Ulmus alata: Difference between revisions

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{{Short description|Species of tree}}
{{Use dmy dates|date=December 2014}}
{{Use dmy dates|date=September 2023}}
{{italic title}}
{{Speciesbox
{{Taxobox
| image = Ulmus alata 071007c.jpg
| name = Winged elm
| image_caption = Leaves of ''U. alata''
| image = Ulmus alata 071007c.jpg
| status = LC
| image_caption = Leaves
| status_system = IUCN3.1
| regnum = [[Plant]]ae
| status_ref = <ref>{{cite iucn |author=Stritch, L. |year=2019 |title=Ulmus alata |amends=2019 |volume=2019 |page=e.T61966604A180056609 |access-date=12 April 2021}}</ref>
| unranked_divisio = [[Angiosperms]]
| genus = Ulmus
| unranked_classis = [[Eudicots]]
| display_parents = 2
| unranked_ordo = [[Rosids]]
| parent = Ulmus sect. Chaetoptelea
| ordo = [[Rosales]]
| species = alata
| familia = [[Ulmaceae]]
| authority = [[André Michaux|Michx.]]
| genus = ''[[Ulmus]]''
| range_map = Ulmus alata range map 2.png
| species = '''''U. alata'''''
| range_map_caption = Natural range
| binomial = ''Ulmus alata''
| synonyms = *''Ulmus pumila'' <small>Walter</small>
| binomial_authority = [[André Michaux|Michx.]]
| range_map = Ulmus alata range map 2.png
| range_map_caption = Natural range
| synonyms = *''Ulmus pumila'' <small>('''not''' L.) Walter </small>
}}
}}

'''''Ulmus alata''''', the '''winged elm''' or '''wahoo''', is a small- to medium-sized [[deciduous]] tree [[endemicity|endemic]] to the woodlands of the southeastern and south-central United States. The species is tolerant of a wide range of soils, and of ponding, but is the least shade-tolerant of the North American elms. Its growth rate is often very slow, the trunk increasing in diameter by less than {{convert|5|mm|in|frac=32|abbr=on}} per year. The tree is occasionally considered a nuisance as it readily invades old fields, forest clearings, and rangelands, proving particularly difficult to eradicate with herbicides.<ref name=University>University of Florida, Environmental Horticulture Department (1994). ''Fact Sheet ST-648''. Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences.</ref>
'''''Ulmus alata''''', the '''winged elm''' or '''wahoo''', is a small- to medium-sized [[deciduous]] tree [[endemicity|endemic]] to the woodlands of the southeastern and south-central United States. The species is tolerant of a wide range of soils, and of ponding, but is the least shade-tolerant of the North American elms. Its growth rate is often very slow, the trunk increasing in diameter by less than {{convert|5|mm|in|frac=32|abbr=on}} per year. The tree is occasionally considered a nuisance as it readily invades old fields, forest clearings, and rangelands, proving particularly difficult to eradicate with herbicides.<ref name=University>University of Florida, Environmental Horticulture Department (1994). ''Fact Sheet ST-648''. Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences.</ref>


