Lip-toothed gobies
Lip-toothed gobies | ||||||||||||
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![]() Sicyopterus fasciatus |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Sicydiinae | ||||||||||||
Gill , 1860 |
Sicydiinae (Sicydiinae) are a sub-family of Grundelartigen (Gobiiformes). They live worldwide in fresh and brackish waters on tropical islands and in fast-flowing rivers and streams in coastal mountain regions up to heights of 1000 meters, in West Africa, South and Southeast Asia, Australia and in tropical America. Some species are adapted to life in fast flowing streams and rapids. The fish are therefore also called climbing gobies. The larvae often develop in the sea.
Lip-toothed gobies feed on growth, diatoms , green algae , blue-green algae and bacteria. Sicyopus species feed on small invertebrates.
features
The fish have a cylindrical body that is often incompletely scaled. Some species have scales only on the tail stalk. The mouth is terminal or below , the lower jaw shorter than the upper jaw. The eyes are wide apart on the top of the head. In the hard-edged first dorsal fin , the middle of the mostly 6 fin rays are higher and are often drawn out like filaments. The pelvic fins have grown together to form a suction funnel and are supported by thick, highly branched fin rays. Except for the genera Sicyopus and Stiphodon , they are fused with the abdominal wall. The species Sicyopterus stimpsoni can generate negative pressure in two places (belly and mouth) and thus stick to the ground even when moving. In contrast, other lip-toothed gobies have to briefly give up the firm surface adhesion in order to move.
Male lip-toothed gobies are often more colorful than the plainly colored females and have larger backs and anal fins . Especially during courtship the territorial males shimmer red and / or metallic.
Reproduction
The fish spawn hidden under stones and in crevices, often after rainfall, so that the larvae are washed into the sea. The clutches are very large, a 10 cm long female can lay up to 200,000 eggs. Late larval stages migrate back into the rivers, often during the new or full moon to take advantage of the spring tide. The contact with fresh water triggers the metamorphosis to the young fish.
Species and genera
There are over 120 species from 9 genera.
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Akihito Watson, Keith & Marquet 2007
- Akihito futuna Keith, Marquet & Watson, 2008
- Akihito vanuatu Watson, Keith & Marquet, 2007
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Cotylopus Guichenot, 1863
- Cotylopus acutipinnis Guichenot 1863
- Cotylopus rubripinnis Keith, Hoareau & Bosc 2005
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Lentipes Günther, 1861
- Lentipes argenteus Keith, Hadiaty & Lord, 2014
- Lentipes caroline Lynch et al., 2013
- Lentipes concolor Gill, 1860
- Lentipes crittersius Watson & Allen, 1999
- Lentipes ikeae Keith, Hadiaty & Lord, 2014
- Lentipes kaaea Watson, Keith & Marquet, 2002
- Lentipes mekonggaensis Keith, Hadiaty & Lord, 2014
- Lentipes multiradiatus Allen, 2001
- Lentipes watsoni Allen, 1997
- Lentipes whittenorum Watson & Kottelat, 1994
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Parasicydium
- Parasicydium bandama Risch, 1980
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Sicydium Valenciennes in Cuvier & Valenciennes, 1837
- Sicydium adelum Bussing, 1996
- Sicydium altum Meek, 1907
- Sicydium antillarum Ogilvie-Grant, 1884
- Sicydium brevifile Ogilvie-Grant, 1884
- Sicydium buscki Evermann & Clark, 1906
- Sicydium cocoensis (Heller & Snodgrass, 1903)
- Sicydium crenilabrum Harrison, 1993
- Sicydium fayae Brock, 1942
- Sicydium gilberti Watson, 2000
- Sicydium gymnogaster Ogilvie-Grant, 1884
- Sicydium hildebrandi Eigenmann, 1918
- Sicydium multipunctatum Regan, 1906
- Sicydium plumieri (Bloch, 1786)
- Sicydium punctatum Perugia, 1896
- Sicydium rosenbergii (Boulenger, 1899)
- Sicydium salvini Ogilvie-Grant, 1884
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Sicyopterus Gill, 1860
- Sicyopterus aiensis Keith, Watson & Marquet, 2004
- Sicyopterus brevis de Beaufort, 1912
