Benthophilinae
| Benthophilinae | ||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
||||||||||||
| Systematics | ||||||||||||
|
||||||||||||
| Scientific name | ||||||||||||
| Benthophilinae | ||||||||||||
| Beling & Ilyin , 1927 |
The Benthophilinae are a subfamily of the gobies . The subfamily was established in 1927 by the Russian ichthyologists D. Beling and BS Iljin, but the associated genera were later added to the subfamily Gobiinae . In 2009 the Benthophilinae were newly diagnosed by Neilson and Stepien as an endemic goby subfamily in the Pontocaspic region .
distribution
The small fresh and brackish water fish are found on the shores of the Black , Azov and Caspian Seas and in the incoming rivers. Five species from this subfamily spread over Central Europe in the last few decades and were able to colonize other catchment areas , e.g. B. the Rhine . Two species of the Benthophilinae, the marbled goby and the black-mouthed goby , got into the North American Great Lakes through the ballast water of ships and are now among the most common bottom fish there.
features
The Benthophilinae are small to medium-sized gobies. Their hallmarks include sensory papillae , which lie in six to seven rows below the eyes, tubular anterior nostrils, the posterior ones are near the eye sockets , and upper pectoral fin rays, which are surrounded by fin membrane. A swim bladder is missing. The eggs are medium to large in size and contain little yolk . A pelagic larval stage is absent. From the subfamily Gobiinae, to which the genera of the Benthophilinae were previously counted, the Benthophilinae can mainly be distinguished by the number of vertebrae. They have a total of 28 or more vertebrae, including 18 to 22 caudal vertebrae (all vertebrae behind the anus).
Internal system
-
Benthophilini
tribe
- Genus Anatirostrum
- Anatirostrum profundorum (Berg, 1927)
- Genus Benthophiloides
- Banded tadpole goby ( Benthophiloides brauneri Beling & Iljin, 1927 )
- Benthophiloides turcomanus (Iljin, 1941)
- Genus Benthophilus (type genus)
- Benthophilus abdurahmanovi Ragimov, 1978
- Benthophilus baeri Kessler, 1877
- Benthophilus casachicus Ragimov, 1978
- Benthophilus ctenolepidus Kessler, 1877
- Benthophilus durrelli Boldyrev & Bogutskaya, 2004
- Rough tadpole goby ( Benthophilus granulosus Kessler, 1877 )
- Benthophilus grimmi Kessler, 1877
- Benthophilus kessleri Berg, 1927
- Benthophilus leobergius Berg, 1949
- Benthophilus leptocephalus Kessler, 1877
- Benthophilus leptorhynchus Kessler, 1877
- Tadpole goby ( Benthophilus macrocephalus (Pallas, 1787) )
- Benthophilus magistri Iljin, 1927
- Benthophilus mahmudbejovi Ragimov, 1976
- Benthophilus nudus Berg, 1898
- Benthophilus pinchuki Ragimov, 1982
- Benthophilus ragimovi Boldyrev & Bogutskaya, 2004
- Benthophilus spinosus Kessler, 1877
- Star tadpole goby ( Benthophilus stellatus (Sauvage, 1874) )
- Benthophilus svetovidovi Pinchuk & Ragimov, 1979
- Genus Caspiosoma
- Caspi's goby ( Caspiosoma caspium (Kessler, 1877) )
- Genus Anatirostrum
- Tribe Neogobiini
- Genus Neogobius
- Neogobius caspius (Eichwald, 1831)
- River goby ( Neogobius fluviatilis (Pallas, 1814) )
- Black-mouthed goby ( Neogobius melanostomus (Pallas, 1814) )
- Neogobius pallasi (Berg, 1916)
- Genus Neogobius
- Tribe Ponticolini
- Genus Babka
- Babka gymnotrachelus (Kessler, 1857)
- Genus Mesogobius
- Toad goby ( Mesogobius batrachocephalus (Pallas, 1814) )
- Mesogobius nigronotatus (Kessler, 1877)
- Mesogobius nonultimus (Ilyin, 1936)
- Genus Ponticola (type genus)
- Ponticola bathybius (Kessler, 1877)
- Ponticola cephalargoides (Pinchuk, 1976)
- Ponticola constructor (Nordmann, 1840)
- Ponticola cyrius (Kessler, 1874)
- Ponticola eurycephalus (Kessler, 1874)
- Ponticola gorlap (Ilyin, 1949)
- Ponticola iranicus Vasiléva, Mousavi-Sabet & Vasilév, 2015
- Kessler goby ( Ponticola kessleri (Günther, 1861) )
- Ponticola platyrostris (Pallas, 1814)
- Ponticola ratan (Nordmann, 1840)
- Ponticola rhodioni (Vasiléva & Vasilév, 1994)
- Ponticola rizensis (Kovačić & Engin, 2008)
- Syrman goby ( Ponticola syrman (Nordmann, 1840) )
- Ponticola turani (Kovačić & Engin, 2008)
- Genus Proterorhinus
- Marbled goby ( Proterorhinus marmoratus (Pallas, 1814) )
- Proterorhinus nasalis (De Filippi, 1863)
- Proterorhinus semilunaris (Heckel, 1837)
- Proterorhinus semipellucidus (Kessler, 1863)
- Proterorhinus tataricus Freyhof & Naseka, 2007
- Genus Babka
literature
- Matthew E. Neilson, Carol A. Stepien: Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei). Molecular Phylogenetics and Evolution, Volume 52, Issue 1, July 2009, Pages 84-102 doi : 10.1016 / j.ympev.2008.12.023