== Description ==
== Description ==
As its common and scientific<ref>''Alata'' is [[Latin]] for "winged".</ref> names imply, winged elm is most easily recognized by the very broad, thin pair of corky wings that form along the branchlets after a couple of years. The tree generally grows to a maximum height and breadth of about {{convert|13|×|13|m|ft|abbr=on}}, although on fertile alluvial soils such as those of the [[Mississippi Delta|Mississippi River Delta]], some specimens have reached double this height (see 'Notable trees' below). The crown can be either rounded or pyramidal; the branches are pendulous.<ref>{{cite web |url=http://www.ces.ncsu.edu/depts/hort/consumer/factsheets/trees-new/ulmus_alata.html |title=Trees: Ulmus alata |website=www.ces.ncsu.edu |url-status=dead |archive-url=https://web.archive.org/web/20040803215625/http://www.ces.ncsu.edu/depts/hort/consumer/factsheets/trees-new/ulmus_alata.html |archive-date=2004-08-03}}</ref> The leaves are comparatively small for the genus, less than {{convert|6.5|cm|in|frac=8|abbr=on}} long and less than {{convert|2.0|cm|in|frac=4|abbr=on}} broad, [[oblong-lanceolate]] to narrowly [[Elliptic leaf|elliptic]], thin in texture, and smooth above with serrate or doubly serrate margins. The leaves turn bright yellow in Autumn. The wind-pollinated [[perfect flower|perfect]] [[apetalous]] flowers are borne on long [[Pedicel (botany)|pedicels]] in March and April before the leaves appear. The reddish [[Samara (fruit)|samaras]] are also relatively small, less than {{convert|8|mm|in|frac=32|abbr=on}} long, narrowly elliptic with two long incurving stigmas at the tip,<ref>Photo of ''U. alata'' samarae, jimbotany.com/16-Catalog_Ra_through_Z-Ackn-LitCitd.htm, [http://jimbotany.com/ULMUALAT.JPG]</ref> and usually disperse before the end of April.<ref name=Elwes>Elwes, H. J. & Henry, A. (1913). ''[http://fax.libs.uga.edu/QK488xE4/1f/trees_of_britain_and_ireland_vol_7.pdf The Trees of Great Britain & Ireland]''. Vol. VII. 1848–1929. Republished 2014 Cambridge University Press, {{ISBN|978-1-108-06938-0}}</ref><ref name=Schnelle>Schnelle, M. (1999). Field Notes: Ulmus alata. ''American Nurseryman'', page 1998, 1 March 1999. p. 98. [[Chicago]]</ref>
[[File:Winged Elm Ulmus alata 2009-05-10.jpg|thumb|left|The corky wings of ''Ulmus alata'']]
<gallery>
As its common and scientific<ref>''Alata'' is [[Latin]] for "winged".</ref> names imply, winged elm is most easily recognized by the very broad, thin pair of corky wings that form along the branchlets after a couple of years. The tree generally grows to a maximum height and breadth of about {{convert|13|×|13|m|ft|abbr=on}}, although on fertile alluvial soils such as those of the [[Mississippi Delta|Mississippi River Delta]], some specimens have reached double this height (see 'Notable trees' below). The crown can be either rounded or pyramidal; the branches are pendulous.<ref>[http://www.ces.ncsu.edu/depts/hort/consumer/factsheets/trees-new/ulmus_alata.html]</ref>
File:Garden of the Gods - Winged Elm, Wahoo (Ulmus alata) - Flickr - Jay Sturner (1).jpg|Young ''Ulmus alata'', showing corky bark-ridges

File:Garden of the Gods - Winged Elm, Wahoo (Ulmus alata) - Flickr - Jay Sturner.jpg|Leaves of ''Ulmus alata''
The leaves are comparatively small for the genus, less than {{convert|6.5|cm|in|frac=8|abbr=on}} long and less than {{convert|2.0|cm|in|frac=4|abbr=on}} broad, [[oblong-lanceolate]] to narrowly [[elliptic]], thin in texture, and smooth above. The wind-pollinated [[perfect flower|perfect]] [[apetalous]] flowers are borne on long [[Pedicel (botany)|pedicels]] in March and April before the leaves appear. The reddish [[Samara (fruit)|samaras]] are also relatively small, less than {{convert|8|mm|in|frac=32|abbr=on}} long, narrowly elliptic with two long incurving stigmas at the tip,<ref>Photo of ''U. alata'' samarae, jimbotany.com/16-Catalog_Ra_through_Z-Ackn-LitCitd.htm, [http://jimbotany.com/ULMUALAT.JPG]</ref> and usually disperse before the end of April.<ref name=Elwes>Elwes, H. J. & Henry, A. (1913). ''[http://fax.libs.uga.edu/QK488xE4/1f/trees_of_britain_and_ireland_vol_7.pdf The Trees of Great Britain & Ireland]''. Vol. VII. 1848–1929. Republished 2014 Cambridge University Press, ISBN 978-1-108-06938-0</ref><ref name=Schnelle>Schnelle, M. (1999). Field Notes: Ulmus alata. ''American Nurseryman'', page 1998, 1 March 1999. p. 98. [[Chicago]]</ref>
</gallery>


== Pests and diseases ==
== Pests and diseases ==
Line 31: Line 30:


