Sphaeroceridae: Difference between revisions

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{{Short description|Family of insects}}
:"Lesser dung fly" redirects here. This can also refer to the distantly related fly [[species]] ''[[Sepsis fulgens]].
{{Hatnote|"Lesser dung fly" redirects here. This can also refer to the distantly related ''[[Sepsis fulgens]]''.}}
{{Taxobox
{{Automatic taxobox
| image = Copromyza.equina.jpg
|image=Copromyza.equina.jpg
| image_width = 240px
| image_caption = ''[[Copromyza equina]]'' of the [[Copromyzinae]]
|image_caption=''[[Copromyza equina]]'' ([[Copromyzinae]])
|taxon=Sphaeroceridae
| regnum = [[Animal]]ia
|authority=[[Pierre-Justin-Marie Macquart|Macquart]], 1835<ref name="Macquart1835">{{cite book|last1=Macquart1835|first1=P. J. M.|title=Histoire Naturelle des insectes. Dipteres. Tome deuxieme.|publisher=Roret|location=Paris|pages=710, 12 pl}}</ref>
| phylum = [[Arthropod]]a
|subdivision_ranks=Subfamilies
| classis = [[Insect]]a
|subdivision=*[[Copromyzinae]]
| ordo = [[Fly|Diptera]]
| zoosectio = [[Schizophora]]
| zoosubsectio = [[Acalyptratae]]
| superfamilia = [[Sphaeroceroidea]]
| familia = '''Sphaeroceridae'''
| familia_authority = [[Justin Pierre Marie Macquart|Macquart]], 1835<ref name="Macquart1835">{{cite book|last1=Macquart1835|first1=P. J. M.|title=Histoire Naturelle des insectes. Dipteres. Tome deuxieme.|publisher=Roret|location=Paris|pages=710, 12 pl}}</ref>
| subdivision_ranks = [[Subfamilies]]
| subdivision = *[[Copromyzinae]]
*[[Homalomitrinae]]
*[[Homalomitrinae]]
*[[Limosininae]]
*[[Limosininae]]
*[[Sphaerocerinae]]
*[[Sphaerocerinae]]
*[[Tucminae]]
*[[Tucminae]]
| synonyms =
|synonyms={{collapsible list|bullets=on
*Borboridae <small>[[Edward Newman (entomologist)|Newman]], 1834 ([[International Commission on Zoological Nomenclature|suppressed]])</small>
|Borboridae <small>[[Edward Newman (entomologist)|Newman]], 1834 ([[International Commission on Zoological Nomenclature|suppressed]])</small>
*Borborites <small>[[Edward Newman (entomologist)|Newman]], 1834 ([[International Commission on Zoological Nomenclature|suppressed]])</small>
|Borborites <small>[[Edward Newman (entomologist)|Newman]], 1834 (suppressed)</small>
*Copromyzidae <small>[[Christian Stenhammar|Stenhammar]], 1855</small>
|Copromyzidae <small>[[Christian Stenhammar|Stenhammar]], 1855</small>
*Cypselidae <small>[[Friedrich Georg Hendel|Hendel]], 1910 ([[International Commission on Zoological Nomenclature|suppressed]])</small>
|Cypselidae <small>[[Friedrich Georg Hendel|Hendel]], 1910 ([suppressed)</small>
*Sphoeroceridae <small>[[Justin Pierre Marie Macquart|Macquart]], 1846 (''[[lapsus]]'')</small>
|Sphoeroceridae <small>[[Pierre-Justin-Marie Macquart|Macquart]], 1846 (''[[lapsus]]'')</small>
<ref name="Rohček2001">{{cite book|last1=Rohček|first1=Jindřich|last2=Marshall|first2=Stephen A.|last3=Norrbom|first3=Allen L.|last4=Buck|first4=Matthias|last5=Quiros|first5=Dora Isabel|last6=Smith|first6=Ian|editor1-last=Rohček|editor1-first=Jindřich|title=World Catalog of Sphaeroceridae (Diptera)|date=2001|publisher=Slezské Zemské Muzeum|location=Opava|isbn=978-8086224213|pages=1–414|url=http://www.uoguelph.ca/debu/catalog.htm|accessdate=3 September 2017}}</ref>
}}
}}
|synonyms_ref=<ref name="Rohček2001">{{cite book|last1=Rohček|first1=Jindřich|last2=Marshall|first2=Stephen A.|last3=Norrbom|first3=Allen L.|last4=Buck|first4=Matthias|last5=Quiros|first5=Dora Isabel|last6=Smith|first6=Ian|editor1-last=Rohček|editor1-first=Jindřich|title=World Catalog of Sphaeroceridae (Diptera)|date=2001|publisher=Slezské Zemské Muzeum|location=Opava|isbn=978-8086224213|pages=1–414|url=http://www.uoguelph.ca/debu/catalog.htm|access-date=3 September 2017}}</ref>
}}

'''Sphaeroceridae''' are a [[family (biology)|family]] of true flies in the [[order (biology)|order]] [[Fly|Diptera]], often called '''small dung flies''', '''lesser dung flies''' or '''lesser corpse flies''' due to their [[saprophagous]] habits. They belong to the typical fly [[suborder]] [[Brachycera]] as can be seen by their short [[antenna (biology)|antenna]]e, and more precisely they are members of the [[section (zoology)|section]] [[Schizophora]]. There are over 1,300 [[species]] and about 125 [[genera]] accepted as valid today, but new [[taxa]] are still being described.<ref name="Rohček2001"/>
'''Sphaeroceridae''' are a [[family (biology)|family]] of true flies in the [[order (biology)|order]] [[Fly|Diptera]], often called '''small dung flies''', '''lesser dung flies''' or '''lesser corpse flies''' due to their [[saprophagous]] habits. They belong to the typical fly [[suborder]] [[Brachycera]] as can be seen by their short [[antenna (biology)|antenna]]e, and more precisely they are members of the [[section (zoology)|section]] [[Schizophora]]. There are over 1,300 [[species]] and about 125 [[genera]] accepted as valid today, but new [[taxa]] are still being described.<ref name="Rohček2001"/>


Unlike the large "corpse flies" or [[blow-flies]] of the family Calliphoridae and the large dung flies of the family [[Scathophagidae]], the small dung flies are members of the schizophoran [[subsection (zoology)|subsection]] [[Acalyptratae]]. Among their [[superfamily (zoology)|superfamily]] [[Sphaeroceroidea]], they seem to be particularly close relatives of the family [[Heleomyzidae]].<ref name="Rohček2001"/>
Unlike the large "corpse flies" or [[Calliphoridae|blow-flies]] of the family Calliphoridae and the large dung flies of the family [[Scathophagidae]], the small dung flies are members of the schizophoran [[subsection (zoology)|subsection]] [[Acalyptratae]]. Among their [[superfamily (zoology)|superfamily]] [[Sphaeroceroidea]], they seem to be particularly close relatives of the family [[Heleomyzidae]].<ref name="Rohček2001"/>


[[File:WillistonSphaeroceridae.jpg|thumb|230px|left|Sphaeroceridae morphological details]]
[[File:WillistonSphaeroceridae.jpg|thumb|230px|Sphaeroceridae morphological details]]


==Description and ecology==
==Description and ecology==
[[File:Smallerdungfly.jpg|thumb|right|Small dung fly v.s HouseFly]]
[[File:Smallerdungfly.jpg|thumb|right|Housefly (left) vs. small dung fly (right)]]
For terms see [[Morphology of Diptera]]
{{Hatnote|For terms see [[Morphology of Diptera]]}}

Dung flies are small to minute, usually dull-colored flies with characteristically thickened first tarsomere of the posterior leg. The first tarsal segment is less than 1+1/2 times as long as the second tarsal segment and dilated. The crossvein separating the 2nd basal and discal cells is missing. Veins 4 and 5 often fade apically. They occur [[Cosmopolitan distribution|all over the world]] except in regions with permanent ice-cover. Despite their ubiquity and abundance, little is known about their economic or [[ecology|ecological]] impact. Some species are known to be [[parthenogenetic]].
Dung flies are small to minute, usually dull-colored flies with characteristically thickened first tarsomere of the posterior leg. The first tarsal segment is less than {{sfrac|1|1|2}} times as long as the second tarsal segment and dilated. The crossvein separating the second basal and discal cells is missing. Veins four and five often fade apically. They occur [[Cosmopolitan distribution|all over the world]] except in regions with permanent ice-cover. Despite their ubiquity and abundance, little is known about their economic or [[ecology|ecological]] impact. Some species are known to be [[parthenogenetic]].


Larval stages are poorly known, but those described are slender, narrowed anteriorly, with groups of ventral spicules on creeping welts. The larva is amphipneustic (having only the anterior and posterior pairs of spiracles). The mandibles are simple, hooked, and without additional teeth. The parastomal bars are long, thin structures, fused to the tentoropharyngeal sclerite. The hypopharyngeal sclerites are long separate or connected by a sclerotized bridge; the anterior spiracle (prothoracic spiracle) is a rosette or branched. The posterior spiracles (on the anal segment) are usually on two cylindrical lobes. Each spiracle has three slit or oval openings and three or five groups of interspiracular hairs that are branched in some species.{{citation needed|reason=very specific, need citation for whole paragraph|date=June 2016}}
Larval stages are poorly known, but those described are slender, narrowed anteriorly, with groups of ventral spicules on creeping welts. The larva is amphipneustic (having only the anterior and posterior pairs of spiracles). The mandibles are simple, hooked, and without additional teeth. The parastomal bars are long, thin structures, fused to the tentoropharyngeal sclerite. The hypopharyngeal sclerites are long separate or connected by a sclerotized bridge; the anterior spiracle (prothoracic spiracle) is a rosette or branched. The posterior spiracles (on the anal segment) are usually on two cylindrical lobes. Each spiracle has three slit or oval openings and three or five groups of interspiracular hairs that are branched in some species.{{citation needed|reason=very specific, need citation for whole paragraph|date=June 2016}}
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The larvae are [[microbe|microbial]] grazers found in abundance in many microenvironments with decomposing [[organic material]]. Most species appear to be associated with decaying [[plants]] or [[fungi]] and they are a part of the [[nutrient cycle]]. Some species, especially [[Troglobite|cave species]], are [[polysaprophagous]]. Many species are associated with various kinds of [[faeces]] including human faeces; there are a few [[carrion]]-feeding species. These, however, are extremely abundant and are important components of the carrion-insect community. Sphaerocerids that abound in economically important decomposer communities such as compost and manure, and some [[decay cycle]]s such as the [[wrack (science)|wrack]] (seaweed) cycle are mediated by sphaerocerid-dominated insect communities.
The larvae are [[microbe|microbial]] grazers found in abundance in many microenvironments with decomposing [[organic material]]. Most species appear to be associated with decaying [[plants]] or [[fungi]] and they are a part of the [[nutrient cycle]]. Some species, especially [[Troglobite|cave species]], are [[polysaprophagous]]. Many species are associated with various kinds of [[faeces]] including human faeces; there are a few [[carrion]]-feeding species. These, however, are extremely abundant and are important components of the carrion-insect community. Sphaerocerids that abound in economically important decomposer communities such as compost and manure, and some [[decay cycle]]s such as the [[wrack (science)|wrack]] (seaweed) cycle are mediated by sphaerocerid-dominated insect communities.


