Dinematichthyidae
Dinematichthyidae | ||||||||||||
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Dinematichthyidae | ||||||||||||
Whitley , 1926 |
The Dinematichthyidae are bony fish (Osteichthyes) from the order of the intestinal fish-like (Ophidiiformes). The family wasn't introduced until April 2016. Before that, the species of the family formed the tribe Dinematichthyini, within the livebearers Brotulas (Bythitidae). Most of the species in the family live tucked away in reefs, and a few live in fresh water. The freshwater species often inhabit cave systems such as Typhliasina pearsei on the Yucatán Peninsula and Ogilbia galapagosensis on the Galapagos Island of Santa Cruz .
features
The species of Dinematichthyidae reach standard lengths of 22 cm to almost two meters. Like all gut fish-like, they are elongated. The number of vertebrae is 36 to 87, of which 10 to 25 are trunk vertebrae and 25 to 65 are tail vertebrae. The dorsal and anal fins are long. The caudal fin is free and has not grown together with the dorsal and anal fin to form a continuous fin edge. The first fin support of the anal fin is more or less elongated.
- Fin formula : dorsal 56–191, anal 40–124, pectoral 14–28, ventral 1.
The sensory pores on the head are usually not reduced. There are 6 pores on the lower jaw, 2 to 4 on the preoperculum , 5 to 7 on the infraorbitalia (eye ring bones) and 3 to 4 on the supraorbitalia (skull bones above the eyes). The rearmost supraorbital pore is tubular. The fish have up to seven extended gill rakes . The male's reproductive organ, which consists of a penis-like organ and one to two (less often three) pairs of pseudoclases , lies in a fold of the skin of the abdomen and is covered by a fleshy hood, is a diagnostic feature of the family .
Genera and species
A total of 25 genera with 114 species belong to the Dinematichthyidae. Many of these species have only been described in recent years .
- Genus Alionematichthys Møller & Schwarzhans, 2008
- Alionematichthys ceylonensis Møller & Schwarzhans, 2008
- Alionematichthys crassiceps Møller & Schwarzhans, 2008
- Alionematichthys minyomma (Sedor & Cohen, 1987)
- Alionematichthys phuketensis Møller & Schwarzhans, 2008
- Alionematichthys piger (Alcock, 1890)
- Alionematichthys plicatosurculus Møller & Schwarzhans, 2008
- Alionematichthys riukiuensis (Aoyagi, 1954)
- Alionematichthys samoaensis Møller & Schwarzhans, 2008
- Alionematichthys shinoharai Møller & Schwarzhans, 2008
- Alionematichthys suluensis Møller & Schwarzhans, 2008
- Alionematichthys winterbottomi Møller & Schwarzhans, 2008
- Genus Beaglichhys Machida, 1993
- Beaglichhys bleekeri Schwarzhans & Møller, 2007
- Beaglichhys larsonae Schwarzhans & Møller, 2007
- Beaglichhys macrophthalmus Machida, 1993
- Genus Brosmolus Machida, 1993
- Brosmolus longicaudus Machida, 1993
- Genus Brotulinella Schwarzhans, Møller & Nielsen, 2005
- Brotulinella taiwanensis Schwarzhans, Møller & Nielsen, 2005
- Genus Dactylosurculus Schwarzhans & Møller, 2007
- Dactylosurculus gomoni Schwarzhans & Møller, 2007
- Genus Dermatopsis Ogilby, 1896
- Dermatopsis greenfieldi Moller & Schwarzhans, 2006
- Dermatopsis hoesei Moller & Schwarzhans, 2006
- Dermatopsis joergennielseni Moller & Schwarzhans, 2006
- Dermatopsis macrodon Ogilby , 1896
