Melittidae

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Melittidae
Melitta nigricans male

Melitta nigricans male

Systematics
Order : Hymenoptera (Hymenoptera)
Subordination : Waist Wasps (Apocrita)
Superfamily : Apoidea
Bees (Apiformes)
Family : Melittidae
Scientific name
Melittidae
Michener , 2000
Subfamilies

Meganomiinae
Melittinae
Dasypodainae

The Melittidae is a small family of bees . About 200 species and 14 genera are known. All species are solitary and nest in the ground. Many of the species are oligolectic, that is, they specialize in certain types of plants. A peculiarity of the Melittidae is that "collecting oil" occurs in this family. Species of the genera Rediviva and Macropis have special hairs on the front and middle legs with which they collect oil, which the larvae deposit in the nests for pollen as food. The oil comes from flowers, e.g. B. of Lysimachia , in elaiophores secrete oil. An extinct species ( Paleomacropis eocenicus ) that was found in amber also had hair that was probably used to collect oil.

features

The Melittidae are similar to the Andrenidae . They are short-tongued bees and leg collectors like these . Unlike most Andrenidae, however, they have reduced scopae on the femur to the coxa . The scopae lie on the tibia and basitarsus . The Melittidae generally have quite short, pointed tongues and unspecialized labial palps , the first two limbs of which are quite short. Their middle coxae are similar to those of the long-tongued bees. There is no characteristic that is common to all Melittidae and cannot be found in any other bee family. The Meganomiinae are large species over 17 mm.

distribution

The species of the family are Palearctic and common in southern Africa . The greatest diversity is found in southern Africa. The family is rare in North America, only Hesperapis is quite common here in the western United States and Mexico. Species of Dasypodainae and Melittinae occur in the Palearctic area. The Meganomiinae are restricted to Africa. No Melittidae are known in South America, Australia and Antarctica.

Systematics

External system

The systematic position of the Melittidae is not yet clearly understood. Traditionally and by several authors, they are regarded as the sister group of the "long-tongued bees" (see also cladogram below). There are molecular biological studies according to which the Melittidae represent the most basic bee family, i.e. the sister group of all other bees, and it is discussed whether the Melittidae are monophyletic.

 Anthophila bees 

 long-tongued bees 

Apidae Apis mellifera Western honey bee.jpg


   

Megachilidae Leafcutter bee (Megachile sp.) Collecting leaves (7519316920) .jpg



   

Melittidae Macropis europaea4.jpg



 short-tongued bees 


Halictidae narrow and furrow bees ˜Iridescent.green.sweat.bee1.jpg


   

Colletidae Colletes cunicularius m1.JPG


   

Stenotritidae Stenotritus pubescens, f, side, australia 2014-07-05-12.18.33 ZS PMax.jpg




   

Andrenidae Andrena haemorrhoa f20140330.30 17-39-10.jpg




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Internal system

The following is the internal system of the Melittidae according to Michez et al. (2009).

 Melittidae 
 Meganomiinae 

Ceratomonia Michener, 1981, Namibia


   

Meganomia Cockerell, 1909, Ethiopia


   

Pseudophilanthus Alfken, 1939, East Africa (including Madagascar)


   

Uromonia Michener, 1981, East Africa (including Madagascar)


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 Melittinae 
 Macropini 

Promelitta Warncke, 1977, North Africa


   

Afrodasypoda Engel, 2005, South Africa


   

Macropis Klug, 1906, Holarctic


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 Melttini 

Rediviva Friese, 1911, South Africa


   

Redivivoides Michener, 1981, South Africa


   

Sawhorn bees ( Melitta Kirby, 1802), Holarctic and Africa


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 Dasypodainae 
 Hesperapini 

Eremaphanta Popov, 1940, Central Asia


   

Capicola Friese, 1911, South Africa


   

Hesperapis Cockerell, 1898, near-table


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 Dasypodaini 

Haplomelitta Cockerell, 1934, South Africa


   

Dasypoda Latreille, 1802, Palearctic


   

Samba friezes, 1908, Kenya


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Individual evidence

  1. ^ A b c John Plant & Hannes F. Paulus : Evolution and phylogeny of bees review and cladistic analysis in light of morphological evidence (Hymenoptera, Apoidea) . Schweizerbart Science Publishers, Stuttgart 2016, ISBN 978-3-510-55048-7 .
  2. a b c d Denis Michez, Sébastien Patiny, Bryan N. Danforth (2009): Phylogeny of the bee family Melittidae (Hymenoptera: Anthophila) based on combined molecular and morphological data. Systematic Entomology 34: 574-597. [1]
  3. Ch. D. Michener: Bees of the World . 2007, ISBN 978-0-8018-8573-0 , pp. 972 ( com.ua [PDF]).

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