As their [[microbe]]-associated habits suggest, sphaerocerids may carry many [[pathogenic]] microorganisms.<ref>Greenberg, B., 1971. ''Flies and Disease'', volume I: Ecology, Classification,and Biotic Association. Princeton University Press, Princeton, New Jersey.</ref> Although their reclusive habits preclude a major role in disease transmission; some can present a [[public health]] hazard on occasion or act as a warning of one. For instance ''Leptocera caenosa'' and other sphaerocerids are associated with blocked sewage drains.<ref>{{cite journal | last1 = Fredeen | first1 = F. J. H. | last2 = Taylor | first2 = M. E. | year = 1964 | title = Borborids (Diptera: Sphaeroceridae) infesting sewage disposal tanks, with notes on the life cycle, behavior and control of Leptocera(Leptocera) caenosa ([[Camillo Róndani|Róndani]]) | url = | journal = Can. Ent. | volume = 96 | issue = | pages = 801–808 | doi=10.4039/ent96801-5}}</ref> Some species occasionally reach high population levels in food-processing plants and other buildings where they may indicate blocked drains, waste accumulation and inadequate hygiene. One species, ''[[Poecilosomella angulata]]'', has been implicated in human [[intestinal]] [[myiasis]]<ref>{{cite journal | last1 = McKibben | first1 = J.W. | last2 = Micks | first2 = D.W. | year = 1956 | title = Report of a case of human intestinal myiasis caused by Leptocera venalicia | url = | journal = American Journal of Tropical Medicine and Hygiene | volume = 5 | issue = 5| pages = 929–32 }}</ref> They have been implicated as the major means by which [[nematode]]s are disseminated among mushroom houses. Sphaeoceridae often coexist with muscoids especially ''[[Fannia canicularis]]'' and ''[[Musca domestica]]'' in the complex manure ecosystem of [[Poultry farming|poultry houses]], and other confined-animal facilities. Here the sphaeocerids are prey for mites and beetles, which themselves also feed on the immatures of muscoid flies reducing the population of the more problematic muscoids.<ref>Axtell, R. C. 1985. Poultry Pests. Chapter 16, p. 269-293, In: ''Livestock Entomology'' (Williams et al., editors), Wiley & Sons, New York.</ref> Carrion-feeding species are useful [[post mortem interval]] indicators in [[forensic entomology]].
As their [[microbe]]-associated habits suggest, sphaerocerids may carry many [[pathogenic]] microorganisms.<ref>Greenberg, B., 1971. ''Flies and Disease'', volume I: ''Ecology, Classification, and Biotic Association''. Princeton University Press, Princeton, New Jersey.</ref> Although their reclusive habits preclude a major role in disease transmission; some can present a [[public health]] hazard on occasion or act as a warning of one. For instance ''Leptocera caenosa'' and other sphaerocerids are associated with blocked sewage drains.<ref>{{cite journal |last1=Fredeen |first1=F. J. H. |last2=Taylor |first2=M. E. |year=1964 |title=Borborids (Diptera: Sphaeroceridae) infesting sewage disposal tanks, with notes on the life cycle, behavior and control of Leptocera(Leptocera) caenosa (Rondani) |journal=Can. Entomol. |volume=96 |issue=5|pages=801–808 |doi=10.4039/ent96801-5}}</ref> Some species occasionally reach high population levels in food-processing plants and other buildings where they may indicate blocked drains, waste accumulation and inadequate hygiene. One species, ''[[Poecilosomella angulata]]'', has been implicated in human [[intestinal]] [[myiasis]]<ref>{{cite journal |last1=McKibben |first1=J.W. |last2=Micks |first2=D.W. |year=1956 |title=Report of a case of human intestinal myiasis caused by Leptocera venalicia |journal=American Journal of Tropical Medicine and Hygiene |volume=5 |issue=5|pages=929–32 |doi=10.4269/ajtmh.1956.5.929 |pmid=13362764 }}</ref> They have been implicated as the major means by which [[nematode]]s are disseminated among mushroom houses. Sphaeoceridae often coexist with muscoids especially ''[[Fannia canicularis]]'' and ''[[Musca domestica]]'' in the complex manure ecosystem of [[Poultry farming|poultry houses]], and other confined-animal facilities. Here the sphaeocerids are prey for mites and beetles, which themselves also feed on the immatures of muscoid flies reducing the population of the more problematic muscoids.<ref>Axtell, R. C. (1985). Chapter 16: "Poultry Pests". In: ''Livestock Entomology'' (Williams et al., editors), Wiley & Sons, New York. pp. 269–293.</ref> Carrion-feeding species are useful [[post mortem interval]] indicators in [[forensic entomology]].


==Genera==
==Genera==
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'''Subfamily [[Tucminae]] <small>Marshall, 1996</small>'''<ref name="Marshall1996">{{cite journal|last1=Marshall|first1=S. A.|title=''Tucma fritzi'', a new species in the enigmatic genus ''Tucma'' Mourguès-Schurter (Diptera, Sphaeroceridae, Tucminae new subfamily)|journal=Studia Dipterologica|date=1996|issue=3|pages=283–288}}</ref>
'''Subfamily [[Tucminae]] <small>Marshall, 1996</small>'''<ref name="Marshall1996">{{cite journal|last1=Marshall|first1=S. A.|title=''Tucma fritzi'', a new species in the enigmatic genus ''Tucma'' Mourguès-Schurter (Diptera, Sphaeroceridae, Tucminae new subfamily)|journal=Studia Dipterologica|date=1996|issue=3|pages=283–288}}</ref>
{{columns-list|colwidth=25em|1=
{{Columns-list|colwidth=25em|1=
* ''[[Tucma (fly)|Tucma]]'' <small>Mourgués-Schurter, 1987</small><ref name="Mourguès-Schurter1987b">{{cite journal|last1=Mourguès-Schurter|first1=L.|title=''Tucma'', novo gènero de Copromyzinae para a AmÈrica do Sul (Diptera, Sphaeroceridae).|journal=Revista Brasileira de Entomologia, S„o Paulo|date=1987|volume=31|issue=1|pages=118–122}}</ref>
* ''[[Tucma (fly)|Tucma]]'' <small>Mourgués-Schurter, 1987</small><ref name="Mourguès-Schurter1987b">{{cite journal|last1=Mourguès-Schurter|first1=L.|title=''Tucma'', novo gènero de Copromyzinae para a AmÈrica do Sul (Diptera, Sphaeroceridae).|journal=Revista Brasileira de Entomologia, S"o Paulo|date=1987|volume=31|issue=1|pages=118–122}}</ref>
}}
}}


'''Subfamily [[Copromyzinae]] <small>[[Christian Stenhammar|Stenhammar]], 1855</small>'''<ref name="Stenhammar1855">{{cite journal|last1=Stenhammar|first1=C.|title=Skandinaviens Copromyzine granskade och beskrifne|journal=Kongliga Vetenskaps-Akademiens Handlingar |series=Ser. 3 |date=1855|volume=1853|pages=257–442|location=Stockholm }}</ref>
'''Subfamily [[Copromyzinae]] <small>[[Christian Stenhammar|Stenhammar]], 1855</small>'''<ref name="Stenhammar1855">{{cite journal|last1=Stenhammar|first1=C.|title=Skandinaviens Copromyzine granskade och beskrifne|journal=Kongliga Vetenskaps-Akademiens Handlingar |series=Ser. 3 |date=1855|volume=1853|pages=257–442|location=Stockholm }}</ref>
{{columns-list|colwidth=25em|1=
{{Columns-list|colwidth=25em|1=
* ''[[Achaetothorax]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923">{{cite journal|last1=Hedicke|first1=H.|title=Nomina nova. IV.|journal=Deutsche entomologische Zeitschrift|date=1923|volume=1923|pages=431|doi=10.1002/mmnd.48019230413}}</ref>
* ''[[Achaetothorax]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923">{{cite journal|last1=Hedicke|first1=H.|title=Nomina nova. IV.|journal=Deutsche entomologische Zeitschrift|date=1923|volume=1923|issue=4|pages=431|doi=10.1002/mmnd.48019230413}}</ref>
* ''[[Alloborborus]]'' <small>[[Oswald Duda|Duda]], 1923</small><ref name="Duda1923">{{cite journal|last1=Duda|first1=Oswald|title=Revision der altweltlichen Arten der Gattung ‘’Borborus’’ (Cypsela) Meigen (Dipteren)|journal=Archiv für Naturgeschichte |series=Abteilung A|date=1923|volume=89|issue=4|pages=35–112|url=https://www.biodiversitylibrary.org/item/113951#page/41/mode/1up|accessdate=3 September 2017}}</ref>
* ''[[Alloborborus]]'' <small>[[Oswald Duda|Duda]], 1923</small><ref name="Duda1923">{{cite journal|last1=Duda|first1=Oswald|title=Revision der altweltlichen Arten der Gattung ''Borborus'' (Cypsela) Meigen (Dipteren)|journal=Archiv für Naturgeschichte |series=Abteilung A|date=1923|volume=89|issue=4|pages=35–112|url=https://www.biodiversitylibrary.org/item/113951#page/41/mode/1up|access-date=3 September 2017}}</ref>
* ''[[Antrops]]'' <small>[[Günther Enderlein|Enderlein]], 1909</small><ref name="Enderlein1909">{{cite journal|last1=Enderlein|first1=Günther|title=''Antrops truncipennis'', eine neue Borboridengattung von Feuerland|journal=Zoologischer Anzeiger|date=1909|volume=34|pages=225–230|url=https://www.biodiversitylibrary.org/item/95292#page/241/mode/1up|accessdate=4 September 2017}}</ref>
* ''[[Antrops]]'' <small>[[Günther Enderlein|Enderlein]], 1909</small><ref name="Enderlein1909">{{cite journal|last1=Enderlein|first1=Günther|title=''Antrops truncipennis'', eine neue Borboridengattung von Feuerland|journal=Zoologischer Anzeiger|date=1909|volume=34|pages=225–230|url=https://www.biodiversitylibrary.org/item/95292#page/241/mode/1up|access-date=4 September 2017}}</ref>
* ''[[Archiborborus]]'' <small>[[Oswald Duda|Duda]], 1921</small><ref name="Duda1921">{{cite journal|last1=Duda|first1=Oswald|title=Fiebrigella und Archiborborus, zwei neue südamerikanische Borboriden-gattungen (Dipteren)|journal=Tijdschrift voor Entomologie|date=1921|volume=64|pages=119–146|url=https://www.biodiversitylibrary.org/page/10858427#page/231/mode/1up|accessdate=4 September 2017}}</ref>
* ''[[Archiborborus]]'' <small>[[Oswald Duda|Duda]], 1921</small><ref name="Duda1921">{{cite journal|last1=Duda|first1=Oswald|title=Fiebrigella und Archiborborus, zwei neue südamerikanische Borboriden-gattungen (Dipteren)|journal=Tijdschrift voor Entomologie|date=1921|volume=64|pages=119–146|url=https://www.biodiversitylibrary.org/page/10858427#page/231/mode/1up|access-date=4 September 2017}}</ref>
* ''[[Borborillus]]'' <small>[[Oswald Duda|Duda]], 1923</small><ref name="Duda1923"/>
* ''[[Borborillus]]'' <small>[[Oswald Duda|Duda]], 1923</small><ref name="Duda1923"/>
* ''[[Copromyza]]'' <small>[[Carl Fredrik Fallén|Fallén]], 1810</small><!-- FaunAbh26:3. -->
* ''[[Copromyza]]'' <small>[[Carl Fredrik Fallén|Fallén]], 1810</small><!-- FaunAbh26:3. -->
* ''[[Crumomyia]]'' <small>[[Justin Pierre Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
* ''[[Crumomyia]]'' <small>[[Pierre-Justin-Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
* ''[[Dudaia]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923"/>
* ''[[Dudaia]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923"/>
* ''[[Frutillaria]]'' <small>Richards, 1961</small><ref name="Richards1961a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera (Sphaeroceridae) from south Chile|journal=Proceedings of the Royal Entomological Society of London B|date=1961|volume=30|pages=57–68}}</ref>
* ''[[Frutillaria]]'' <small>Richards, 1961</small><ref name="Richards1961a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera (Sphaeroceridae) from south Chile|journal=Proceedings of the Royal Entomological Society of London B|date=1961|volume=30|pages=57–68}}</ref>
* ''[[Gymnometopina]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923"/>
* ''[[Gymnometopina]]'' <small>Hedicke, 1923</small><ref name="Hedicke1923"/>
* ''[[Lotophila]]'' <small>[[Paolo Lioy|Lioy]], 1864</small><ref name="Lioy1864">{{cite book|last1=Lioy|first1=P.|title=I Ditteri distribuiti secondo un nuovo metodo di classifi cazione naturale Serie III, 9|date=1864|publisher=Atti dellíIstituto Veneto di Scienze, Lettere ed Arti|location=Venezia|pages=1087–1126}}</ref><!-- FaunAbh26:3. -->
* ''[[Lotophila]]'' <small>[[Paolo Lioy|Lioy]], 1864</small><ref name="Lioy1864">{{cite book|last1=Lioy|first1=P.|title=I Ditteri distribuiti secondo un nuovo metodo di classifi cazione naturale Serie III, 9|date=1864|publisher=Atti dellíIstituto Veneto di Scienze, Lettere ed Arti|location=Venezia|pages=1087–1126}}</ref><!-- FaunAbh26:3. -->
* ''[[Immoderatus]]'' <small>Papp, 2004</small><ref name="Papp2004">{{cite journal|last1=Papp|first1=L.|title=''Immoderatus'' gen. n. of Copromyzinae (Diptera: Sphaeroceridae)|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=2004|volume=50|issue=1|pages=45–53|url=http://actazool.nhmus.hu/50/1/papp.pdf|accessdate=2 September 2017}}</ref>
* ''[[Immoderatus]]'' <small>Papp, 2004</small><ref name="Papp2004">{{cite journal|last1=Papp|first1=L.|title=''Immoderatus'' gen. n. of Copromyzinae (Diptera: Sphaeroceridae)|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=2004|volume=50|issue=1|pages=45–53|url=http://actazool.nhmus.hu/50/1/papp.pdf|access-date=2 September 2017}}</ref>
* ''[[Metaborborus]]'' <small>Vanschuytbroeck, 1948</small><ref name="Vanschuytbroeck1948">{{cite journal|last1=Vanschuytbroeck|first1=P.|title=Sphaeroceridae (Diptera, Acalyptratae)|journal=Exploration du Parc National Albert, Mission G. F. de Witte (1933-35)|date=1948|volume=52|pages=1–43}}</ref>
* ''[[Metaborborus]]'' <small>Vanschuytbroeck, 1948</small><ref name="Vanschuytbroeck1948">{{cite journal|last1=Vanschuytbroeck|first1=P.|title=Sphaeroceridae (Diptera, Acalyptratae)|journal=Exploration du Parc National Albert, Mission G. F. De Witte (1933-35)|date=1948|volume=52|pages=1–43}}</ref>
* ''[[Norrbomia]]'' <small>Papp, 1988</small><ref name="Papp1988a">{{cite journal|last1=Papp|first1=L.|title=A review of the Afrotropical species of Norrbomia gen.n. (Diptera: Sphaeroceridae, Copromyzini)|journal=Acta Zoologica Hungarica|date=1988|volume=34|pages=393–408}}</ref><ref name="MarshallNorrbom1992">{{cite journal|last1=Marshall|first1=Stephen A.|last2=Norrbom|first2=Allen L.|title=A revision of the New World species of ''Norrbomia'' (Diptera: Sphaeroceridae), including all American species previously placed in ''Borborillus''|journal=Insecta Mundi|date=1992|volume=6|issue=3–4|pages=151–181|url=http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1441&context=insectamundi|accessdate=3 September 2017|publisher=Center for Systematic Entomology, Gainesville, Florida}}</ref>
* ''[[Norrbomia]]'' <small>Papp, 1988</small><ref name="Papp1988a">{{cite journal|last1=Papp|first1=L.|title=A review of the Afrotropical species of Norrbomia gen.n. (Diptera: Sphaeroceridae, Copromyzini)|journal=Acta Zoologica Hungarica|date=1988|volume=34|pages=393–408}}</ref><ref name="MarshallNorrbom1992">{{cite journal|last1=Marshall|first1=Stephen A.|last2=Norrbom|first2=Allen L.|title=A revision of the New World species of ''Norrbomia'' (Diptera: Sphaeroceridae), including all American species previously placed in ''Borborillus''|journal=Insecta Mundi|date=1992|volume=6|issue=3–4|pages=151–181|url=http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1441&context=insectamundi|access-date=3 September 2017|publisher=Center for Systematic Entomology, Gainesville, Florida}}</ref>
* ''[[Palaeoceroptera]]'' <small>[[Oswald Duda|Duda]], 1929</small><ref name="Duda1929">{{cite journal|last1=Duda|first1=Oswald|title=Die Ausbeute der deutschen Chako-Expedition 1925/1926 (Diptera). VI. Sepsidae, VII. Piophilidae, VIII. Cypselidae, IX. Drosophilidae und X. Chloropidae|journal=Konowia|date=1929|volume=8|issue=1|pages=33–50}}</ref>
* ''[[Palaeoceroptera]]'' <small>[[Oswald Duda|Duda]], 1929</small><ref name="Duda1929">{{cite journal|last1=Duda|first1=Oswald|title=Die Ausbeute der deutschen Chako-Expedition 1925/1926 (Diptera). VI. Sepsidae, VII. Piophilidae, VIII. Cypselidae, IX. Drosophilidae und X. Chloropidae|journal=Konowia|date=1929|volume=8|issue=1|pages=33–50}}</ref>
* ''[[Palaeolimosina]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b">{{cite journal|last1=Duda|first1=Oswald|title=Vorläufige Mitteilung zur Kenntnis der außereuropäischen Arten der Gattungen Leptoceras Olivier = Limosina Macq. und Borborus Meigen (Dipteren)|journal=Zoologische Jahrbücher. Zeitschrift für Systematik, Geographie und Biologie der Tier|date=1920|volume=43|pages=433–446|url=https://www.biodiversitylibrary.org/page/9476663#page/443/mode/1up|accessdate=17 September 2017}}</ref>
* ''[[Palaeolimosina]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b">{{cite journal|last1=Duda|first1=Oswald|title=Vorläufige Mitteilung zur Kenntnis der außereuropäischen Arten der Gattungen Leptoceras Olivier=Limosina Macq. und Borborus Meigen (Dipteren)|journal=Zoologische Jahrbücher. Zeitschrift für Systematik, Geographie und Biologie der Tier|date=1920|volume=43|pages=433–446|url=https://www.biodiversitylibrary.org/page/9476663#page/443/mode/1up|access-date=17 September 2017}}</ref>
* ''[[Penola (fly)|Penola]]'' <small>Richards, 1941</small><ref name="Richards1941">{{cite journal|last1=Richards|first1=O. W.|title=Sphaeroceridae (Diptera) collected by the British Graham Land Expedition, 1934-1937|journal=British Graham Land Expedition 1934-37, Scientific Reports|date=1941|volume=1|issue=7|pages=323–326}}</ref>
* ''[[Penola (fly)|Penola]]'' <small>Richards, 1941</small><ref name="Richards1941">{{cite journal|last1=Richards|first1=O. W.|title=Sphaeroceridae (Diptera) collected by the British Graham Land Expedition, 1934-1937|journal=British Graham Land Expedition 1934-37, Scientific Reports|date=1941|volume=1|issue=7|pages=323–326}}</ref>
* ''[[Pycnopota]]'' <small>[[Mario Bezzi|Bezzi]], 1927</small><ref name="Bezzi1927">{{cite journal|last1=Bezzi|first1=Mario|title=A new genus and species of Borborid flies from South America.|journal=Proceedings of the United States National Museum|date=1927|volume=68 | issue = 1926 |issue=20|pages=1–6.|url=https://www.biodiversitylibrary.org/page/7610465#page/719/mode/1up|accessdate=17 September 2017}}</ref>
* ''[[Pycnopota]]'' <small>[[Mario Bezzi|Bezzi]], 1927</small><ref name="Bezzi1927">{{cite journal|last1=Bezzi|first1=Mario|title=A new genus and species of Borborid flies from South America.|journal=Proceedings of the United States National Museum|date=1927|volume=68 |number=2621 |pages=1–6|url=https://www.biodiversitylibrary.org/page/7610465#page/719/mode/1up|access-date=17 September 2017|doi=10.5479/si.00963801.68-2621.1}}</ref>
* ''[[Richardsia]]'' <small>Papp, 1973</small><ref name="Papp1973a">{{cite journal|last1=Papp|first1=László|title=Sphaeroceridae (Diptera) from Mongolia|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1973|volume=19|pages=369–425}}</ref>
* ''[[Richardsia]]'' <small>Papp, 1973</small><ref name="Papp1973a">{{cite journal|last1=Papp|first1=László|title=Sphaeroceridae (Diptera) from Mongolia|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1973|volume=19|pages=369–425}}</ref>
}}
}}