- Genus Dermatopsoides Smith, 1948
- Dermatopsoides andersoni Moller & Schwarzhans, 2006
- Dermatopsoides kasougae (Smith, 1943)
- Dermatopsoides morrisonae Moller & Schwarzhans, 2006
- Dermatopsoides talboti Cohen, 1966
- Genus Diancistrus Ogilby 1899
- Diancistrus alatus Schwarzhans, Møller & Nielsen, 2005
- Diancistrus alleni Schwarzhans, Møller & Nielsen, 2005
- Diancistrus altidorsalis Schwarzhans, Møller & Nielsen, 2005
- Diancistrus atollorum Schwarzhans, Møller & Nielsen, 2005
- Diancistrus beateae Schwarzhans, Møller & Nielsen, 2005
- Diancistrus brevirostris Schwarzhans, Møller & Nielsen, 2005
- Diancistrus eremitus Schwarzhans, Møller & Nielsen, 2005
- Diancistrus erythraeus (Fowler, 1946)
- Diancistrus fijiensis Schwarzhans, Møller & Nielsen, 2005
- Diancistrus fuscus (Fowler, 1946)
- Diancistrus jackrandalli Schwarzhans, Møller & Nielsen, 2005
- Diancistrus jeffjohnsoni Schwarzhans, Møller & Nielsen, 2005
- Diancistrus karinae Schwarzhans, Møller & Nielsen, 2005
- Diancistrus katrineae Schwarzhans, Møller & Nielsen, 2005
- Diancistrus leisi Schwarzhans, Møller & Nielsen, 2005
- Diancistrus longifilis Ogilby, 1899
- Diancistrus machidai Schwarzhans, Møller & Nielsen, 2005
- Diancistrus manciporus Schwarzhans, Møller & Nielsen, 2005
- Diancistrus mcgroutheri Schwarzhans, Møller & Nielsen, 2005
- Diancistrus mennei Schwarzhans, Møller & Nielsen, 2005
- Diancistrus niger Schwarzhans, Møller & Nielsen, 2005
- Diancistrus novaeguineae (Machida, 1996)
- Diancistrus pohnpeiensis Schwarzhans, Møller & Nielsen, 2005
- Diancistrus robustus Schwarzhans, Møller & Nielsen, 2005
- Diancistrus springeri Schwarzhans, Møller & Nielsen, 2005
- Diancistrus tongaensis Schwarzhans, Møller & Nielsen, 2005
- Diancistrus vietnamensis Schwarzhans, Møller & Nielsen, 2005
- Genus Didymothallus Schwarzhans & Møller, 2007
- Didymothallus criniceps Schwarzhans & Møller, 2007
- Didymothallus mizolepis (Günther, 1867)
- Didymothallus nudigena Schwarzhans & Møller, 2011
- Didymothallus pruvosti Schwarzhans & Møller, 2007
- Genus Dinematichthys Bleeker, 1855
- Dinematichthys dasyrhynchus Cohen & Hutchins, 1982
- Dinematichthys iluocoeteoides Bleeker , 1855
- Dinematichthys indicus Machida, 1994
- Dinematichthys megasoma Machida, 1994
- Dinematichthys minyomma Sedor & Cohen, 1987
- Dinematichthys randalli Machida, 1994
- Dinematichthys riukiuensis Aoyagi, 1954
- Dinematichthys trilobatus Møller & Schwarzhans, 2008
- Genus Dipulus Waite, 1905
- Dipulus caecus Waite, 1905
- Dipulus hutchinsi Moller & Schwarzhans, 2006
- Dipulus multiradiatus (McCulloch & Waite, 1918)
- Dipulus norfolkanus Machida, 1993
- Genus Eusurculus Schwarzhans & Møller, 2007
- Eusurculus andamanensis Schwarzhans & Møller, 2007
- Eusurculus pistillum Schwarzhans & Møller, 2007
- Eusurculus pristinus Schwarzhans & Møller, 2007
- Genus Gunterichthys Dawson, 1966
- Gunterichthys bussingi Møller, Schwarzhans & Nielsen, 2004
- Gunterichthys coheni Møller, Schwarzhans & Nielsen, 2004
- Gunterichthys longipenis Dawson, 1966
- Genus Lapitaichthys
- Lapitaichthys frickei Schwarzhans & Møller, 2007
- Genus Majungaichthys Schwarzhans & Møller, 2007
- Majungaichthys agalegae Schwarzhans & Møller, 2011
- Majungaichthys simplex Schwarzhans & Møller, 2007
- Genus Mascarenichthys Schwarzhans & Møller, 2007