== Cultivation ==
== Cultivation ==
''Ulmus alata'' is rarely cultivated beyond its natural range. It remains in commercial production in the USA, and is occasionally available in Europe; several specimens are also grown in New Zealand.<ref name=Auckland>''Auckland Botanical Society Journal'' (2003). Vol. 58 (1), June 2003. {{ISSN|0113-4132}}</ref>
''Ulmus alata'' is rarely cultivated beyond its natural range. It remains in commercial production in the US, and is occasionally available in Europe. At the beginning of the 19th century, the tree was one of the three American elm species cultivated in ornamental plantations in Britain,<ref name=Main>{{cite book|last1=Main|first1=James|title=The Forest Planter and Pruner's Assistant|date=1839|publisher=Ridgway|location=London|page=113|url=https://archive.org/stream/forestplanterpru00main#page/113/mode/2up}}</ref> but is now rare there. Several specimens are grown in New Zealand.<ref name=Auckland>{{cite journal|first1=Mike|last1=Wilcox|first2=Chris|last2=Inglis|journal=Auckland Botanical Society Journal|title=Auckland's elms|volume=58|issue=1|date=2003|pages=38–45|publisher=Auckland Botanical Society|url=http://bts.nzpcn.org.nz/bts_pdf/ABJ58%281%292003-38-45-Elms.pdf}}</ref>


== Notable trees ==
== Notable trees ==
On the silty uplands of the Mississippi Delta, ''Ulmus alata'' can attain 27&nbsp;m (89&nbsp;ft) in height, although the trunk diameter rarely exceeds 60&nbsp;cm (24&nbsp;in) [[diameter at breast height|d.b.h]]. In the old growth [[Fernbank Forest]] in [[Atlanta, Georgia]], the species attains heights up to {{convert|39|m}}. A tree measuring {{convert|40|m}} high has been reported from the [[Congaree National Park]] in [[South Carolina]].<ref>http://www.nativetreesociety.org/events/congaree2009/NewCongMaxList.xls</ref> However, the USA National Champion, measuring {{convert|27|m}} high in 2009, grows in [[Hopewell County]], Virginia.<ref name=American>American Forests. (2012). The 2012 National Register of Big Trees.</ref>[http://www.americanforests.org/our-programs/bigtree]
On the silty uplands of the Mississippi Delta, ''Ulmus alata'' can attain 27&nbsp;m (89&nbsp;ft) in height, although the trunk diameter rarely exceeds 60&nbsp;cm (24&nbsp;in) [[diameter at breast height|d.b.h]]. In the old growth [[Fernbank Forest]] in [[Atlanta, Georgia]], the species attains heights up to {{convert|39|m}}. A tree measuring {{convert|40|m}} high has been reported from the [[Congaree National Park]] in [[South Carolina]].<ref>[http://www.nativetreesociety.org/events/congaree2009/NewCongMaxList.xls Event list] nativetreesociety.org {{dead link|date=September 2023}}</ref> However, the USA National Champion, measuring {{convert|27|m}} high in 2009, grows in [[Hopewell, Virginia]].<ref name=American>American Forests. (2012). The 2012 National Register of Big Trees.</ref>[http://www.americanforests.org/our-programs/bigtree ]


== Cultivars ==
== Cultivars ==
*[[Lace Parasol (elm cultivar)|Lace Parasol]]
* [[Lace Parasol (elm cultivar)|Lace Parasol]]


== Other uses ==
== Other uses ==
''Ulmus alata'' is of minimal commercial significance, its hard timber considered no more remarkable than that of other American elms, and of limited use because of the commonly small size of the trees. However, owing to its resistance to splitting, it is used to make high-quality [[hockey]] sticks.{{cn|date=October 2015}}
''Ulmus alata'' is of minimal commercial significance, its hard timber considered no more remarkable than that of other American elms, and of limited use because of the commonly small size of the trees. However, owing to its resistance to splitting, it is used to make high-quality [[hockey stick]]s.<ref>{{cite web |url=https://www.srs.fs.usda.gov/pubs/misc/ag_654/volume_2/ulmus/alata.htm |title=''Ulmus alata'' Michx. Winged Elm |last=Snow |first=G. A. |date= |website= |publisher=United States Department of Agriculture. Southern Research Station |access-date=February 2, 2021|quote=}}</ref>