'''Subfamily [[Sphaerocerinae]] <small>[[Justin Pierre Marie Macquart|Macquart]], 1835</small>'''<ref name="Macquart1835"/>
'''Subfamily [[Sphaerocerinae]] <small>[[Pierre-Justin-Marie Macquart|Macquart]], 1835</small>'''<ref name="Macquart1835"/>
[[File:Sphaerocera cf curvipes - Marburg 2012.ogv|thumb|thumbtime=0:53|''Sphaerocera'' cf. ''curvipes'' of the [[Sphaerocerinae]]]]
[[File:Sphaerocera cf curvipes - Marburg 2012.ogv|thumb|thumbtime=0:53|''Sphaerocera'' cf. ''curvipes'' of the [[Sphaerocerinae]]]]
{{columns-list|colwidth=25em|1=
{{Columns-list|colwidth=25em|1=
* ''[[Afromyia]]'' <small>Kim, 1968</small><ref name="Kim1968">{{cite journal|last1=Kim|first1=K. C.|title=Revision of Sphaerocera, with description of a new genus ''Afromyia'' (Diptera: Sphaeroceridae).|journal=Annals of the Entomological Society of America|date=1968|volume=61|issue=2|pages=296–312|doi=10.1093/aesa/61.2.296}}</ref>
* ''[[Afromyia]]'' <small>Kim, 1968</small><ref name="Kim1968">{{cite journal|last1=Kim|first1=K. C.|title=Revision of Sphaerocera, with description of a new genus ''Afromyia'' (Diptera: Sphaeroceridae).|journal=Annals of the Entomological Society of America|date=1968|volume=61|issue=2|pages=296–312|doi=10.1093/aesa/61.2.296}}</ref>
* ''[[Ischiolepta]]'' <small>[[Paolo Lioy|Lioy]], 1864</small><ref name="Lioy1864"/>
* ''[[Ischiolepta]]'' <small>[[Paolo Lioy|Lioy]], 1864</small><ref name="Lioy1864"/>
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* ''[[Mesosphaerocera]]'' <small>Kim, 1972</small><ref name="Kim1972b">{{cite journal|last1=Kim|first1=K. C.|title=The New World genus ''Parasphaerocera'' and allied groups, with descriptions of new genera and species (Diptera: Sphaeroceridae)|journal=Miscellaneous Publications of the Entomological Society of America|volume=8|issue=6|pages=377–444}}</ref>
* ''[[Mesosphaerocera]]'' <small>Kim, 1972</small><ref name="Kim1972b">{{cite journal|last1=Kim|first1=K. C.|title=The New World genus ''Parasphaerocera'' and allied groups, with descriptions of new genera and species (Diptera: Sphaeroceridae)|journal=Miscellaneous Publications of the Entomological Society of America|volume=8|issue=6|pages=377–444}}</ref>
* ''[[Neosphaerocera]]'' <small>Kim, 1972</small><ref name="Kim1972b"/>
* ''[[Neosphaerocera]]'' <small>Kim, 1972</small><ref name="Kim1972b"/>
* ''[[Parasphaerocera]]'' <small>Spuler, 1924</small><ref name="Spuler1924c">{{cite journal|last1=Spuler|first1=Anthony|title=North American species of the genus Sphaerocera and Aptilotus (Diptera, Borboridae)|journal=The Pan-Pacific Entomologist|date=1924|volume=1|pages=66–71|url=https://www.biodiversitylibrary.org/page/53385719#page/72/mode/1up|accessdate=17 September 2017|publisher=The Pacific Coast Entomological Society|location=San Francisco}}</ref>
* ''[[Parasphaerocera]]'' <small>Spuler, 1924</small><ref name="Spuler1924c">{{cite journal|last1=Spuler|first1=Anthony|title=North American species of the genus Sphaerocera and Aptilotus (Diptera, Borboridae)|journal=The Pan-Pacific Entomologist|date=1924|volume=1|pages=66–71|url=https://www.biodiversitylibrary.org/page/53385719#page/72/mode/1up|access-date=17 September 2017|publisher=The Pacific Coast Entomological Society|location=San Francisco}}</ref>
* ''[[Safaria]]'' <small>Richards, 1950</small><ref name="Richards1950">{{cite journal|last1=Richards|first1=O. W.|title=New species of Sphaeroceridae (Diptera) found with driver ants (Dorylinae) by S. Patrizi and F. Meneghetti|journal=Bollettino dellí Istituto di Entomologia della Università degli Studi di Bologna|date=1950|volume=18|pages=14–23}}</ref>
* ''[[Safaria]]'' <small>Richards, 1950</small><ref name="Richards1950">{{cite journal|last1=Richards|first1=O. W.|title=New species of Sphaeroceridae (Diptera) found with driver ants (Dorylinae) by S. Patrizi and F. Meneghetti|journal=Bollettino dellí Istituto di Entomologia della Università degli Studi di Bologna|date=1950|volume=18|pages=14–23}}</ref>
* ''[[Sphaerocera]]'' <small>[[Pierre André Latreille|Latreille]], 1804</small><ref name="Latreille1805">{{cite book|last1=Latreille|first1=Pierre André|title=Histoire naturelle, générale et particulière des crustacés et des insectes |date=1805|publisher=Dufart|volume=14|location=Paris|pages=1–432|url=https://www.biodiversitylibrary.org/item/53501#page/453/mode/1up|accessdate=17 September 2017}}</ref>
* ''[[Sphaerocera]]'' <small>[[Pierre André Latreille|Latreille]], 1804</small><ref name="Latreille1805">{{cite book|last1=Latreille|first1=Pierre André|title=Histoire naturelle, générale et particulière des crustacés et des insectes |date=1805|publisher=Dufart|volume=14|location=Paris|pages=1–432|url=https://www.biodiversitylibrary.org/item/53501#page/453/mode/1up|access-date=17 September 2017}}</ref>
* ''[[Trichosphaerocera]]'' <small>Papp, 1978</small><ref name="Papp1978d">{{cite journal|last1=Papp|first1=László|title=Sphaeroceridae (Diptera) in the collection of the Hungarian Natural History Museum. IV. Sphaerocerinae|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1978|volume=24|pages=371–395}}</ref>
* ''[[Trichosphaerocera]]'' <small>Papp, 1978</small><ref name="Papp1978d">{{cite journal|last1=Papp|first1=László|title=Sphaeroceridae (Diptera) in the collection of the Hungarian Natural History Museum. IV. Sphaerocerinae|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1978|volume=24|pages=371–395}}</ref>
}}
}}