- Sicyopterus caeruleus (Lacepède, 1800)
- Sicyopterus caudimaculatus Maugé, Marquet & Laboute, 1992
- Sicyopterus crassus Herre, 1927
- Sicyopterus cynocephalus (Valenciennes, 1837)
- Sicyopterus eudentatus Parenti & Maciolek, 1993
- Sicyopterus fasciatus (Day, 1874)
- Sicyopterus franouxi (Pellegrin, 1935)
- Sicyopterus fuliag Herre, 1927
- Sicyopterus griseus (Day, 1877)
- Sicyopterus hageni Popta, 1921
- Sicyopterus japonicus (Tanaka, 1909)
- Sicyopterus lacrymosus Herre, 1927
- Sicyopterus lagocephalus (Pallas, 1770)
- Sicyopterus laticeps (Valenciennes, 1837)
- Sicyopterus lividus Parenti & Maciolek, 1993
- Sicyopterus longifilis de Beaufort, 1912
- Sicyopterus macrostetholepis (Bleeker, 1853)
- Sicyopterus marquesensis Fowler, 1932
- Sicyopterus microcephalus (Bleeker, 1854)
- Sicyopterus micrurus (Bleeker, 1853)
- Sicyopterus ouwensi Weber, 1913
- Sicyopterus panayensis Herre, 1927
- Sicyopterus parvei (Bleeker, 1853)
- Sicyopterus pugnans (Ogilvie-Grant, 1884)
- Sicyopterus punctissimus Sparks & Nelson, 2004
- Sicyopterus rapa Parenti & Maciolek, 1996
- Sicyopterus sarasini Weber & de Beaufort, 1915
- Sicyopterus squamosissimus Keith et al., 2015
- Sicyopterus stimpsoni (Gill, 1860)
- Sicyopterus wichmanni (Weber, 1894)
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Sicyopus Gill, 1860
- Sicyopus auxilimentus Watson & Kottelat, 1994
- Sicyopus cebuensis Chen & Shao, 1998
- Sicyopus discordipinnis Watson, 1995
- Sicyopus jonklaasi (Axelrod, 1972)
- Sicyopus multisquamatus de Beaufort, 1912
- Sicyopus nigriradiatus Parenti & Maciolek, 1993
- Sicyopus rubicundus Keith, P., Hadiaty, Busson & Hubert, 2014
- Sicyopus zosterophorum (Bleeker, 1857)
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Smilosicyopus
- Smilosicyopus bitaeniatus (Maugé, Marquet & Laboute, 1992)
- Smilosicyopus chloe (Watson, Keith & Marquet, 2001)
- Smilosicyopus Fehlmanni (Parenti & Maciolek, 1993)
- Smilosicyopus leprurus (Sakai & Nakamura, 1979)
- Smilosicyopus mystax (Watson & Allen, 1999)
- Smilosicyopus pentecost (Keith, Lord & Taillebois, 2010)
- Smilosicyopus sasali (Keith & Marquet, 2005)
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Stiphodon Weber, 1895
- Stiphodon allen Watson, 1996
- Stiphodon annieae Keith & Hadiaty, 2014
- Stiphodon astilbos Ryan, 1986
- Stiphodon atratus Watson, 1996
- Stiphodon atropurpureus (Herre, 1927)
- Stiphodon aureofuscus Keith et al., 2015
- Stiphodon birdsong Watson, 1996
- Stiphodon caeruleus Parenti & Maciolek, 1993
- Stiphodon carisa Watson, 2008
- Stiphodon discotorquatus Watson, 1995
- Stiphodon elegans (Steindachner, 1879)
- Stiphodon hydroreibatus Watson, 1999
- Stiphodon imperiorientis Watson & Chen, 1998
- Stiphodon julieni Keith, Watson & Marquet, 2002
- Stiphodon kalfatak Keith, Marquet & Watson, 2007
- Stiphodon larson Watson, 1996
- Stiphodon martenstyni Watson, 1998
- Stiphodon maculidorsalis Maeda & Tan, 2013
- Stiphodon mele Keith, Marquet & Pouilly, 2009
- Stiphodon multisquamus Wu & Ni, 1986
- Stiphodon niraikanaiensis Maeda, 2013
- Stiphodon oatea Keith, Feunteun & Vigneux, 2009
- Stiphodon olivaceus Watson & Kottelat, 1995
- Stiphodon ornatus Meinken, 1974
- Stiphodon palawanensis Maeda & Palla, 2015
- Stiphodon pelewensis Herre, 1936
- Stiphodon percnopterygionus Watson & Chen, 1998
- Stiphodon rubromaculatus Keith & Marquet, 2007
- Stiphodon rutilaureus Watson, 1996
- Stiphodon sapphirinus Watson, Keith & Marquet, 2005
- Stiphodon semoni Weber, 1895
- Stiphodon surrufus Watson & Kottelat, 1995
- Stiphodon tuivi Watson, 1995
- Stiphodon weberi Watson, Allen & Kottelat, 1998
- Stiphodon zebrinus Watson, Allen & Kottelat, 1998
literature
- Joseph S. Nelson : Fishes of the World. 4th edition. John Wiley & Sons, Hoboken NJ 2006, ISBN 0-471-25031-7 .
- Hans Horsthemke: Sicydiinae. In: Claus Schaefer, Torsten Schröer (Hrsg.): The large lexicon of aquaristics. Eugen Ulmer, Stuttgart 2004, ISBN 3-8001-7497-9 , pp. 888 f.
Web links
- Lip-toothed gobies on Fishbase.org (English)
proof
- ↑ Michael Stang : Diverse fish mouths - gobies on Hawaii use their mouths for climbing. In: Deutschlandfunk . Retrieved January 7, 2013 .