Terebellomorpha
Terebellomorpha | ||||||||||||
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Scientific name | ||||||||||||
Terebellomorpha | ||||||||||||
Hatschek , 1893 |
Terebellomorpha and Terebelliformia two interchangeably used name of mostly as fine categorized or subordination taxon mostly sessile and röhrenbauender , as filter feeders living polychaete (Polychaeta), which are found in oceans worldwide.
features
According to the original description of Berthold Hatschek the Terebellomorpha have usually only at the foremost segments Parapodial - gills . The prostomium is reduced and bears tufts of thread-like buccal tentacles equipped with eyelashes (in Hatschek "head threads" and in Carl Claus " Fühler "). The throat ( pharynx ) cannot be turned forward. The parapodia are two-branched, so "divided into two bristle humps".
The body of the terebelliformia is divided on the basis of external morphological features into a thorax, on whose segments the parapodia are equipped with bristle-bearing notopodia and neuropodia, and an abdomen, on whose segments only neuropodia with bristles are located. While notopodia are typically absent on the abdomen, there are some forms in which there are bristle-less notopodia on the abdomen that are reduced to short papillae. In the Pectinariidae , the body consists of three sections: a thorax with two bristle-free and gill-free segments, two gill-bearing segments and three segments with notopodia only, an abdomen with both bristle-bearing notopodia and bristle-bearing neuropodia as well as a suction cup-shaped end-scaphe, a bowl-shaped, Structure formed from the last five fused segments and laterally covered with spiky bristles. According to Nogueira, Hutchings and Fukuda (2010), however, the first two sections mentioned are homologous with the thorax, whereas the end scaphe is homologous with the abdomen of the remaining terebelliformia.
Systematics
In 1893, Berthold Hatschek described the Terebellomorpha (in addition to the Spiomorpha, Amphinomorpha, Nereimorpha, Drilomorpha and Serpulimorpha) in his system of annelids as one of six suborders within the order of the Polychaeta and included the Terebellidae and the Grube Amphictenida , which was later synonymous with the Pectinariidae , 1850 . Rodney Phillips Dales established the more extensive order Terebellida in 1962 , in which he included the family Ampharetidae in addition to the Pectinariidae and Terebellidae . Gregory W. Rouse and Kristian Fauchald introduced in 1998 in the subordination Terebellida within the order Canalipalpata also previously by Kristian Fauchald within the order of Spionida as Cirratuliformia summarized families Cirratulidae and Acrocirridae with which they as common features the sharp fillet membrane and the production of blood cells have in the dorsal blood vessel . Struck, Golombek and others (2015), on the other hand, see the terebelliformia on the basis of phylogenetic studies on a molecular genetic basis as a sister group of the Arenicolidae , with which they in turn form a sister group to the clitellata , and find no close relationship to the cirratuliformia.
According to the extent of the terebelliformia also mentioned by Nogueira, Hutchings and Fukuda (2010), Geoffrey Read and Kristian Fauchald 2018 included the following families in the suborder Terebellomorpha within the order Terebellida :
- Alvinellidae Desbruyères & Laubier, 1986
- Ampharetidae Malmgren, 1866
- Pectinariidae Quatrefages, 1866
- Terebellidae Johnston, 1846
- Trichobranchidae Malmgren, 1866
literature
- Berthold Hatschek (1893): System der Anneliden, a preliminary report. Meeting reports of the German Natural Science and Medicine Association for Bohemia "Lotos" in Prague. New episode 13, pp. 123–126 ( download pdf ).
- Carl Claus, Karl Grobben, Alfred Kühn: Textbook of Zoology. Published by Julius Springer, Berlin and Vienna 1932, p. 551.
- Stanley J. Edmonds: Fauna of Australia, Volume 4A. Polychaetes & Allies. The Southern Synthesis 4. Commonwealth of Australia, 2000. Class Polychaeta. Pp. 271f., Terebelliformia.
- Gregory W. Rouse, Kristian Fauchald (1998): Recent views on the status, delineation, and classification of the Annelida. (PDF). American Zoologist. 38 (6), pp. 953-964. doi: 10.1093 / icb / 38.6.953
- Peter Ax: The system of Metazoa II. A textbook on phylogenetic systematics. Gustav Fischer Verlag, Stuttgart / Jena 1999. pp. 47–56, chapter Polychaeta : Terebellida , p. 53.
- João Miguel de Matos Nogueira, Pat A. Hutchings, Marcelo Veronesi Fukuda (2010): Morphology of terebelliform polychaetes (Annelida: Polychaeta: Terebelliformia), with a focus on Terebellidae. Zootaxa 2460 (2460), pp. 1-185.
- Torsten Hugo Struck, Anja Golombek, Anne Weigert, Franziska Anni Franke, Wilfried Westheide, Günter Purschke, Christoph Bleidorn, Kenneth Michael Halanych (2015): The Evolution of Annelids Reveals Two Adaptive Routes to the Interstitial Realm Current Biology. Current Biology 25 (15), pp. 1993-1999. DOI: 10.1016 / j.cub.2015.06.007
- Anne Weigert, Christoph Bleidorn (2016), Current status of annelid phylogeny. Organisms Diversity and Evolution 16 (2), pp. 345-362. DOI: 10.1007 / s13127-016-0265-7
Individual evidence
- ↑ Geoffrey Read, Kristian Fauchald (Ed.) (2018): Terebellomorpha Hatschek, 1893. WoRMS , 2018. Accessed December 9, 2018.