'''Subfamily [[Homalomitrinae]] <small>Roháček & Marshall, 1998</small>'''<ref name="RoháčekMarshall1998a">{{cite journal|last1=Roháček|first1=J.|last2=Marshall|first2=S. A.|title=Revision of Homalomitrinae subfam.n. (Diptera:Sphaeroceridae), with the description of a new genus and three new species|journal=European Journal of Entomology|date=1998|volume=95|issue=3|pages=455–491|url=https://www.eje.cz/pdfs/eje/1998/03/20.pdf|accessdate=12 October 2017}}</ref>
'''Subfamily [[Homalomitrinae]] <small>Roháček & Marshall, 1998</small>'''<ref name="RoháčekMarshall1998a">{{cite journal|last1=Roháček|first1=J.|last2=Marshall|first2=S. A.|title=Revision of Homalomitrinae subfam.n. (Diptera:Sphaeroceridae), with the description of a new genus and three new species|journal=European Journal of Entomology|date=1998|volume=95|issue=3|pages=455–491|url=https://www.eje.cz/pdfs/eje/1998/03/20.pdf|access-date=12 October 2017}}</ref>
{{columns-list|colwidth=25em|1=
{{Columns-list|colwidth=25em|1=
* ''[[Homalomitra]]'' <small>Borgmeier, 1931</small><ref name="Borgmeier">{{cite journal|last1=Borgmeier|first1=T.|title=Um cypselideviso ecitophilo de Goyaz (Dipt., Cypselidae)|journal=Reta de Entomologia, São Paulo|date=1931|volume=1|issue=1|pages=30–37}}</ref>
* ''[[Homalomitra]]'' <small>Borgmeier, 1931</small><ref name="Borgmeier">{{cite journal|last1=Borgmeier|first1=T.|title=Um cypselideviso ecitophilo de Goyaz (Dipt., Cypselidae)|journal=Reta de Entomologia, São Paulo|date=1931|volume=1|issue=1|pages=30–37}}</ref>
* ''[[Sphaeromitra]]'' <small>Roháček & Marshall, 1998</small><ref name="RoháčekMarshall1998a"/>
* ''[[Sphaeromitra]]'' <small>Roháček & Marshall, 1998</small><ref name="RoháčekMarshall1998a"/>
Line 96: Line 92:


