Abida vergniesiana

Abida vergniesiana
	Abida vergniesiana
Systematics
Subordination :	Land snails (Stylommatophora)
Superfamily :	Pupilloidea
Family :	Corn snails (Chondrinidae)
Subfamily :	Chondrininae
Genre :	Abida
Type :	Abida vergniesiana
Scientific name
	Abida vergniesiana
	( Charpentier in Küster , 1850)

Abida vergniesiana is a species of grain snail (Chondrinidae) from the suborder of land snails (Stylommatophora).

features

The slim to very slim-spindle-shaped to cylindrical-spindle-shaped housing is 5.5 to 8.7 mm high and 2.0 to 2.4 mm wide. It has 8 to 10 slightly to very slightly curved turns. The greatest width is reached approximately in the middle of the housing. The surface is regularly and finely striped. At the edge of the mouth, a moderately strong to strong thickened white lip is formed, but it is not swell-shaped. The mouth edges are connected to one another in the parietal area and can be pulled forward by up to 0.8 mm. The parietalis is strongly developed in the mouth reinforcement, and an angularis is always present. In contrast, a spiral is rarely present, it is then not connected to the angularis. Usually a subangularis is also missing; occasionally there is a small hump at this point, connected to the angularis. The columellaris is more strongly developed than the infracolumellaris. The latter, however, extends further forward to the edge of the mouth. The superior palatalis, the inferior palatalis and the infrapalatalis are designed as double humps and extend forward to the lip of the mouth. Occasionally a weak suprapalatalis is also present. The last turn is somewhat narrowed and flattened more or less at an angle. The base is strongly keeled. In the area of the infrapalatalis, the housing is slightly dented. The mouth rim runs almost vertically in the palate area (palatal). The navel is narrow and barely recognizable when viewed vertically from below.

In the male part of the sexual apparatus, the penis and epiphallus in the lower penis area are fused together and form a loop. Both halves of the bow are about the same thickness. the penis / epiphallus transition is sometimes marked by a constriction. The transition is below the loop bend to the epiphallus. The "free" penis, i.e. H. from the confluence with the atrium to the point of adhesion with the epiphallus is short to very short. ** In the proximal part of the penis, i.e. H. towards the confluence with the atrium, the first third to the first half of the inner wall shows no particular structure. in the further course, an indistinct longitudinal structure and very fine transverse wrinkles appear, which become somewhat coarser after the bend shortly before the transition to the epiphallus. The lumen widens further towards the epiphallus, and a rough transverse wrinkle can be seen on the inner wall. This initially becomes finer as the epiphallus progresses, and a longitudinal structure appears again on the inner wall. In the last half or last third of the epiphallus (in front of the confluence of the vas deferens) this longitudinal structure is twisted spirally, and fine transverse wrinkles appear. The penis retractor starts at the boundary between the first and second third of the penile loop. In the female part of the reproductive system, the vagina is comparatively very long, the free fallopian tube very short; usually the ratio is greater than 6: 1. The vagina is also three to four times longer than the "free" penis. The seminal vesicle is very long and reaches or towers above the albumin gland. The actual bladder is hardly thicker than the comparatively thick stem. At the branch / free fallopian tube, this is about two to three times as wide as the free fallopian tube. The radula has 15 to 16 posterior teeth in a semi-transverse row next to the central tooth.

Similar species

Abida vergniesiana is similar to Abida occidentalis , but differs in the differently designed palatal folds. In Abida pyrenaearia the columellaris is also more strongly developed than the subcolumellaris. However, in this species the Columellaris extends further forward.

Geographical distribution and habitat

The distribution area of Abida vergniesiana extends from Andorra to the upper valley of the Ariège and into the valley of its tributary Videssos in the French northern Pyrenees.

