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Cophyla tuberifera

Cophyla tuberifera

without rank: Amphibians (Lissamphibia)
Order : Frog (anura)
Subordination : Neobatrachia
Family : Narrow-mouth frogs (Microhylidae)
Subfamily : Madagascar narrow -mouth frogs (Cophylinae)
Genre : Cophyla
Scientific name
Boettger , 1880

Cophyla is a genus of the frog from the subfamily of the Madagascar narrow -mouth frogs (Cophylinae) within the family of the narrow-mouth frogs (Microhylidae). It was merged with the genus Platypelis in 2015.


The pupils are horizontal. The tongue is large, oval, with entire margins and freely liftable at the back. The palatal teeth are arranged in two small, closely spaced groups behind the line connecting the choans . The eardrum is not very clearly visible. The fingers and toes are webbed together. These do not intervene between the metatarsi of the 4th and 5th toes. The tips of the fingers and toes are widened to form adhesive discs and have t-shaped or y-shaped bony terminal phalanges. The precoracoid are weak and lie on the coracoid . The omosternum is missing. The sternum is a small plate of cartilage. The transverse processes of the sacral vertebra are moderately widened.


The genus is endemic to Madagascar . The species of the formerly independent genus Platypelis live in the north and north-west of the island, the other species, which are assigned to Cophyla , are distributed in the north-east and south of Madagascar.


The genus Cophyla was first described by Oskar Boettger in 1880 . The distinction to the genus Platypelis , described by George Albert Boulenger in 1882, was based on a few skeletal features that are subject to a wide range of variation in narrow-mouthed frogs. Peloso et al. in their molecular genetic study, which was published in March 2015, could not determine a clear monophyly of the two genera and have therefore synonymized them under the name Cophyla . Several authors who are intensively researching Madagascar's frog fauna, however, do not agree with the merger. Therefore, species are still described in either one or the other genus.


The genus currently has a total of 23 species.

As of June 23, 2020

In the genus Cophyla there were six species before they were merged with Platypelis :

Up until this point in time, the genus Platypelis comprised 13 species, which are now included in Cophyla :

Two more species were described in 2019.

  • Cophyla ando (Scherz, Köhler, Vences & Glaw, 2019) as Platypelis ando
  • Cophyla fortuna Rakotoarison, Scherz, Bletz, Razafindraibe, Glaw & Vences, 2019

Two types were added in 2020:

  • Cophyla ranjomena (Glaw, Scherz, Rakotoarison, Crottini, Raselimanana, Andreone, Köhler & Vences, 2020) originally described as Platypelis ranjomena
  • Cophyla laeta (Rakotoarison, Scherz, Köhler, Ratsoavina, Hawlitschek, Megson, Vences & Glaw, 2020) originally described as Platypelis laetus

Individual evidence

  1. ^ Fritz Nieden: Anura II. In: FE Schulze, W. Kükenthal, K. Heider (Eds.): Das Tierreich, Walter de Gruyter & Co., Berlin and Leipzig 1926, p. 8
  2. ^ Oskar Boettger : Diagnoses Batrachiorum novorum insulae Madagascar. Zoologischer Anzeiger, 3, pp. 567-568, 1880
  3. Pedro LV Peloso, Darrel R. Frost, Stephen J. Richards, Miguel T. Rodrigues, Stephen Donnellan, Masafumi Matsui, Cristopher J. Raxworthy, SD Biju, Emily Moriarty Lemmon, Alan R. Lemmon & Ward C. Wheeler: The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae). Cladistics, 3, 1-28, March 2015 doi : 10.1111 / cla.12118
  4. ^ Darrel R. Frost: Cophyla Boettger, 1880 . In: Amphibian Species of the World: an Online Reference . Version 6.0, accessed October 16, 2015
  5. a b c Rakotoarison, A., A. Crottini, J. Müller, M.-O. Rödel, F. Glaw & M. Vences: Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species. Zootaxa, 3937, pp. 61-89, 2015
  6. Gonçalo M. Rosa et al .: A new phytotelmic species of Platypelis (Microhylidae: Cophylinae) from the Betampona Reserve, eastern Madagascar . SALAMANDRA 50, 4, pp. 201-214, 2014 PDF; 3.49 MB , accessed December 22, 2014.
  7. ^ Mark D. Scherz, Jörn Koehler, Miguel Vences, and Frank Glaw. 2019. A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar. Evolutionary Systematics, 3, pp. 75-83, 2019 doi : 10.3897 / evolsyst.3.33417 .
  8. A. Rakotoarison, MD Scherz, M. Bletz, JH Razafindraibe, Frank Glaw & Miguel Vences: Description of the lucky Cophyla (Microhylidae, Cophylinae), a new arboreal frog from Marojejy National Park in north-eastern Madagascar. Zootaxa, 4651, pp. 271-288, 2019
  9. F. Glaw, MD Scherz, A. Rakotoarison, A. Crottini, AP Raselimanana, F. Andreone, J. Köhler & M. Vences: Genetic variability and partial integrative revision of Platypelis frogs (Microhylidae) with red flash marks from eastern Madagascar . Vertebrate Zoology, Senckenberg, 70, pp. 141-156, 2020 doi : 10.26049 / VZ70-2-2020-04 .
  10. A. Rakotoarison, MD Scherz, J. Köhler, FM Ratsoavina, O. Hawlitschek, S. Megson, M. Vences & F. Glaw: Frogs of the genus Platypelis from the Sorata massif in northern Madagascar: description of a new species and reports of range extensions. Zoosystematics and Evolution, Museum für Naturkunde, 96, pp. 263–274, Berlin 2020 doi : 10.3897 / zse.96.47088

Web links

Commons : Platypelis  - collection of images, videos and audio files