[[File:Limosina.silvatica.jpg|thumb|right|''[[Limosina silvatica]]'' of the [[Limosininae]]]]
[[File:Limosina.silvatica.jpg|thumb|right|''[[Limosina silvatica]]'' of the [[Limosininae]]]]
'''Subfamily [[Limosininae]] <small>[[Heinrich Frey|Frey]], 1921</small>'''<ref name="Frey1921">{{cite journal|last1=Frey|first1=R.|title=Studien über der Bau des Mundes der niederen Diptera Schizophora nebst Bemerkungen über die Systematik dieser Dipteren-Gruppe |journal=Acta Societatis pro Fauna et Flora Fennica |location=Helsinki |date=1921|volume=48|issue=3|pages=1–247|url=https://www.biodiversitylibrary.org/item/28237#page/239/mode/1up|accessdate=21 October 2017}}</ref>
'''Subfamily [[Limosininae]] <small>[[Richard Karl Hjalmar Frey|Frey]], 1921</small>'''<ref name="Frey1921">{{cite journal|last1=Frey|first1=R.|title=Studien über der Bau des Mundes der niederen Diptera Schizophora nebst Bemerkungen über die Systematik dieser Dipteren-Gruppe |journal=Acta Societatis Pro Fauna et Flora Fennica |location=Helsinki |date=1921|volume=48|issue=3|pages=1–247|url=https://www.biodiversitylibrary.org/item/28237#page/239/mode/1up|access-date=21 October 2017}}</ref>
{{columns-list|colwidth=25em|1=
{{Columns-list|colwidth=25em|1=
*''[[Acuminiseta]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925">{{cite journal|last1=Duda|first1=Oswald|title=Die außereuropäischen Arten der Gattung Leptocera Olivier - Limosina Macquart (Dipteren) mit Berücksichtigung der europäischen Arten|journal=Archiv für Naturgeschichte |location=Berlin |series=Abteilung A|date=1925|volume=90|issue=(11)(1924|pages=5–21|url=http://www.zobodat.at/pdf/Archiv-Naturgeschichte_90A_11_0001-0215.pdf|accessdate=21 October 2017}}</ref>
*''[[Acuminiseta]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925">{{cite journal|last1=Duda|first1=Oswald|title=Die außereuropäischen Arten der Gattung Leptocera Olivier - Limosina Macquart (Dipteren) mit Berücksichtigung der europäischen Arten|journal=Archiv für Naturgeschichte |location=Berlin |series=Abteilung A|date=1925|volume=90|issue=(11)(1924|pages=5–21|url=http://www.zobodat.at/pdf/Archiv-Naturgeschichte_90A_11_0001-0215.pdf|access-date=21 October 2017}}</ref>
*''[[Afropterogramma]]'' <small>Papp, 2008</small><ref name="Papp2008b">{{cite journal|last1=Papp|first1=László|title=New genera of the Old World Limosininae (Diptera, Sphaeroceridae)|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=2008|volume=54(supplement 2)|pages=47–209|url=http://actazool.nhmus.hu/54/Suppl2/azh54_S2_Limos.pdf|accessdate=24 October 2017}}</ref>
*''[[Afropterogramma]]'' <small>Papp, 2008</small><ref name="Papp2008b">{{cite journal|last1=Papp|first1=László|title=New genera of the Old World Limosininae (Diptera, Sphaeroceridae)|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=2008|volume=54(supplement 2)|pages=47–209|url=http://actazool.nhmus.hu/54/Suppl2/azh54_S2_Limos.pdf|access-date=24 October 2017}}</ref>
*''[[Aluligera]]'' <small>Richards, 1951</small><ref name="Richards1951a">{{cite journal|last1=Richards|first1=O. W.|title=Brachypterous Sphaeroceridae|journal=British Museum (Natural History) Ruwenzori Expedition, 1934-1935|date=1951|volume=2|issue=8|pages=829–851}}</ref>
*''[[Aluligera]]'' <small>Richards, 1951</small><ref name="Richards1951a">{{cite journal|last1=Richards|first1=O. W.|title=Brachypterous Sphaeroceridae|journal=British Museum (Natural History) Ruwenzori Expedition, 1934-1935|date=1951|volume=2|issue=8|pages=829–851}}</ref>
*''[[Anatalanta]]'' <small>[[Alfred Edwin Eaton|Eaton]], 1875</small><ref name="Eaton1875b">{{cite journal|last1=Eaton|first1=A. E.|title=Breves dipterarum uniusque lepidopterarum insulae Kerguelensi indigenarum diagnoses|journal=The Entomologists Monthly Magazine|date=1875|volume=12|pages= 58–61}}</ref>
*''[[Anatalanta]]'' <small>[[Alfred Edwin Eaton|Eaton]], 1875</small><ref name="Eaton1875b">{{cite journal|last1=Eaton|first1=A. E.|title=Breves dipterarum uniusque lepidopterarum insulae Kerguelensi indigenarum diagnoses|journal=The Entomologist's Monthly Magazine|date=1875|volume=12|pages= 58–61}}</ref>
*''[[Anomioptera]]'' <small>[[Ignaz Rudolph Schiner|Schiner]], 1868</small><ref name="Schiner1868">{{cite book|last1=Schiner|first1=J. R.|title=Diptera. In: Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil.|date=1868|volume=2|publisher=B. K. Gerold’s Sohn|location=Wien|pages=Pt. 1, vi + 388 pp}}</ref>
*''[[Anomioptera]]'' <small>[[Ignaz Rudolph Schiner|Schiner]], 1868</small><ref name="Schiner1868">{{cite book|last1=Schiner|first1=J. R.|title=Diptera. In: Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil.|date=1868|volume=2|publisher=B. K. Gerold’s Sohn|location=Wien|pages=Pt. 1, vi + 388 pp}}</ref>
*''[[Anommonia]]'' <small>Schmitz, 1917</small><ref name="Schmitz1917c">{{cite journal|last1=Schmitz|first1=H.|title= ''Anommonia'', eine neue myrmecophile Borboridengattung aus Kamerun|journal=Zoologische Mededelingen | location=Leiden |date=1917|volume=3|pages=121–126}}</ref>
*''[[Anommonia]]'' <small>Schmitz, 1917</small><ref name="Schmitz1917c">{{cite journal|last1=Schmitz|first1=H.|title= ''Anommonia'', eine neue myrmecophile Borboridengattung aus Kamerun|journal=Zoologische Mededelingen |location=Leiden |date=1917|volume=3|pages=121–126}}</ref>
*''[[Apterobiroina]]'' <small>Papp, 1979</small><ref name="Papp1979e">{{cite journal|last1=Papp|first1=László|title= On apterous and reduced-winged forms of the family Drosophilidae, Ephydridae and Sphaeroceridae (Diptera)|journal= Acta Zoologica Academiae Scientiarum Hungaricae|url=http://www.drosophila.jp/jdd/class/030709/03070947.pdf|accessdate=22 October 2017|volume=25|pages=357–374}}</ref>
*''[[Apterobiroina]]'' <small>Papp, 1979</small><ref name="Papp1979e">{{cite journal|last1=Papp|first1=László|title= On apterous and reduced-winged forms of the family Drosophilidae, Ephydridae and Sphaeroceridae (Diptera)|journal= Acta Zoologica Academiae Scientiarum Hungaricae|url=http://www.drosophila.jp/jdd/class/030709/03070947.pdf|access-date=22 October 2017|volume=25|pages=357–374}}</ref>
*''[[Apteromyia]]'' <small>Vimmer, 1929</small><ref name="Vimmer1929a">{{cite journal|last1=Vimmer|first1=A.|title=Dvě nové bezkřídlé mouchy (čel. Cypselidae) [Zwei neue fluegellosen Di- pteren - Fam. Cypselidae]|journal=Časopis Československé společnosti entomologické|date=1929|volume=26|pages=64–68}}</ref>
*''[[Apteromyia]]'' <small>Vimmer, 1929</small><ref name="Vimmer1929a">{{cite journal|last1=Vimmer|first1=A.|title=Dvě nové bezkřídlé mouchy (čel. Cypselidae) [Zwei neue fluegellosen Di- pteren - Fam. Cypselidae]|journal=Časopis Československé Společnosti Entomologické|date=1929|volume=26|pages=64–68}}</ref>
*''[[Aptilotella]]'' <small>[[Oswald Duda|Duda]], 1924</small><ref name="Duda1924c">{{cite journal|last1=Duda|first1=Oswald|title= ''Aptilotella Borgmeieri'' ♂, eine neue flügellose Borboride (Dipt.) aus Brasilien|journal= Natuurhistorisch Maandblad, Maastricht|url=http://opuscula.elte.hu/PDF/Opuscula14_1977/Papp,%20L_14_8.pdf|accessdate=22 October 2017|volume=13|issue=5|pages=74–76}}</ref>
*''[[Aptilotella]]'' <small>[[Oswald Duda|Duda]], 1924</small><ref name="Duda1924c">{{cite journal|last1=Duda|first1=Oswald|title= ''Aptilotella Borgmeieri'' ♂, eine neue flügellose Borboride (Dipt.) aus Brasilien|journal= Natuurhistorisch Maandblad, Maastricht|url=http://opuscula.elte.hu/PDF/Opuscula14_1977/Papp,%20L_14_8.pdf|access-date=22 October 2017|volume=13|issue=5|pages=74–76}}</ref>
*''[[Aptilotus]]'' <small>[[Josef Mik|Mik]], 1898</small><ref name="Mik1898">{{cite journal|last1=Mik|first1=Josef|title=Altes und neues über Dipteren.|journal=Wiener Entomologische Zeitung|volume=17|pages=196–219|url=https://www.biodiversitylibrary.org/page/12109787#page/242/mode/1up|accessdate=22 October 2017|doi=10.5962/bhl.part.3129|year=1898}}</ref>
*''[[Aptilotus]]'' <small>[[Josef Mik|Mik]], 1898</small><ref name="Mik1898">{{cite journal|last1=Mik|first1=Josef|title=Altes und neues über Dipteren.|journal=Wiener Entomologische Zeitung|volume=17|pages=196–219|url=https://www.biodiversitylibrary.org/page/12109787#page/242/mode/1up|access-date=22 October 2017|doi=10.5962/bhl.part.3129|year=1898|doi-access=free}}</ref>
*''[[Archiceroptera]]'' <small>Papp, 1977</small><ref name="Papp1977c">{{cite journal|last1=Papp|first1=László|title= A contribution to the knowledge of species of the subfamily Ceropterinae (Diptera: Sphaeroceridae)|journal= Acta Zoologica Academiae Scientiarum Hungaricae|url=http://actazool.nhmus.hu/60/2/ActaZH_2014_Vol_60_2_109.pdf|accessdate=22 October 2017|volume=23|issue=2|pages=371–385}}</ref>
*''[[Archiceroptera]]'' <small>Papp, 1977</small><ref name="Papp1977c">{{cite journal|last1=Papp|first1=László|title= A contribution to the knowledge of species of the subfamily Ceropterinae (Diptera: Sphaeroceridae)|journal= Acta Zoologica Academiae Scientiarum Hungaricae|url=http://actazool.nhmus.hu/60/2/ActaZH_2014_Vol_60_2_109.pdf|access-date=22 October 2017|volume=23|issue=2|pages=371–385}}</ref>
*''[[Archicollinella]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925"/>
*''[[Archicollinella]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925"/>
*''[[Archileptocera]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Archileptocera]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Archipterogrammoides]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Archipterogrammoides]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Aspinilimosina]]'' <small>Papp, 2004</small><ref name="Papp2004a">{{cite journal|last1=Papp|first1=László|title=Two new genera of the Oriental Sphaeroceridae (Diptera)|journal=Annales Historico-Naturales Musei Nationalis Hungarici|date=2004|volume=96|pages=103–114|url=http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_2004_Vol_96_103.pdf|accessdate=24 October 2017}}</ref>
*''[[Aspinilimosina]]'' <small>Papp, 2004</small><ref name="Papp2004a">{{cite journal|last1=Papp|first1=László|title=Two new genera of the Oriental Sphaeroceridae (Diptera)|journal=Annales Historico-Naturales Musei Nationalis Hungarici|date=2004|volume=96|pages=103–114|url=http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_2004_Vol_96_103.pdf|access-date=24 October 2017}}</ref>
*''[[Australimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Australimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Bentrovata]]'' <small>Richards, 1973</small><ref name="Richards1973">{{cite journal|last1=Richards|first1=O. W.|title=The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australian Region|journal=Australian Journal of Zoology, supplementary series|date=1973|volume=21|issue=22|pages=297–401|doi=10.1071/ajzs022}}</ref>
*''[[Bentrovata]]'' <small>Richards, 1973</small><ref name="Richards1973">{{cite journal|last1=Richards|first1=O. W.|title=The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australian Region|journal=Australian Journal of Zoology, Supplementary Series|date=1973|volume=21|issue=22|pages=297–401|doi=10.1071/ajzs022}}</ref>
*''[[Biconnecta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Biconnecta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Biroina]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Biroina]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Bitheca]]'' <small>Marshall, 1987</small><ref name="Marshall1987c">{{cite journal|last1=Marshall|first1=S. A.|title=Systematics of ''Bitheca'', a new genus of New World Sphaeroceridae (Diptera)|journal=Systematic Entomology|date=1987|volume=12|pages=355–380|issn=0307-6970|doi=10.1111/j.1365-3113.1987.tb00207.x}}</ref>
*''[[Bitheca]]'' <small>Marshall, 1987</small><ref name="Marshall1987c">{{cite journal|last1=Marshall|first1=S. A.|title=Systematics of ''Bitheca'', a new genus of New World Sphaeroceridae (Diptera)|journal=Systematic Entomology|date=1987|volume=12|issue=3|pages=355–380|issn=0307-6970|doi=10.1111/j.1365-3113.1987.tb00207.x}}</ref>
*''[[Bromeloecia]]'' <small>Spuler, 1924</small><ref name="Spuler1924d">{{cite journal|last1=Spuler|first1=Anthony|title=North American genera and subgenera of the dipterous family Borboridae|journal=Proceedings of the Academy of Natural Sciences of Philadelphia|date=1924|volume=75 | issue = 1923 |pages=369–378}}</ref>
*''[[Bromeloecia]]'' <small>Spuler, 1924</small><ref name="Spuler1924d">{{cite journal|last1=Spuler|first1=Anthony|title=North American genera and subgenera of the dipterous family Borboridae|journal=Proceedings of the Academy of Natural Sciences of Philadelphia|date=1924|volume=75 |issue=1923 |pages=369–378}}</ref>
*''[[Cephalimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Cephalimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Ceroptera]]'' <small>[[Justin Pierre Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
*''[[Ceroptera]]'' <small>[[Pierre-Justin-Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
*''[[Ceropterella]]'' <small>Richards, 1953</small><ref name="Richards1953">{{cite journal|last1=Richards|first1=O. W.|title=Two new African species of ''Ceroptera'' Meigen (Diptera, Sphaeroceridae)|journal=Proceedings of the Royal Entomological Society of London B|date=1953|volume=22|pages=5–8}}</ref>
*''[[Ceropterella]]'' <small>Richards, 1953</small><ref name="Richards1953">{{cite journal|last1=Richards|first1=O. W.|title=Two new African species of ''Ceroptera'' Meigen (Diptera, Sphaeroceridae)|journal=Proceedings of the Royal Entomological Society of London B|date=1953|volume=22|pages=5–8}}</ref>
*''[[Chaetopodella]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Chaetopodella]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Chaetosifemur]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Chaetosifemur]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Chespiritos]]'' <small>Marshall, 2000</small><ref name="Marshall2000a">{{cite journal|last1=Marshall|first1=S. A.|title=''Chespiritos'', a new genus of Limosininae (Diptera: Sphaeroceridae) from Costa Rica|journal=Proceedings of the Entomological Society of Washington|date=2000|volume=102|pages=609–612|url=https://www.biodiversitylibrary.org/page/16172520#page/135/mode/1up|accessdate=22 October 2017}}</ref>
*''[[Chespiritos]]'' <small>Marshall, 2000</small><ref name="Marshall2000a">{{cite journal|last1=Marshall|first1=S. A.|title=''Chespiritos'', a new genus of Limosininae (Diptera: Sphaeroceridae) from Costa Rica|journal=Proceedings of the Entomological Society of Washington|date=2000|volume=102|pages=609–612|url=https://www.biodiversitylibrary.org/page/16172520#page/135/mode/1up|access-date=22 October 2017}}</ref>
*''[[Coproica]]'' <small>[[Camillo Róndani|Róndani]], 1861</small>
*''[[Coproica]]'' <small>[[Camillo Rondani|Rondani]], 1861</small>
*''[[Druciatus]]'' <small>Marshall, 1995</small><ref name="Marshall1995">{{cite journal|last1=Marshall|first1=S. A .|title=''Sclerocoelus'' and ''Druciatus'', new genera of New World Sphaeroceridae (Diptera; Sphaeroceridae; Limosininae)|journal=Insecta Mundi|date=1995|volume=9|pages=283–289|url=http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1142&context=insectamundi|accessdate=22 October 2017|publisher=University of Nebraska - Lincoln}}</ref>
*''[[Druciatus]]'' <small>Marshall, 1995</small><ref name="Marshall1995">{{cite journal|last1=Marshall|first1=S. A .|title=''Sclerocoelus'' and ''Druciatus'', new genera of New World Sphaeroceridae (Diptera; Sphaeroceridae; Limosininae)|journal=Insecta Mundi|date=1995|volume=9|pages=283–289|url=http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1142&context=insectamundi|access-date=22 October 2017|publisher=University of Nebraska - Lincoln}}</ref>
*''[[Elachisoma]]'' <small>[[Camillo Róndani|Róndani]], 1880</small>
*''[[Elachisoma]]'' <small>[[Camillo Rondani|Rondani]], 1880</small>
*''[[Eulimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Eulimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Eximilimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Eximilimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Gigalimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e">{{cite journal|last1=Roháček|first1=Jindřich|title=A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II|journal=Beiträge zur Entomologie, Berlin|date=1983|volume=33|pages=3–195|url=http://www.contributions-to-entomology.org/index.php/beitraege_entomologie/article/view/1184/1183|accessdate=23 October 2017}}</ref>
*''[[Gigalimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e">{{cite journal|last1=Roháček|first1=Jindřich|title=A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II|journal=Beiträge zur Entomologie, Berlin|date=1983|volume=33|pages=3–195|url=http://www.contributions-to-entomology.org/index.php/beitraege_entomologie/article/view/1184/1183|access-date=23 October 2017}}</ref>
*''[[Giraffimyiella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Giraffimyiella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Gobersa]]'' <small>de Coninck, 1983</small><ref name="Coninck1983">{{cite journal|last1=Coninck|first1=E. de|title=''Gobersa leleupi'' n.g., n.sp., a brachypterous Sphaerocerid from the Uluguru Mountains, Tanzania (Diptera)|journal=Revue de Zoologie Africaine, Tervuren|date=1983|volume=97|issue=2|pages=337–344}}</ref>
*''[[Gobersa]]'' <small>de Coninck, 1983</small><ref name="Coninck1983">{{cite journal|last1=Coninck|first1=E. de|title=''Gobersa leleupi'' n.g., n.sp., a brachypterous Sphaerocerid from the Uluguru Mountains, Tanzania (Diptera)|journal=Revue de Zoologie Africaine, Tervuren|date=1983|volume=97|issue=2|pages=337–344}}</ref>
*''[[Gonioneura]]'' <small>[[Camillo Róndani|Róndani]], 1880</small>
*''[[Gonioneura]]'' <small>[[Camillo Rondani|Rondani]], 1880</small>
*''[[Gonitella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Gonitella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Gyretria]]'' <small>[[Günther Enderlein|Enderlein]], 1938</small><ref name="Enderlein1938">{{cite journal|last1=Enderlein|first1=Günther|editor1-last=Skottsberg|editor1-first=C.|title=60. Die Dipterenfauna der Juan-Fernandez-Inseln und der Oster-Insel|journal=The natural history of Juan Fernández and Easter Island (Zool.) Uppsala|date=1938|volume=3|pages=643–680}}</ref>
*''[[Gyretria]]'' <small>[[Günther Enderlein|Enderlein]], 1938</small><ref name="Enderlein1938">{{cite journal|last1=Enderlein|first1=Günther|editor1-last=Skottsberg|editor1-first=C.|title=60. Die Dipterenfauna der Juan-Fernandez-Inseln und der Oster-Insel|journal=The Natural History of Juan Fernández and Easter Island (Zool.) Uppsala|date=1938|volume=3|pages=643–680}}</ref>
*''[[Hellerella]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Hellerella]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Herniosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Herniosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Howickia]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
*''[[Howickia]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
*''[[Indiosina]]'' <small>Papp, 1981</small><ref name="Papp1981">{{cite journal|last1=Papp|first1=László|title=Two new genera of the apterous and subapterous Sphaeroceridae (Diptera) from India|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1981|volume=27|pages=381–388}}</ref>
*''[[Indiosina]]'' <small>Papp, 1981</small><ref name="Papp1981">{{cite journal|last1=Papp|first1=László|title=Two new genera of the apterous and subapterous Sphaeroceridae (Diptera) from India|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1981|volume=27|pages=381–388}}</ref>
*''[[Kabaria (genus)|Kabaria]]'' <small>Richards, 1966</small><ref name="Richards1966a">{{cite journal|last1=Richards|first1=O. W.|title=New Sphaeroceridae (Diptera) from the Kivu district, with a revision of the genus ''Aluligera'' Richards|journal=Revue de Zoologie et de Botanique Africaines|date=1966|pages = 201–254|volume=73}}</ref>
*''[[Kabaria (fly)|Kabaria]]'' <small>Richards, 1966</small><ref name="Richards1966a">{{cite journal|last1=Richards|first1=O. W.|title=New Sphaeroceridae (Diptera) from the Kivu district, with a revision of the genus ''Aluligera'' Richards|journal=Revue de Zoologie et de Botanique Africaines|date=1966|pages=201–254|volume=73}}</ref>
*''[[Lepidosina]]'' <small>Marshall & Buck, 2007</small><ref name="MarshallBuckLonsdale2007">{{cite journal|last1=Marshall|first1=S. A.|last2=Buck|first2=M.|last3=Lonsdale|first3=O.|title=''Lepidosina'', a new genus of New World Limosininae (Diptera: Sphaeroceridae)|journal=European Journal of Entomolog|date=2007|volume=104|issue=3|pages=573–599|url=https://www.eje.cz/pdfs/eje/2007/03/22.pdf|accessdate=24 October 2017|doi=10.14411/eje.2007.074}}</ref>
*''[[Lepidosina]]'' <small>Marshall & Buck, 2007</small><ref name="MarshallBuckLonsdale2007">{{cite journal|last1=Marshall|first1=S. A.|last2=Buck|first2=M.|last3=Lonsdale|first3=O.|title=''Lepidosina'', a new genus of New World Limosininae (Diptera: Sphaeroceridae)|journal=European Journal of Entomology|date=2007|volume=104|issue=3|pages=573–599|url=https://www.eje.cz/pdfs/eje/2007/03/22.pdf|access-date=24 October 2017|doi=10.14411/eje.2007.074|doi-access=free}}</ref>
*''[[Leptocera]]'' <small>[[Guillaume-Antoine Olivier|Olivier]], 1813</small>
*''[[Leptocera]]'' <small>[[Guillaume-Antoine Olivier|Olivier]], 1813</small>
*''[[Limomyza]]'' <small>Marshall, 1997</small><ref name="Marshall1997a">{{cite journal|last1=Marshall|first1=S. A.|title=''Limomyza'', a new genus of primitive Limosininae (Diptera: Sphaeroceridae), with five new species from United States, Mexico and Central America|journal=Proceedings of the Entomological Society of Washington|date=1997|volume=99|pages=279–289|url=https://www.biodiversitylibrary.org/page/16212946#page/289/mode/1up|accessdate=23 October 2017}}</ref>
*''[[Limomyza]]'' <small>Marshall, 1997</small><ref name="Marshall1997a">{{cite journal|last1=Marshall|first1=S. A.|title=''Limomyza'', a new genus of primitive Limosininae (Diptera: Sphaeroceridae), with five new species from United States, Mexico and Central America|journal=Proceedings of the Entomological Society of Washington|date=1997|volume=99|pages=279–289|url=https://www.biodiversitylibrary.org/page/16212946#page/289/mode/1up|access-date=23 October 2017}}</ref>
*''[[Limosina]]'' <small>[[Justin Pierre Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
*''[[Limosina]]'' <small>[[Pierre-Justin-Marie Macquart|Macquart]], 1835</small><ref name="Macquart1835"/>
*''[[Limosinella]]'' <small>Richards, 1968</small><ref name="Richards1968c">{{cite journal|last1=Richards|first1=O. W.|title=Some Sphaeroceridae (Diptera), partly short-winged, collected in South Africa by Monsieur N. Leleup|journal=Annals of the Natal Museum, Pietermaritzburg|date=1968|volume=20|pages=93–108|url=http://journals.co.za/docserver/fulltext/annals/20/1/716.pdf?expires=1508744957&id=id&accname=guest&checksum=F47D3DA66D859D7B496A6055F723207B|accessdate=23 October 2017}}</ref>
*''[[Limosinella]]'' <small>Richards, 1968</small><ref name="Richards1968c">{{cite journal|last1=Richards|first1=O. W.|title=Some Sphaeroceridae (Diptera), partly short-winged, collected in South Africa by Monsieur N. Leleup|journal=Annals of the Natal Museum, Pietermaritzburg|date=1968|volume=20|pages=93–108|url=http://journals.co.za/docserver/fulltext/annals/20/1/716.pdf?expires=1508744957&id=id&accname=guest&checksum=F47D3DA66D859D7B496A6055F723207B|access-date=23 October 2017}}</ref>
*''[[Lobeliomyia]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
*''[[Lobeliomyia]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
*''[[Mesaptilotus]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
*''[[Mesaptilotus]]'' <small>Richards, 1951</small><ref name="Richards1951a"/>
Line 156: Line 152:
*''[[Nearcticorpus]]'' <small>Roháček & Marshall, 1982</small><ref name="RoháčekMarshall1982">{{cite journal|last1=Roháček|first1=Jindřich|last2=Marshall|first2=S. A.|title=A monograph of the genera ''Puncticorpus'' Duda, 1918 and ''Nearcticorpus'' gen.n. (Diptera, Sphaeroceridae)|journal=Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere|volume=109|pages=357–398}}</ref>
*''[[Nearcticorpus]]'' <small>Roháček & Marshall, 1982</small><ref name="RoháčekMarshall1982">{{cite journal|last1=Roháček|first1=Jindřich|last2=Marshall|first2=S. A.|title=A monograph of the genera ''Puncticorpus'' Duda, 1918 and ''Nearcticorpus'' gen.n. (Diptera, Sphaeroceridae)|journal=Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere|volume=109|pages=357–398}}</ref>
*''[[Ocellipsis]]'' <small>Richards, 1938</small><ref name="Richards1938a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera Sphaeroceridae (Borboridae, Cypselidae). Mission scientifique de l'Omo |journal=Mémoires du Museum National d'Histoire Naturelle|date=1938a|volume=4|issue=40|pages=381–405|location=Paris}}</ref>
*''[[Ocellipsis]]'' <small>Richards, 1938</small><ref name="Richards1938a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera Sphaeroceridae (Borboridae, Cypselidae). Mission scientifique de l'Omo |journal=Mémoires du Museum National d'Histoire Naturelle|date=1938a|volume=4|issue=40|pages=381–405|location=Paris}}</ref>
*''[[Ocellusia]]'' <small>[[Eugène Séguy|Séguy]], 1955</small><ref name="Séguy1955">{{cite journal|last1=Séguy|first1=Eugène|title=Deux Cypsélides aptères de Madagascar (Diptères)|journal=Naturaliste Malgache|date=1955|volume=7|pages=77–81|url=http://madadoc.irenala.edu.mg/documents/8797_deux%20cypselides.pdf|accessdate=23 October 2017}}</ref>
*''[[Ocellusia]]'' <small>[[Eugène Séguy|Séguy]], 1955</small><ref name="Séguy1955">{{cite journal|last1=Séguy|first1=Eugène|title=Deux Cypsélides aptères de Madagascar (Diptères)|journal=Naturaliste Malgache|date=1955|volume=7|pages=77–81|url=http://madadoc.irenala.edu.mg/documents/8797_deux%20cypselides.pdf|access-date=23 October 2017}}</ref>
*''[[Opacifrons]]'' <small>[[Oswald Duda|Duda]], 1918</small><ref name="Duda1918">{{cite journal|last1=Duda|first1=Oswald|title=Revision der europäischen Arten der Gattung ''Limosina'' Macquart (Dipteren)|journal=Abhandlungen der k.k. zoologisch-botanischen Gesellschaft in Wien|date=1918|volume=10|issue=1|pages=1–240}}</ref>
*''[[Opacifrons]]'' <small>[[Oswald Duda|Duda]], 1918</small><ref name="Duda1918">{{cite journal|last1=Duda|first1=Oswald|title=Revision der europäischen Arten der Gattung ''Limosina'' Macquart (Dipteren)|journal=Abhandlungen der k.k. Zoologisch-botanischen Gesellschaft in Wien|date=1918|volume=10|issue=1|pages=1–240}}</ref>
*''[[Opalimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Opalimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Oribatomyia]]'' <small>Richards, 1960</small><ref name="Richards1960a">{{cite journal|last1=Richards|first1=O. W.|title=Further brachypterous and apterous Sphaeroceridae in the collection of the Musée royal du Congo belge|journal=Revue de Zoologie et de Botanique Africaines|date=1960|volume=61|pages=320–332}}</ref>
*''[[Oribatomyia]]'' <small>Richards, 1960</small><ref name="Richards1960a">{{cite journal|last1=Richards|first1=O. W.|title=Further brachypterous and apterous Sphaeroceridae in the collection of the Musée royal du Congo belge|journal=Revue de Zoologie et de Botanique Africaines|date=1960|volume=61|pages=320–332}}</ref>
Line 163: Line 159:
*''[[Palaeocoprina]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Palaeocoprina]]'' <small>[[Oswald Duda|Duda]], 1920</small><ref name="Duda1920b"/>
*''[[Papualimosina]]'' <small>Hayashi, 2006</small><ref name="Hayashi2006b">{{cite journal|last1=Hayashi|first1=T.|title=Description of a new genus ''Papualimosina'' (Diptera, Sphaeroceridae) from the Australasian Region|journal=Japanese Journal of Systematic Entomology|date=2006|volume=12|pages=225–229}}</ref>
*''[[Papualimosina]]'' <small>Hayashi, 2006</small><ref name="Hayashi2006b">{{cite journal|last1=Hayashi|first1=T.|title=Description of a new genus ''Papualimosina'' (Diptera, Sphaeroceridae) from the Australasian Region|journal=Japanese Journal of Systematic Entomology|date=2006|volume=12|pages=225–229}}</ref>
*''[[Papuella]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Papuellicesa]]'' <small>Koçak & Kemal, 2010</small><ref name="KoçakKemal2010">{{cite journal|last1=Koçak|first1=A. Ö|last2=Kemal|first2=M.|title=Nomenclatural notes on the genus group names of the order Diptera|journal=Miscellaneous Papers, Centre for Entomological Studies Ankara|date=2010|volume=151|pages=5–7}}</ref>
*''[[Papuellicesa]]'' <small>Koçak & Kemal, 2010</small><ref name="KoçakKemal2010">{{cite journal|last1=Koçak|first1=A. Ö|last2=Kemal|first2=M.|title=Nomenclatural notes on the genus group names of the order Diptera|journal=Miscellaneous Papers, Centre for Entomological Studies Ankara|date=2010|volume=151|pages=5–7}}</ref>
*''[[Paracuminiseta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Paracuminiseta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
Line 173: Line 168:
*''[[Paraptilotus]]'' <small>Richards, 1938</small><ref name="Richards1938a"/>
*''[[Paraptilotus]]'' <small>Richards, 1938</small><ref name="Richards1938a"/>
*''[[Parasclerocoelus]]'' <small>Marshall & Dong, 2008</small><ref name="MarshallDong2008">{{cite journal|last1=Marshall|first1=Stephen A.|last2=Dong|first2=H.|title=''Parasclerocoelus'', a new south temperate genus of Limosininae (Diptera: Sphaeroceridae)|journal=Studia Dipterologica|date=2008|volume=15|pages=223–230}}</ref>
*''[[Parasclerocoelus]]'' <small>Marshall & Dong, 2008</small><ref name="MarshallDong2008">{{cite journal|last1=Marshall|first1=Stephen A.|last2=Dong|first2=H.|title=''Parasclerocoelus'', a new south temperate genus of Limosininae (Diptera: Sphaeroceridae)|journal=Studia Dipterologica|date=2008|volume=15|pages=223–230}}</ref>
*''[[Paraspelobia]]'' <small>[[Oswald Duda|Duda]], 1938</small><ref name="Duda1938">{{cite journal|last1=Duda|first1=Oswald|editor1-last=Lindner|editor1-first=E.|title=Sphaeroceridae (Cypselidae)|journal=Die Fliegen der palaearktischen Region|date=1938|volume=6|issue=57|pages=1–182|publisher=E. Schweizerbartísche Verlagsbuchhandlung|location=Stuttgart}}</ref>
*''[[Paraspelobia]]'' <small>[[Oswald Duda|Duda]], 1938</small><ref name="Duda1938">{{cite journal|last1=Duda|first1=Oswald|editor1-last=Lindner|editor1-first=E.|title=Sphaeroceridae (Cypselidae)|journal=Die Fliegen der Paläarktischen Region|date=1938|volume=6|issue=57|pages=1–182|publisher=E. Schweizerbartísche Verlagsbuchhandlung|location=Stuttgart}}</ref>
*''[[Pellucialula]]'' <small>Papp, 2004</small><ref name="Papp2004a"/>
*''[[Pellucialula]]'' <small>Papp, 2004</small><ref name="Papp2004a"/>
*''[[Philocoprella]]'' <small>Richards, 1929</small><ref name="Richards1929">{{cite journal|last1=Richards|first1=O. W.|title=Systematic notes on the Borboridae (Diptera) with description of new species of Leptocera (Limosina)|journal=The Entomologist’s Monthly Magazine|date=1929|volume=65|pages=171–176}}</ref>
*''[[Philocoprella]]'' <small>Richards, 1929</small><ref name="Richards1929">{{cite journal|last1=Richards|first1=O. W.|title=Systematic notes on the Borboridae (Diptera) with description of new species of Leptocera (Limosina)|journal=The Entomologist's Monthly Magazine|date=1929|volume=65|pages=171–176}}</ref>
*''[[Phthitia]]'' <small>[[Günther Enderlein|Enderlein]], 1938</small><ref name="Enderlein1938"/>
*''[[Phthitia]]'' <small>[[Günther Enderlein|Enderlein]], 1938</small><ref name="Enderlein1938"/>
*''[[Piliterga]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Piliterga]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
Line 181: Line 176:
*''[[Pleuroseta]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Pleuroseta]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Poecilosomella]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925"/>
*''[[Poecilosomella]]'' <small>[[Oswald Duda|Duda]], 1925</small><ref name="Duda1925"/>
*''[[Popondetta (genus)|Popondetta]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Popondetta (fly)|Popondetta]]'' <small>Richards, 1973</small><ref name="Richards1973"/>
*''[[Pseudacuminiseta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pseudacuminiseta]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pseudaspinilimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pseudaspinilimosina]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pseudocollinella]]'' <small>[[Oswald Duda|Duda]], 1924</small><ref name="Duda1924b">{{cite journal|last1=Duda|first1=O.|title=Berichtigungen zur Revision der europäischen Arten der Gattung Limosina Macq. (Dipteren), nebst Beschreibung von sechs neuen Arten|journal=Verhandlungen der zoologisch-botanischen Gesellschaft in Wien|date=1924|volume=73 | issue = 1923 |pages=163–180|url=http://www.zobodat.at/stable/pdf/VZBG_73_0163-0180.pdf|accessdate=23 October 2017}}</ref>
*''[[Pseudocollinella]]'' <small>[[Oswald Duda|Duda]], 1924</small><ref name="Duda1924b">{{cite journal|last1=Duda|first1=O.|title=Berichtigungen zur Revision der europäischen Arten der Gattung Limosina Macq. (Dipteren), nebst Beschreibung von sechs neuen Arten|journal=Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien|date=1924|volume=73 |issue=1923 |pages=163–180|url=http://www.zobodat.at/stable/pdf/VZBG_73_0163-0180.pdf|access-date=23 October 2017}}</ref>
*''[[Pseudopterogramma]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pseudopterogramma]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Pteremis]]'' <small>[[Camillo Róndani|Róndani]], 1856</small>
*''[[Pteremis]]'' <small>[[Camillo Rondani|Rondani]], 1856</small>
*''[[Pterogramma]]'' <small>Spuler, 1923</small><ref name="Spuler1924d"/>
*''[[Pterogramma]]'' <small>Spuler, 1923</small><ref name="Spuler1924d"/>
*''[[Pterogrammoides]]'' <small>Papp, 1972</small><ref name="Papp1972c">{{cite journal|last1=Papp|first1=László|title=New genera and species of Sphaeroceridae (Diptera) from New Guinea|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1972|volume=18|pages=101–115}}</ref>
*''[[Pterogrammoides]]'' <small>Papp, 1972</small><ref name="Papp1972c">{{cite journal|last1=Papp|first1=László|title=New genera and species of Sphaeroceridae (Diptera) from New Guinea|journal=Acta Zoologica Academiae Scientiarum Hungaricae|date=1972|volume=18|pages=101–115}}</ref>
Line 197: Line 192:
*''[[Rudolfina]]'' <small>Roháček, 1987</small><ref name="Roháček1987a">{{cite journal|last1=Roháček|first1=Jindřich|title=Replacement name for ''Rudolfia'' Roháček, 1982 (Diptera, Sphaeroceridae), with first record of ''R. rozkosnyi'' from northern Europe|journal=Acta Entomologica Bohemoslovaca|date=1987a|volume=84|pages=474–476}}</ref>
*''[[Rudolfina]]'' <small>Roháček, 1987</small><ref name="Roháček1987a">{{cite journal|last1=Roháček|first1=Jindřich|title=Replacement name for ''Rudolfia'' Roháček, 1982 (Diptera, Sphaeroceridae), with first record of ''R. rozkosnyi'' from northern Europe|journal=Acta Entomologica Bohemoslovaca|date=1987a|volume=84|pages=474–476}}</ref>
*''[[Sclerocoelus]]'' <small>Marshall, 1995</small><ref name="Marshall1995"/>
*''[[Sclerocoelus]]'' <small>Marshall, 1995</small><ref name="Marshall1995"/>
*''[[Scutelliseta]]'' <small>Richards, 1960</small><ref name="Richards1960b">{{cite journal|last1=Richards|first1=O. W.|title=A genus and species of wingless fly from South Africa (Diptera, Sphaeroceridae)|journal=Annals of the Natal Museum|date=1960|volume=15|pages=71–73|url=http://journals.co.za/docserver/fulltext/annals/15/5/810.pdf?expires=1508766003&id=id&accname=guest&checksum=9C8FBED1989A447A09274D2BA8C614F9|accessdate=23 October 2017}}</ref>
*''[[Scutelliseta]]'' <small>Richards, 1960</small><ref name="Richards1960b">{{cite journal|last1=Richards|first1=O. W.|title=A genus and species of wingless fly from South Africa (Diptera, Sphaeroceridae)|journal=Annals of the Natal Museum|date=1960|volume=15|pages=71–73|url=http://journals.co.za/docserver/fulltext/annals/15/5/810.pdf?expires=1508766003&id=id&accname=guest&checksum=9C8FBED1989A447A09274D2BA8C614F9|access-date=23 October 2017}}</ref>
*''[[Setositibiella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Setositibiella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Siphlopteryx]]'' <small>[[Günther Enderlein|Enderlein]], 1908</small><ref name="Enderlein1908b">{{cite book|last1=Enderlein|first1=Günther|editor1-last=Drygalski|editor1-first=E. D. von|title=Die Insekten des antarktischen Gebietes. In Deutsch Südpolar-Expedition 1901-1903 im Auftrage des Reichsamtes des Innern heraus- gegeben von Erich von Drygalski, Leiter der Expedition. X. Zoologie|volume=2 | issue = 4 |date=1908|publisher=G. Reimer,|location=Berlin|pages=361–528}}</ref>
*''[[Siphlopteryx]]'' <small>[[Günther Enderlein|Enderlein]], 1908</small><ref name="Enderlein1908b">{{cite book|last1=Enderlein|first1=Günther|editor1-last=Drygalski|editor1-first=E. D. von|title=Die Insekten des antarktischen Gebietes. In Deutsch Südpolar-Expedition 1901-1903 im Auftrage des Reichsamtes des Innern heraus- gegeben von Erich von Drygalski, Leiter der Expedition. X. Zoologie|volume=2 |issue=4 |date=1908|publisher=G. Reimer|location=Berlin|pages=361–528}}</ref>
*''[[Spelobia]]'' <small>Spuler, 1924</small><ref name="Spuler1924d"/>
*''[[Spelobia]]'' <small>Spuler, 1924</small><ref name="Spuler1924d"/>
*''[[Spinilimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Spinilimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
Line 208: Line 203:
*''[[Trachyopella]]'' <small>[[Oswald Duda|Duda]], 1918</small><ref name="Duda1918"/>
*''[[Trachyopella]]'' <small>[[Oswald Duda|Duda]], 1918</small><ref name="Duda1918"/>
*''[[Trilobitella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Trilobitella]]'' <small>Papp, 2008</small><ref name="Papp2008b"/>
*''[[Trisetomyia]]'' <small>Richards, 1965</small><ref name="Richards1965a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera, Flightless Sphaeroceridae and Chloropidae. Mission zoologique de l'I.R.S.A.C. en Afrique orientale. (P. Basilewsky et N. Leleup, 1957)|journal=Annales du Musée Royal de l’Afrique Central, Tervuren, Série in 8 o , Sciences Zoologiques|date=1965|volume=138|pages=415–464}}</ref>
*''[[Trisetomyia]]'' <small>Richards, 1965</small><ref name="Richards1965a">{{cite journal|last1=Richards|first1=O. W.|title=Diptera, Flightless Sphaeroceridae and Chloropidae. Mission zoologique de l'I.R.S.A.C. en Afrique orientale. (P. Basilewsky et N. Leleup, 1957)|journal=Annales du Musée Royal de l'Afrique Centrale, Tervuren, Série in 8 o, Sciences Zoologiques|date=1965|volume=138|pages=415–464}}</ref>
*''[[Xenolimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
*''[[Xenolimosina]]'' <small>Roháček, 1983</small><ref name="Roháček1983e"/>
}}
}}
Line 222: Line 217:
==References==
==References==
* {{aut|Roháček, Jindřich (ed.)}} (2001): ''World Catalogue of Sphaeroceridae''. Slezské zemské muzeum, Opava, Czech Republic. <small>{{ISBN|80-86224-21-X}}</small> [http://www.uoguelph.ca/~samarsha/catalog.htm PDF fulltext without images]
* {{aut|Roháček, Jindřich (ed.)}} (2001): ''World Catalogue of Sphaeroceridae''. Slezské zemské muzeum, Opava, Czech Republic. <small>{{ISBN|80-86224-21-X}}</small> [http://www.uoguelph.ca/~samarsha/catalog.htm PDF fulltext without images]
*K. G. V. Smith, 1989 An introduction to the immature stages of British Flies. Diptera Larvae, with notes on eggs, puparia and pupae.''[[Royal Entomological Society Handbooks|Handbooks for the Identification of British Insects]]'' Vol 10 Part 14. [http://www.royensoc.co.uk/content/out-print-handbooks pdf ] download manual (two parts Main text and figures index)
*K. G. V. Smith, 1989 An introduction to the immature stages of British Flies. Diptera Larvae, with notes on eggs, puparia and pupae.''[[Royal Entomological Society Handbooks|Handbooks for the Identification of British Insects]]'' Vol 10 Part 14. [https://web.archive.org/web/20140209094357/http://www.royensoc.co.uk/content/out-print-handbooks pdf ] download manual (two parts Main text and figures index)