- Mascarenichthys heemstrai Schwarzhans & Møller, 2007
- Mascarenichthys microphthalmus Schwarzhans & Møller, 2007
- Mascarenichthys remotus Schwarzhans & Møller, 2011
- Genus Monothrix Ogilby, 1897
- Monothrix polylepis Ogilby, 1897
- Genus Nielsenichthys Schwarzhans & Møller, 2011
- Nielsenichthys pullus Schwarzhans & Møller, 2011
- Genus Ogilbia Jordan & Evermann 1898
- Ogilbia boehlkei Møller, Schwarzhans & Nielsen, 2005
- Ogilbia boydwalkeri Møller, Schwarzhans & Nielsen, 2005
- Ogilbia cayorum Evermann & Kendall, 1898
- Ogilbia cocoensis Møller, Schwarzhans & Nielsen, 2005
- Ogilbia davidsmithi Møller, Schwarzhans & Nielsen, 2005
- Ogilbia deroyi (Poll & van Mol, 1966)
- Ogilbia galapagosensis (Poll & LeLeup, 1965)
- Ogilbia jeffwilliamsi Møller, Schwarzhans & Nielsen, 2005
- Ogilbia jewettae Møller, Schwarzhans & Nielsen, 2005
- Ogilbia mccoskeri Møller, Schwarzhans & Nielsen, 2005
- Ogilbia nigromarginata Møller, Schwarzhans & Nielsen, 2005
- Ogilbia nudiceps Møller, Schwarzhans & Nielsen, 2005
- Ogilbia robertsoni Møller, Schwarzhans & Nielsen, 2005
- Ogilbia sabaji Møller, Schwarzhans & Nielsen, 2005
- Ogilbia sedorae Møller, Schwarzhans & Nielsen, 2005
- Ogilbia suarezae Møller, Schwarzhans & Nielsen, 2005
- Ogilbia tyleri Møller, Schwarzhans & Nielsen, 2005
- Ogilbia ventralis (Gill, 1863)
- Genus Ogilbichthys Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys ferocis Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys haitiensis Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys kakuki Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys longimanus Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys microphthalmus Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys puertoricoensis Møller, Schwarzhans & Nielsen, 2004
- Ogilbichthys tobagoensis Møller, Schwarzhans & Nielsen, 2004
- Genus Paradiancistrus Schwarzhans, Møller & Nielsen, 2005
- Paradiancistrus acutirostris Schwarzhans, Møller & Nielsen, 2005
- Paradiancistrus christmasensis Schwarzhans & Møller, 2011
- Paradiancistrus cuyoensis Schwarzhans, Møller & Nielsen, 2005
- Paradiancistrus lombokensis Schwarzhans & Møller, 2007
- Genus Porocephalichthys Møller & Schwarzhans, 2008
- Porocephalichthys dasyrhynchus (Cohen & Hutchins, 1982)
- Genus Pseudogilbia Møller, Schwarzhans & Nielsen, 2004
- Pseudogilbia sanblasensis Møller, Schwarzhans & Nielsen, 2004
- Genus Typhlias Whitley, 1951
- Typhlias pearsei Hubbs, 1938
- Genus Ungusurculus Schwarzhans & Møller, 2007
- Ungusurculus collettei Schwarzhans & Møller, 2007
- Ungusurculus komodoensis Schwarzhans & Møller, 2007
- Ungusurculus philippinensis Schwarzhans & Møller, 2007
- Ungusurculus riauensis Schwarzhans & Møller, 2007
- Ungusurculus sundaensis Schwarzhans & Møller, 2007
- Ungusurculus williamsis Schwarzhans & Møller, 2007
- Genus Zephyrichthys Schwarzhans & Møller, 2007
- Zephyrichthys barryi Schwarzhans & Møller, 2007
literature
- Peter Rask Møller, Steen Wilhelm Knudsen, Werner Schwarzhans, Jørgen G. Nielsen: A new classification of viviparous brotulas (Bythitidae) - with the establishment of a new family Dinematichthyidae - based on molecular, morphological and fossil data. Molecular Phylogenetics and Evolution, April 2016, doi: 10.1016 / j.ympev.2016.04.008