powell: The New Zealand recent and fossil Mollusca of the family Turridae. With general notes on turrid nomenclature and systematics. Bulletin of the Auckland Institute and Museum 1942, 2, pp. 1–192, 1942
  2. ↑ a b J. D. Taylor, YI Kantor & AV Sysoev: Foregut anatomy, feeding mechanisms, relationships and classification of the Conoidea (= Toxoglossa) (Gastropoda) . Bulletin of the Natural History Museum (Zool.), 5, 2, pp. 125-170, London 1993
  3. ↑ Philippe Bouchet & Jean-Pierre Rocroi (eds.): Classification and nomenclator of gastropod families. Part 2. Working classification of the Gastropoda . Malacologia, 47, pp. 239-283, Ann Arbor 2005 ISSN  0076-2997 ISBN 3-925919-72-4
  4. ↑ a b N. Puillandre, Yu. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, JA Todd & Philippe Bouchet: The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies, 77, 3, pp. 259–272, 2011 ( PDF )
  5. ↑ a b c P. Bouchet, YI Kantor, A. Sysoev & N. Puillandre: A new operational classification of the Conoidea. Journal of Molluscan Studies, 77, pp. 273–308, 2011 ( online )
  6. ^ Yuri I. Kantor: Formation of marginal radular teeth in Conoidea (Neogastropoda) and the evolution of the hypodermic envenomation mechanism. Journal of Zoology, 252, 2, pp. 251-262, 2000 doi : 10.1111 / j.1469-7998.2000.tb00620.x
  7. ↑ Walter Oliver Cernohorsky: Comments on the Authorship of Some subfamilial names in the Turridae, Mollusca: Gastropoda . Veliger, 15, pp. 127-128, 1972
  8. ^ Philippe Bouchet: Cochlespiridae Powell, 1942 . In: WoRMS, World Register of Marine Species, 2015, accessed March 23, 2015

literature

  • Philippe Bouchet & Jean-Pierre Rocroi (Eds.): Classification and nomenclator of gastropod families. Part 2. Working classification of the Gastropoda . Malacologia, 47, pp. 239-283, Ann Arbor 2005 ISSN  0076-2997 ISBN 3-925919-72-4
  • Arthur William Baden Powell: The New Zealand recent and fossil Mollusca of the family Turridae. With general notes on turrid nomenclature and systematics. Bulletin of the Auckland Institute and Museum 1942, 2, pp. 1–192, 1942 (first description)

Web links

Commons : Cochlespiridae  - collection of images, videos and audio files
  • Philippe Bouchet: Cochlespiridae Powell, 1942 . In: WoRMS, World Register of Marine Species, 2015, accessed March 23, 2015
  • Cochlespiridae , pictures of various genera from Gastropods.com, accessed March 23, 2015