==Further reading==
==Further reading==
* [[Oswald Duda]],1938. 57. ''Sphaeroceridae'' (Cypselidae). In Lindner, E. (ed.): ''Die Fliegen der Paläarktischen Region'' Vol.6, 182 pp., E. Schweizerbart.sche Verlagsbuchhandlung, Stuttgart.
'''Important works on Sphaeroceridae'''
* [[Oswald Duda]],1938. 57. ''Sphaeroceridae'' (Cypselidae). In Lindner, E. (ed.): ''Die Fliegen der palaearktischen Region'' Vol.6, 182 pp., E. Schweizerbart.sche Verlagsbuchhandlung, Stuttgart.
* Scientific papers by [[Theodor Becker]]
* Scientific papers by [[Theodor Becker]]
*{{cite journal | last1 = Hackman | first1 = W | year = 1965 | title = On the genus Copromyza Fall. (Dipt., Sphaeroceridae), with special reference to the Finnish species | url = | journal = Notul. ent. | volume = 45 | issue = | pages = 33–46 }}
*{{cite journal |last1=Hackman |first1=W |year=1965 |title=On the genus Copromyza Fall. (Dipt., Sphaeroceridae), with special reference to the Finnish species |journal=Notul. Ent. |volume=45 |pages=33–46 }}
* Pitkin, B.R. (1988). Lesser dung flies. Diptera: Sphaeroceridae. ''[[Royal Entomological Society Handbooks|Handbooks for the Identification of British Insects]]'' 10(5e). London: Royal Entomological Society. {{ISBN|0-901546-67-4}}
* Pitkin, B.R. (1988). Lesser dung flies. Diptera: Sphaeroceridae. ''[[Royal Entomological Society Handbooks|Handbooks for the Identification of British Insects]]'' 10(5e). London: Royal Entomological Society. {{ISBN|0-901546-67-4}}
* {{cite journal | last1 = Richards | first1 = O.W. | year = 1930 | title = The British species of Sphaeroceridae (Borboridae:Diptera) | url = | journal = Proceedings of the Zoological Society of London | volume = 18 | issue = | pages = 261–345 }}
* {{cite journal |last1=Richards |first1=O.W. |year=1930 |title=The British species of Sphaeroceridae (Borboridae:Diptera) |journal=Proceedings of the Zoological Society of London |volume=18 |pages=261–345 }}
* Rohácek, J. (1982-5). A monograph and reclassification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe, parts 1 to 4. Beitrage zur Entomologie
* Rohácek, J. (1982-5). A monograph and reclassification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe, parts 1 to 4. Beitrage zur Entomologie
*[[Eugene Seguy]]. 1934. Cypselidae. ''[[Faune de France]]'' volume 28, pp.&nbsp;444–473. [http://www.faunedefrance.org/BibliothequeVirtuelleNumerique|Bibliothèque virtuelle numérique]
*[[Eugene Seguy]]. 1934. Cypselidae. ''[[Faune de France]]'' volume 28, pp.&nbsp;444–473. [https://web.archive.org/web/20131014031511/http://www.faunedefrance.org/BibliothequeVirtuelleNumerique%7CBiblioth%C3%A8que virtuelle numérique]