== Accessions ==
== Accessions ==
;North America
;North America
*[[Arnold Arboretum]]. Acc. no. 404-95, wild collected.
* [[Arnold Arboretum]], US. Acc. no. 404-95, wild collected.
*Bartlett Tree Experts<ref>[http://www.bartlett.com]</ref> Acc. no. 1438, unrecorded provenance.
* Bartlett Tree Experts,<ref>{{Cite web|url=https://www.bartlett.com/|title=Bartlett Tree Experts: Tree Services|website=www.bartlett.com}}</ref> US. Acc. no. 1438, unrecorded provenance.
*[[Brooklyn Botanic Garden]],<ref>[http://www.bbg.org/cgi/bgbase/search.cgi]</ref> [[New York City]]. Acc. nos. 730275, X00886
* [[Brooklyn Botanic Garden]],<ref>{{cite web |url=http://www.bbg.org/cgi/bgbase/search.cgi |title=BROOKLYN BOTANIC GARDEN >> Plant Records |website=www.bbg.org |url-status=dead |archive-url=https://web.archive.org/web/20030430105750/http://www.bbg.org/cgi/bgbase/search.cgi |archive-date=2003-04-30}}</ref> [[New York City]], US. Acc. nos. 730275, X00886
*[[Bernheim Arboretum and Research Forest]] [http://www.bernheim.org/species_lists.htm], [[Clermont, Kentucky]]. No details available.
* [[Bernheim Arboretum and Research Forest]] [http://www.bernheim.org/species_lists.htm Bernheim Arboretum and Research Forest - Connecting People with Nature], [[Clermont, Kentucky]], US. No details available.
*[[Morton Arboretum]]. Acc. no. 116-96, wild collected [[Papoose Lake]], [[Illinois]].
* [[Morton Arboretum]], US. Acc. no. 116-96, wild collected [[Papoose Lake]], [[Illinois]].<ref>[https://www.cirrusimage.com/tree_winged_elm/ Photographs of Morton Arboretum ''U. alata'', Acc. no. 116-96; cirrusimage.com]</ref>
;Europe
;Europe
*[[Brighton & Hove]] City Council, UK, [[NCCPG]] Elm Collection.<ref>{{cite web|title=List of plants in the {elm} collection|publisher=Brighton & Hove City Council|access-date=23 September 2016|url=http://www.brighton-hove.gov.uk/content/leisure-and-libraries/parks-and-green-spaces/list-plants-collection}}</ref> One tree at East Brighton Park, UK champion {{convert|13|m|ft|abbr=on}} high, {{convert|31|cm|in|frac=4|abbr=on}} [[d.b.h.]] in 2001.<ref name=Johnson>Johnson, Owen (ed.) (2003). ''Champion Trees of Britain & Ireland''. Whittet Press, ISBN 978-1-873580-61-5.</ref>
* [[Brighton & Hove]] City Council, UK. [[NCCPG]] Elm Collection.<ref>{{cite web|title=List of plants in the {elm} collection|publisher=Brighton & Hove City Council|access-date=23 September 2016|url=http://www.brighton-hove.gov.uk/content/leisure-and-libraries/parks-and-green-spaces/list-plants-collection}}</ref> One tree at East Brighton Park, UK champion {{convert|13|m|ft|abbr=on}} high, {{convert|31|cm|in|frac=4|abbr=on}} [[d.b.h.]] in 2001.<ref name=Johnson>Johnson, Owen (ed.) (2003). ''Champion Trees of Britain & Ireland''. Whittet Press, {{ISBN|978-1-873580-61-5}}.</ref>
*[[Grange Farm Arboretum]], [[Sutton St James]], [[Spalding, Lincolnshire]], UK. Acc. no. 506
* [[Grange Farm Arboretum]], [[Lincolnshire]], UK. Acc. no. 506
*[[Royal Botanic Garden Edinburgh]]. Acc. no. 20080092, from seed wild collected in USA.
* [[Royal Botanic Garden Edinburgh]], UK. Acc. no. 20080092, from seed wild collected in USA.<ref name=Royal>Royal Botanic Garden Edinburgh. (2017). ''List of Living Accessions: Ulmus'' [http://elmer.rbge.org.uk/bgbase/livcol/bgbaselivcol.php?cfg=bgbase/livcol/bgbaselivcol.cfg&startrow=26]</ref>
*[[Thenford]] House, [[Northamptonshire]], UK. No details available.
* [[Thenford]] House, [[Northamptonshire]], UK. No details available.
;Australasia
;Australasia
*Manukau Cemetery & Crematorium, [[Auckland]], New Zealand. No details available.
* Manukau Cemetery & Crematorium, [[Auckland]], New Zealand. No details available.