Taxonomy and nomenclature

The taxon was in today's opinion in 1850 by Carl Heinrich sexton in "Systematic shells Cabinet of Martini and Chemnitz" in the first band, 15 division, on page 103 and Taf.14, Fig. 13-16 as Pupa vergniesiana first published (but see the objections of Welter Schultes). The "Systematic Conchylia Cabinet by Martini and Chemnitz" appeared in separate deliveries, pages 97–128 appeared in 1850, plate 14 was already included in an earlier delivery and was issued in 1847. Küster expressly writes that he received the description from Jean de Charpentier ("Communicated by Mr. Director von Charpentier, who named her to Vicdessos in honor of the Genussies de Boucheren family"). At that time it was quite common for researchers to pass their descriptions on to other researchers for publication in collective works (see also the remark under Pupa villae in the same work by Küster). According to the wording above, there is no doubt about the authorship of Jean de Charpentier.

In the following works, Pupa vergniesiana Küster was often interpreted as a synonym for Pupa pyrenaearia or other species. Edmund Gittenberger ultimately downgraded the taxon to the subspecies of Abida pyrenaearia . He distinguished two subspecies within the Abida pyrenaearia taxon : Abida pyrenaearia pyrenaearia (Michaud, 1831) and Abida pyrenaearia vergniesiana (Küster, 1850). In the molecular genetic analysis of the Chondrinidae taxa in 2010 by Bas Kokshoorn and Edmund Gittenberger, however, it turned out that Abida pyrenaearia vergniesiana is the sister taxon of the taxon Abida secale ateni , which was originally also regarded as a subspecies . In terms of molecular genetics, both taxa are far removed from the respective nominate subspecies Abida pyrenaearia pyrenaearia and Abida secale secale and were therefore regarded as independent species; d. H. Abida vergniesiana and Abida ateni . Gargominy, et al. (2011) and Fauna Europaea accept the taxon as a separate species; Fauna Europaea gives "Küster 1847" as the author. On the other hand, Welter Schultes counts the taxon to Abida pyrenaearia . Edmund Gittenberger put the following synonym for Abida (pyrenaearia) vergniesiana ; the fauna of Europaea followed him in:

Pupa vergniesiana var. Provida Westerlund, 1902

Küster gave the following locations: Vicdessos , Tarascon and Foix . Gittenberger (1973) determined Vicdessos to be the type locality.

Danger

According to the assessment of the International Union for Conservation of Nature and Natural Resources (IUCN), the species is only slightly endangered ("near threatened") despite its small distribution area. The species lives in rocky terrain. Although only 8 to 9 localities are known where the species occurs, a threat is more of a theoretical nature; building quarries or widening roads could affect some occurrences of the species.

supporting documents

literature

Edmund Gittenberger: Contributions to the knowledge of the pupillacea: III. Chondrininae. Zoologische Verhandelingen, 127 (1): 3-267, 1973 ISSN 0024-1652 PDF (pp. 142ff.).
Bas Kokshoorn, Edmund Gittenberger: Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata). Zootaxa, 2539: 1–62, 2010 Preview (PDF; 22 kB)
Francisco W. Welter-Schultes: European non-marine molluscs, a guide for species identification = identification book for European land and freshwater mollusks. A1 – A3 p., 679 p., Q1 – Q78 p., Planet Poster Ed., Göttingen 2012, ISBN 978-3-933922-75-5 .
Francisco W. Welter-Schultes: Systematisches Conchylien-Cabinet by Martini and Chemnitz (1837-1920), bibliography of the volumes in Göttingen. Archives of Natural History, 26: 157-203, 1999 ISSN 0260-9541 doi : 10.3366 / anh.1999.26.2.157

Individual evidence

^ Heinrich Carl Küster: The genera Pupa, Megaspira, Balea and Tornatellina. In illustrations according to nature with descriptions. Systematic Conchylia Cabinet by Martini and Chemnitz, 1 (15): 1–194, plates 1–21, Nuremberg 1847–1852 Online at www.biodiversitylibrary.org (p. 102)
↑ AnimalBase: Pupa vergniesiana. Sexton, 1850
Jump up ↑ Eugene V. Coan, Alan R. Kabat and Richard E. Petit: 2,400 Years of Malacology (10th Edition). Annex 2: Collation of the Systematisches Conchylien-Cabinet. (1837-1920). 65 p., American Malacological Society 2013 PDF
↑ "... discovered by Director von Charpentier and given to me for illustration and description." Küster (1850): p.108
↑ Olivier Gargominy, Vincent Prie, Jean-Michel Bichain, Xavier Cucherat, Benoît Fontaine: List de référence annotée des mollusques continentaux de France. Annotated checklist of the continental molluscs from France. MalaCo, 7: 307–382, 2011 PDF ( Memento from December 30, 2011 in the Internet Archive )
↑ ^a ^b Fauna Europaea: Abida vergniesiana (Kuster 1847)
↑ Gargominy, O. 2011. Abida vergniesiana. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1. on iucnredlist.org. Retrieved September 25, 2013.