==External links==
==External links==
{{commons category}}
{{Commons category}}
{{wikispecies}}
* [http://delta-intkey.com/britin/dip/www/sphaeroc.htm Family description and images]
* [http://delta-intkey.com/britin/dip/www/sphaeroc.htm Family description and images]
* [http://www.ozane.com/profiles/sphaeroceridfly.html Page on Sphaeroceridae in houses]
* [http://www.ozane.com/profiles/sphaeroceridfly.html Page on Sphaeroceridae in houses]
* [http://www.diptera.info/photogallery.php?album_id=47 Image Gallery from Dipter.info]
* [http://www.diptera.info/photogallery.php?album_id=47 Image Gallery from Dipter.info]
* [http://eol.org/pages/9013/media Family Sphaeroceridae at EOL] Images
* [http://popgen0146uns50.unimaas.nl/~jlindsey/commanster/Insects/Flies/SpFlies/Leptocera.limosa.jpg Picture of ''Leptocera limosa'', a typical sphaerocerid]
* [http://popgen0146uns50.unimaas.nl/~jlindsey/commanster/Insects/Flies/SpFlies/Leptocera.limosa.jpg Picture of ''Leptocera limosa'', a typical sphaerocerid]
* [http://www.drawwing.org/insects/diptera/sphaeroceridae Wing venation]
* [http://www.drawwing.org/insects/diptera/sphaeroceridae Wing venation]