== Nurseries ==
== Nurseries ==
Line 61: Line 60:
Widely available.
Widely available.
;Europe
;Europe
*[http://www.arboretum-waasland.be/English.htm Arboretum Waasland], [[Nieuwkerken-Waas]], Belgium.
* [http://www.arboretum-waasland.be/English.htm Arboretum Waasland], [[Nieuwkerken-Waas]], Belgium.
*[[Grange Farm Arboretum|Grange Farm Plants]], [[Spalding, Lincolnshire]], UK.
* [[Grange Farm Arboretum|Grange Farm Plants]], [[Spalding, Lincolnshire]], UK.
*[http://www.esveld.nl/voorraadengels/voorrulmus.htm Plantentuin Esveld], Netherlands.
* [http://www.esveld.nl/voorraadengels/voorrulmus.htm Plantentuin Esveld], Netherlands.


=== Seed suppliers ===
=== Seed suppliers ===
*[http://www.sheffields.com/products/simple_search Sheffield's Seeds Co. Inc.], [[New York (state)|New York]], USA.
* [http://www.sheffields.com/products/simple_search Sheffield's Seeds Co. Inc.], [[New York (state)|New York]], US.


== References ==
== References ==
{{reflist}}
{{Reflist}}
*{{Silvics |volume=2 |genus=Ulmus |species=alata |first=G. A. |last=Snow}}


==External links==
* {{Silvics |volume=2 |genus=Ulmus |species=alata |first=G. A. |last=Snow}}
*[http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233501324 ''U. alata'', Flora of North America, www.efloras.org]


{{Elm species, varieties, hybrids, hybrid cultivars and species cultivars |state=collapsed}}
{{Elm species, varieties, hybrids, hybrid cultivars and species cultivars |state=collapsed}}
{{Taxonbar|from=Q7879360}}
{{Authority control}}


[[Category:Elm species and varieties]]
[[Category:Trees of Northern America]]
[[Category:Trees of the United States]]
[[Category:Plants used in bonsai]]
[[Category:Plants used in bonsai]]
[[Category:Trees of the Southeastern United States]]
[[Category:Trees of the North-Central United States]]
[[Category:Trees of the South-Central United States]]
[[Category:Ulmus articles with images]]
[[Category:Ulmus articles with images]]
[[Category:Ulmus]]
[[Category:Ulmus|alata]]

Latest revision as of 10:37, 25 January 2024

Ulmus alata
Leaves of U. alata
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Rosales
Family: Ulmaceae
Genus: Ulmus
Subgenus: U. subg. Oreoptelea
Section: U. sect. Chaetoptelea
Species:
U. alata
Binomial name
Ulmus alata
Natural range
Synonyms
  • Ulmus pumila Walter

Ulmus alata, the winged elm or wahoo, is a small- to medium-sized deciduous tree endemic to the woodlands of the southeastern and south-central United States. The species is tolerant of a wide range of soils, and of ponding, but is the least shade-tolerant of the North American elms. Its growth rate is often very slow, the trunk increasing in diameter by less than 5 mm (316 in) per year. The tree is occasionally considered a nuisance as it readily invades old fields, forest clearings, and rangelands, proving particularly difficult to eradicate with herbicides.[2]

Description[edit]

As its common and scientific[3] names imply, winged elm is most easily recognized by the very broad, thin pair of corky wings that form along the branchlets after a couple of years. The tree generally grows to a maximum height and breadth of about 13 m × 13 m (43 ft × 43 ft), although on fertile alluvial soils such as those of the Mississippi River Delta, some specimens have reached double this height (see 'Notable trees' below). The crown can be either rounded or pyramidal; the branches are pendulous.[4] The leaves are comparatively small for the genus, less than 6.5 cm (2+12 in) long and less than 2.0 cm (34 in) broad, oblong-lanceolate to narrowly elliptic, thin in texture, and smooth above with serrate or doubly serrate margins. The leaves turn bright yellow in Autumn. The wind-pollinated perfect apetalous flowers are borne on long pedicels in March and April before the leaves appear. The reddish samaras are also relatively small, less than 8 mm (516 in) long, narrowly elliptic with two long incurving stigmas at the tip,[5] and usually disperse before the end of April.[6][7]

