Gut fish
Gut fish | ||||||||||||
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Carapus acus |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Carapidae | ||||||||||||
Jordan & Fowler , 1902 |
Visceral fish (Carapidae) live in tropical and subtropical regions of the Atlantic , Pacific and Indian Oceans at depths of up to 2000 meters. Echiodon drummondii also lives in the North Sea . They got their German name because some species hide in the body cavities of invertebrates .
features
Visceral fish are eel-like, elongated, the body is scaly and often transparent. The dorsal and anal fin form seams along the top and bottom of the body, which is pointed at the end, whereby the fin rays of the anal fin are always longer than the opposite fin rays of the dorsal fin. Ventral fins are always missing, the pectoral fins only in three species of the genus Encheliophis . The anus and genital opening are very far in front, directly behind the head, below the pectoral fins, at the front end of the anal fin. The gill openings are large and reach far forward, there are six to seven gill traps. In addition to the jaws, the ploughshare and the palatine bone are also dentate. Visceral fish grow to be seven to 36 centimeters long.
Way of life
Visceral fish can live freely ( Echiodon , Eurypleuron ) or together with sea cucumbers , starfish , clams or sea squirts as commensals ( Carapus , Onuxodon ) or possibly as parasites ( Encheliophis ). The fish penetrate the host first with the tail or head-on and can also live in it in pairs.
Gut fish eat small invertebrates and fry. It is controversial whether some species, as is often claimed, eat the internal organs of their hosts.
The eggs of the intestinal fish are pelagic , the larvae, which are also pelagic, first go through a stage called vexillifer , in which they carry a long appendage with three to eight leaf-like appendages. The next larval stage is called Tenuis . This relatively long larval form melts the vertebral bodies from back to front and uses the released nutrients and the like. a. for the formation of otoliths . Tenuis larvae live on the sea floor, they already resemble adult fish. The head is still quite small relative to the body and has not yet reached its full length. At this stage, intestinal fish colonize their host.
Systematics
There are 35 species in seven genera and three subfamilies:
Subfamily Carapinae
In the subfamily Carapinae, the pectoral fins are shorter than the length of the head and have 23 or fewer fin rays. The upper jaw is not protractile (cannot be extended). The number of precaudal vertebrae (all vertebrae except caudal vertebrae) is 17 to 35.
- Genus Carapus Rafinesque, 1810
- Carapus acus (Brünnich, 1768)
- Carapus bermudensis (Jones, 1874)
- Carapus dubius (Putnam, 1874)
- Carapus mourlani (Petit, 1934)
- Carapus sluiteri (Weber, 1905)
- Genus Echiodon Thompson, 1837
- Echiodon anchipterus Williams, 1984
- Echiodon atopus Anderson, 2005
- Echiodon coheni Williams, 1984
- Echiodon cryomargarites Markle, Williams & Olney, 1983
- Echiodon dawsoni Williams & Shipp, 1982
- Echiodon dentatus (Cuvier, 1829)
- Echiodon drummondii Thompson, 1837
- Echiodon exsilium rose leaf, 1961
- Echiodon neotes Markle & Olney, 1990
- Echiodon pegasus Markle & Olney, 1990
- Echiodon prionodon Parmentier, 2012
- Echiodon pukaki Markle & Olney, 1990
- Echiodon rendahli (Whitley, 1941)
- Genus Encheliophis Müller, 1842
- Encheliophis boraborensis (Kaup, 1856)
- Encheliophis chardewalli Parmentier, 2004
- Encheliophis gracilis (Bleeker, 1856)
- Encheliophis homei (Richardson, 1846)
- Encheliophis sagamianus (Tanaka, 1908)
- Encheliophis vermicularis Müller, 1842
- Encheliophis vermiops Markle & Olney, 1990
- Genus Eurypleuron Markle & Olney, 1990
- Eurypleuron cinereum (Smith, 1955)
- Eurypleuron owasianum (Matsubara, 1953)
- Genus Onuxodon Smith, 1955
- Onuxodon fowleri (Smith, 1955)
- Onuxodon margaritiferae ( Rendahl , 1921)
- Onuxodon parvibrachium (Fowler, 1927)
Subfamily Pyramodontinae
In the subfamily Pyramodontinae, the pectoral fins are as long as the length of the head and are supported by 24 to 30 fin rays. The upper jaw is protractile (can be extended). The number of precaudal vertebrae is 12 to 15.
- Genus Pyramodon Smith & Radcliffe in Radcliffe, 1913
- Pyramodon lindas Markle & Olney, 1990
- Pyramodon parini Markle & Olney, 1990
- Pyramodon punctatus (Regan, 1914)
- Pyramodon ventralis Smith & Radcliffe in Radcliffe, 1913
- Genus Snyderidia Gilbert, 1905
- Snyderidia canina Gilbert, 1905
Subfamily Tetragondacninae
The only species of the subfamily Tetragondacninae differs in numerous characteristics, e.g. B. their scaling of other intestinal fish.
- Genus Tetragondacnus Anderson & Satria, 2007
- Tetragondacnus spilotus Anderson & Satria, 2007
swell
- Joseph S. Nelson : Fishes of the World. 4th edition. John Wiley & Sons, Hoboken NJ et al. 2006, ISBN 0-471-25031-7 .
- Kurt Fiedler: Fish (= textbook of special zoology. Vol. 2: Vertebrates. Part. 2). Gustav Fischer, Jena 1991, ISBN 3-334-00338-8 .
further reading
- Parmentier Eric et al .: Remodelling of the vertebrate axis during metamorphic shrinkage in pearlfish. (2003), Journal of Fish Biology, Volume 64 Issue 1, Pages 159 - 169. doi : 10.1111 / j.1095-8649.2004.00294.x
Web links
- Pearlfish on Fishbase.org (English)
Individual evidence
- ↑ Brooke Luciano, Ashleigh Lyman, Selena McMillan, Abby Nickels: The symbiotic relationship between Sea cucumbers (Holothuriidae) and Pearlfish (Carapidae) . Text on the symbiotic relationship between sea cucumber and gut fish PDF