'''Species lists'''
'''Species lists'''
* [http://www.faunaeur.org/full_results.php?id=10958 European species list]
* [https://web.archive.org/web/20051031044417/http://www.faunaeur.org/full_results.php?id=10958 European species list]
* [http://www.nearctica.com/nomina/main.htm Nearctic species list]
* [http://www.nearctica.com/nomina/main.htm Nearctic species list]
* [http://hbs.bishopmuseum.org/aocat/sphaero.html Australasian/Oceanian species list]
* [http://hbs.bishopmuseum.org/aocat/sphaero.html Australasian/Oceanian species list]
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{{Diptera|2}}
{{Diptera|2}}
{{Taxonbar|from=Q4584868}}
{{Authority control}}


[[Category:Sphaeroceridae| ]]
[[Category:Sphaeroceridae| ]]
[[Category:Brachycera families]]
[[Category:Brachycera families]]
[[Category:Taxa named by Justin Pierre Marie Macquart]]
[[Category:Taxa named by Pierre-Justin-Marie Macquart]]
[[Category:Articles containing video clips]]
[[Category:Articles containing video clips]]

Latest revision as of 14:08, 18 October 2023

Sphaeroceridae
Copromyza equina (Copromyzinae)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Superfamily: Sphaeroceroidea
Family: Sphaeroceridae
Macquart, 1835[1]
Subfamilies
Synonyms[2]
List

Sphaeroceridae are a family of true flies in the order Diptera, often called small dung flies, lesser dung flies or lesser corpse flies due to their saprophagous habits. They belong to the typical fly suborder Brachycera as can be seen by their short antennae, and more precisely they are members of the section Schizophora. There are over 1,300 species and about 125 genera accepted as valid today, but new taxa are still being described.[2]

Unlike the large "corpse flies" or blow-flies of the family Calliphoridae and the large dung flies of the family Scathophagidae, the small dung flies are members of the schizophoran subsection Acalyptratae. Among their superfamily Sphaeroceroidea, they seem to be particularly close relatives of the family Heleomyzidae.[2]

Sphaeroceridae morphological details

Description and ecology[edit]

Housefly (left) vs. small dung fly (right)

Dung flies are small to minute, usually dull-colored flies with characteristically thickened first tarsomere of the posterior leg. The first tarsal segment is less than 1+1/2 times as long as the second tarsal segment and dilated. The crossvein separating the second basal and discal cells is missing. Veins four and five often fade apically. They occur all over the world except in regions with permanent ice-cover. Despite their ubiquity and abundance, little is known about their economic or ecological impact. Some species are known to be parthenogenetic.

Larval stages are poorly known, but those described are slender, narrowed anteriorly, with groups of ventral spicules on creeping welts. The larva is amphipneustic (having only the anterior and posterior pairs of spiracles). The mandibles are simple, hooked, and without additional teeth. The parastomal bars are long, thin structures, fused to the tentoropharyngeal sclerite. The hypopharyngeal sclerites are long separate or connected by a sclerotized bridge; the anterior spiracle (prothoracic spiracle) is a rosette or branched. The posterior spiracles (on the anal segment) are usually on two cylindrical lobes. Each spiracle has three slit or oval openings and three or five groups of interspiracular hairs that are branched in some species.[citation needed]

The larvae are microbial grazers found in abundance in many microenvironments with decomposing organic material. Most species appear to be associated with decaying plants or fungi and they are a part of the nutrient cycle. Some species, especially cave species, are polysaprophagous. Many species are associated with various kinds of faeces including human faeces; there are a few carrion-feeding species. These, however, are extremely abundant and are important components of the carrion-insect community. Sphaerocerids that abound in economically important decomposer communities such as compost and manure, and some decay cycles such as the wrack (seaweed) cycle are mediated by sphaerocerid-dominated insect communities.

As their microbe-associated habits suggest, sphaerocerids may carry many pathogenic microorganisms.[3] Although their reclusive habits preclude a major role in disease transmission; some can present a public health hazard on occasion or act as a warning of one. For instance Leptocera caenosa and other sphaerocerids are associated with blocked sewage drains.[4] Some species occasionally reach high population levels in food-processing plants and other buildings where they may indicate blocked drains, waste accumulation and inadequate hygiene. One species, Poecilosomella angulata, has been implicated in human intestinal myiasis[5] They have been implicated as the major means by which nematodes are disseminated among mushroom houses. Sphaeoceridae often coexist with muscoids especially Fannia canicularis and Musca domestica in the complex manure ecosystem of poultry houses, and other confined-animal facilities. Here the sphaeocerids are prey for mites and beetles, which themselves also feed on the immatures of muscoid flies reducing the population of the more problematic muscoids.[6] Carrion-feeding species are useful post mortem interval indicators in forensic entomology.

Genera[edit]

The genera are arranged alphabetically according to subfamily; these are arranged in the presumed phylogenetic sequence from the most ancestral to the most advanced:[2]

Subfamily Tucminae Marshall, 1996[7]

Subfamily Copromyzinae Stenhammar, 1855[9]

Subfamily Sphaerocerinae Macquart, 1835[1]

Sphaerocera cf. curvipes of the Sphaerocerinae

Subfamily Homalomitrinae Roháček & Marshall, 1998[31]

Limosina silvatica of the Limosininae

Subfamily Limosininae Frey, 1921[33]

See also[edit]

Footnotes[edit]

  1. ^ a b c d e Macquart1835, P. J. M. Histoire Naturelle des insectes. Dipteres. Tome deuxieme. Paris: Roret. pp. 710, 12 pl.{{cite book}}: CS1 maint: numeric names: authors list (link)
  2. ^ a b c d Rohček, Jindřich; Marshall, Stephen A.; Norrbom, Allen L.; Buck, Matthias; Quiros, Dora Isabel; Smith, Ian (2001). Rohček, Jindřich (ed.). World Catalog of Sphaeroceridae (Diptera). Opava: Slezské Zemské Muzeum. pp. 1–414. ISBN 978-8086224213. Retrieved 3 September 2017.
  3. ^ Greenberg, B., 1971. Flies and Disease, volume I: Ecology, Classification, and Biotic Association. Princeton University Press, Princeton, New Jersey.
  4. ^ Fredeen, F. J. H.; Taylor, M. E. (1964). "Borborids (Diptera: Sphaeroceridae) infesting sewage disposal tanks, with notes on the life cycle, behavior and control of Leptocera(Leptocera) caenosa (Rondani)". Can. Entomol. 96 (5): 801–808. doi:10.4039/ent96801-5.
  5. ^ McKibben, J.W.; Micks, D.W. (1956). "Report of a case of human intestinal myiasis caused by Leptocera venalicia". American Journal of Tropical Medicine and Hygiene. 5 (5): 929–32. doi:10.4269/ajtmh.1956.5.929. PMID 13362764.
  6. ^ Axtell, R. C. (1985). Chapter 16: "Poultry Pests". In: Livestock Entomology (Williams et al., editors), Wiley & Sons, New York. pp. 269–293.
  7. ^ Marshall, S. A. (1996). "Tucma fritzi, a new species in the enigmatic genus Tucma Mourguès-Schurter (Diptera, Sphaeroceridae, Tucminae new subfamily)". Studia Dipterologica (3): 283–288.
  8. ^ Mourguès-Schurter, L. (1987). "Tucma, novo gènero de Copromyzinae para a AmÈrica do Sul (Diptera, Sphaeroceridae)". Revista Brasileira de Entomologia, S"o Paulo. 31 (1): 118–122.
  9. ^ Stenhammar, C. (1855). "Skandinaviens Copromyzine granskade och beskrifne". Kongliga Vetenskaps-Akademiens Handlingar. Ser. 3. 1853. Stockholm: 257–442.
  10. ^ a b c Hedicke, H. (1923). "Nomina nova. IV". Deutsche entomologische Zeitschrift. 1923 (4): 431. doi:10.1002/mmnd.48019230413.
  11. ^ a b Duda, Oswald (1923). "Revision der altweltlichen Arten der Gattung Borborus (Cypsela) Meigen (Dipteren)". Archiv für Naturgeschichte. Abteilung A. 89 (4): 35–112. Retrieved 3 September 2017.
  12. ^ Enderlein, Günther (1909). "Antrops truncipennis, eine neue Borboridengattung von Feuerland". Zoologischer Anzeiger. 34: 225–230. Retrieved 4 September 2017.
  13. ^ Duda, Oswald (1921). "Fiebrigella und Archiborborus, zwei neue südamerikanische Borboriden-gattungen (Dipteren)". Tijdschrift voor Entomologie. 64: 119–146. Retrieved 4 September 2017.
  14. ^ Richards, O. W. (1961). "Diptera (Sphaeroceridae) from south Chile". Proceedings of the Royal Entomological Society of London B. 30: 57–68.
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References[edit]

  • Roháček, Jindřich (ed.) (2001): World Catalogue of Sphaeroceridae. Slezské zemské muzeum, Opava, Czech Republic. ISBN 80-86224-21-X PDF fulltext without images
  • K. G. V. Smith, 1989 An introduction to the immature stages of British Flies. Diptera Larvae, with notes on eggs, puparia and pupae.Handbooks for the Identification of British Insects Vol 10 Part 14. pdf download manual (two parts Main text and figures index)

Further reading[edit]

  • Oswald Duda,1938. 57. Sphaeroceridae (Cypselidae). In Lindner, E. (ed.): Die Fliegen der Paläarktischen Region Vol.6, 182 pp., E. Schweizerbart.sche Verlagsbuchhandlung, Stuttgart.
  • Scientific papers by Theodor Becker
  • Hackman, W (1965). "On the genus Copromyza Fall. (Dipt., Sphaeroceridae), with special reference to the Finnish species". Notul. Ent. 45: 33–46.
  • Pitkin, B.R. (1988). Lesser dung flies. Diptera: Sphaeroceridae. Handbooks for the Identification of British Insects 10(5e). London: Royal Entomological Society. ISBN 0-901546-67-4
  • Richards, O.W. (1930). "The British species of Sphaeroceridae (Borboridae:Diptera)". Proceedings of the Zoological Society of London. 18: 261–345.
  • Rohácek, J. (1982-5). A monograph and reclassification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe, parts 1 to 4. Beitrage zur Entomologie
  • Eugene Seguy. 1934. Cypselidae. Faune de France volume 28, pp. 444–473. virtuelle numérique

External links[edit]

Species lists