Pests and diseases[edit]

Like the other North American species of elm, U. alata is very susceptible to Dutch elm disease and Elm Yellows (Elm phloem necrosis).[8]

Cultivation[edit]

Ulmus alata is rarely cultivated beyond its natural range. It remains in commercial production in the US, and is occasionally available in Europe. At the beginning of the 19th century, the tree was one of the three American elm species cultivated in ornamental plantations in Britain,[9] but is now rare there. Several specimens are grown in New Zealand.[10]

Notable trees[edit]

On the silty uplands of the Mississippi Delta, Ulmus alata can attain 27 m (89 ft) in height, although the trunk diameter rarely exceeds 60 cm (24 in) d.b.h. In the old growth Fernbank Forest in Atlanta, Georgia, the species attains heights up to 39 metres (128 ft). A tree measuring 40 metres (130 ft) high has been reported from the Congaree National Park in South Carolina.[11] However, the USA National Champion, measuring 27 metres (89 ft) high in 2009, grows in Hopewell, Virginia.[12][3]

Cultivars[edit]

Other uses[edit]

Ulmus alata is of minimal commercial significance, its hard timber considered no more remarkable than that of other American elms, and of limited use because of the commonly small size of the trees. However, owing to its resistance to splitting, it is used to make high-quality hockey sticks.[13]

Accessions[edit]

North America
Europe
Australasia
  • Manukau Cemetery & Crematorium, Auckland, New Zealand. No details available.

Nurseries[edit]

North America

Widely available.

Europe

Seed suppliers[edit]

References[edit]

  1. ^ Stritch, L. (2019) [amended version of 2019 assessment]. "Ulmus alata". IUCN Red List of Threatened Species. 2019: e.T61966604A180056609. Retrieved 12 April 2021.
  2. ^ University of Florida, Environmental Horticulture Department (1994). Fact Sheet ST-648. Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences.
  3. ^ Alata is Latin for "winged".
  4. ^ "Trees: Ulmus alata". www.ces.ncsu.edu. Archived from the original on 3 August 2004.
  5. ^ Photo of U. alata samarae, jimbotany.com/16-Catalog_Ra_through_Z-Ackn-LitCitd.htm, [1]
  6. ^ Elwes, H. J. & Henry, A. (1913). The Trees of Great Britain & Ireland. Vol. VII. 1848–1929. Republished 2014 Cambridge University Press, ISBN 978-1-108-06938-0
  7. ^ Schnelle, M. (1999). Field Notes: Ulmus alata. American Nurseryman, page 1998, 1 March 1999. p. 98. Chicago
  8. ^ "Elm Phloem Necrosis".
  9. ^ Main, James (1839). The Forest Planter and Pruner's Assistant. London: Ridgway. p. 113.
  10. ^ Wilcox, Mike; Inglis, Chris (2003). "Auckland's elms" (PDF). Auckland Botanical Society Journal. 58 (1). Auckland Botanical Society: 38–45.
  11. ^ Event list nativetreesociety.org [dead link]
  12. ^ American Forests. (2012). The 2012 National Register of Big Trees.
  13. ^ Snow, G. A. "Ulmus alata Michx. Winged Elm". United States Department of Agriculture. Southern Research Station. Retrieved 2 February 2021.
  14. ^ "Bartlett Tree Experts: Tree Services". www.bartlett.com.
  15. ^ "BROOKLYN BOTANIC GARDEN >> Plant Records". www.bbg.org. Archived from the original on 30 April 2003.
  16. ^ Photographs of Morton Arboretum U. alata, Acc. no. 116-96; cirrusimage.com
  17. ^ "List of plants in the {elm} collection". Brighton & Hove City Council. Retrieved 23 September 2016.
  18. ^ Johnson, Owen (ed.) (2003). Champion Trees of Britain & Ireland. Whittet Press, ISBN 978-1-873580-61-5.
  19. ^ Royal Botanic Garden Edinburgh. (2017). List of Living Accessions: Ulmus [2]

External links[edit]