annotation

↑ Welter Schultes puts up for discussion whether the taxon was not validly published by Ludwig Georg Karl Pfeiffer as early as 1848 . Pfeiffer lists a Pupa Vergnesiana ( sic ) in the synonymy of Pupa pyrenaearia and gives an indication of Plate 14, Fig. 13-16, published as early as 1847, and a brief description (p. 343). According to the International Rules for Zoological Nomenclature, this reference to a published figure and the brief characterization is theoretically sufficient for the name Pupa vergnesiana to be available if it was later (but before 1961) adopted as a valid name or as an older homonym (in this Case of vergniesiana ). Since Küster does not go into Pfeiffer's work, but describes the taxon as new vergniesiana , under a similar but somewhat different name, Küster's vergniesiana is therefore neither an emendation nor a secondary incorrect spelling. The binomial Pupa Vergnesiana Pfeiffer, 1848 was not used by any author before 1961 as an older homonym or adopted as a valid name and is therefore not an available scientific name in the sense of the zoological nomenclature rules. Even Edmund Gittenberger mentioned in 1973 the work of Pfeiffer or the name vergnesiana not.

[Küster-1] Heinrich Carl Küster: The genera Pupa, Megaspira, Balea and Tornatellina. In illustrations according to nature with descriptions. Systematic Conchylia Cabinet by Martini and Chemnitz, 1 (15): 1–194, plates 1–21, Nuremberg 1847–1852 Online at www.biodiversitylibrary.org (p. 102)

[AnimalBase-2] AnimalBase: Pupa vergniesiana. Sexton, 1850

[Coan-4] Jump up ↑ Eugene V. Coan, Alan R. Kabat and Richard E. Petit: 2,400 Years of Malacology (10th Edition). Annex 2: Collation of the Systematisches Conchylien-Cabinet. (1837-1920). 65 p., American Malacological Society 2013 PDF

[villae-5] "... discovered by Director von Charpentier and given to me for illustration and description." Küster (1850): p.108

[Gargominy-6] Olivier Gargominy, Vincent Prie, Jean-Michel Bichain, Xavier Cucherat, Benoît Fontaine: List de référence annotée des mollusques continentaux de France. Annotated checklist of the continental molluscs from France. MalaCo, 7: 307–382, 2011 PDF ( Memento from December 30, 2011 in the Internet Archive )

[Fauna_Europaea_vergniesiana-7] Fauna Europaea: Abida vergniesiana (Kuster 1847)

[IUCN-8] Gargominy, O. 2011. Abida vergniesiana. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1. on iucnredlist.org. Retrieved September 25, 2013.

[3] Welter Schultes puts up for discussion whether the taxon was not validly published by Ludwig Georg Karl Pfeiffer as early as 1848 . Pfeiffer lists a Pupa Vergnesiana ( sic ) in the synonymy of Pupa pyrenaearia and gives an indication of Plate 14, Fig. 13-16, published as early as 1847, and a brief description (p. 343). According to the International Rules for Zoological Nomenclature, this reference to a published figure and the brief characterization is theoretically sufficient for the name Pupa vergnesiana to be available if it was later (but before 1961) adopted as a valid name or as an older homonym (in this Case of vergniesiana ). Since Küster does not go into Pfeiffer's work, but describes the taxon as new vergniesiana , under a similar but somewhat different name, Küster's vergniesiana is therefore neither an emendation nor a secondary incorrect spelling. The binomial Pupa Vergnesiana Pfeiffer, 1848 was not used by any author before 1961 as an older homonym or adopted as a valid name and is therefore not an available scientific name in the sense of the zoological nomenclature rules. Even Edmund Gittenberger mentioned in 1973 the work of Pfeiffer or the name vergnesiana not.