Filosa

Filosa
Chlorarachnion reptans
Systematics
Domain : Eukaryotes (eukaryota) without rank: Diaphoreticks without rank: Sar without rank: Rhizaria without rank: Cercozoa Sub-stem : Filosa
Scientific name
Filosa
Cavalier-Smith , 2003

The Filosa , also known as the core Cercozoa, are a large group of protists within the Cercozoa tribe .

features

The Filosa are morphologically very diverse. Among them are naked or shelled amoebas with filamentous pseudopodia and tubular mitochondria -Cristae; Zoo flagellates with two flagella ; but also photosynthetically active algae. They often have extrusomes , mostly in the form of simple, round kinetocysts , rarely elongated ones similar to trichocysts .

Systematics

The core Cercozoa, the Filosa, appear in most of the works as a monophyletic group. Most representatives, with the exception of the Chlorarachniophyta and some flagellate species, have an insertion of two amino acids on the monomer-monomer connection of the polyubiquitin gene. The size of the Filosa has changed continuously in the last few years, Pavlovsky included the following groups in 2008:

• Euglyphida
• Cercomonadida
• Heteromitida
• Cryomonadida
• Ebriida
• Thaumatomonadida
• Chlorachnea
• Desmothoracida
• Phaeodarea
• Massisteria
• Gymnophrys
• other genera

A more detailed system is presented by Cavalier-Smith:

• Superclass reticulofilosa
  • Class chlorarachnea
    • Order Chlorarachnida
  • Class Spongomonadea
    • Order Spongomonadida
      • z. B. Spongomonas, Rhipododendron
  • (Class Proteomyxidea was added to Endomyxa in 2008 ).
• About class Monadofilosa
  • Order Metromonadida
    • Family Metromonadidae
      • Metromonas
  • Class Sarcomonadea
    • Order Cercomonadida
      • Family Cercomonadidae
      • Family Heteromitidae
  • Class Thecofilosea
    • Order Tectofilosida , with e.g.
      • Family Pseudodifflugiidae , e.g. B. with
        • Cryptodifflugia
      • Family Chlamydophryidae
      • Family Psammonobiotidae
      • Family Amphitremidae
      • Family Volutellidae
    • Order Cryomonadida
  • Class Imbricatea
    • Thaumatomonadida
    • Euglyphida
  • Class Phaeodarea
• Class Granofilosea , for subdivision see there. (not assigned to a superclass)
• incertae sedis Filosa:
  • Order Metopiida
    • Metopiidae family
      • Metopion

Within the Filosa there are a large number of groups that are only known from environmental DNA samples and have not yet been described.

supporting documents

• Thomas Cavalier-Smith, Ema E.-Y. Chao: Phylogeny and Classification of Phylum Cercozoa (Protozoa) . Protist, Volume 154, 2003, pp. 341-358, doi : 10.1078 / 143446103322454112

Individual evidence

1. ^ ^{A b} Jan Pawlowski: The twilight of Sarcodina: a molecular perspective on the polyphyletic origin of amoeboid protists . Protistology, Vol. 5, 2008, pp. 281-302. (pdf, 570 kB)
2. ↑ Thomas Cavalier-Smith, Ema E.-Y. Chao: Phylogeny and Classification of Phylum Cercozoa (Protozoa) . Protist, Volume 154, 2003, pp. 341-358, doi : 10.1078 / 143446103322454112
3. ^ ^{A b} David Bass, Thomas Cavalier-Smith: Phylum-specific environmental DNA analysis reveals remarkably high global biodiversity of Cercozoa (Protozoa) . International Journal of Systematic and Evolutionary Microbiology, Volume 54, 2004, pp. 2393-2404, doi : 10.1099 / ijs.0.63229-0
4. ↑ cf. for example: David Bass, David Moreira, Purificacion Lopez-Garcıa, Stephane Polet, Ema E. Chao, Sophie von der Heyden, Jan Pawlowski, Thomas Cavalier-Smith: Polyubiquitin Insertions and the Phylogeny of Cercozoa and Rhizaria . Protist, Volume 156, 2005, pp. 149-161, doi : 10.1016 / j.protis.2005.03.001
5. ↑ David Bass, Ema E.-Y.Chao, Sergey Nikolaev, Akinori Yabuki, Ken-ichiro Ishida, Cédric Berney, Ursula Pakzad, Claudia Wylezich, Thomas Cavalier-Smith: Phylogeny of Novel Naked Filose and Reticulose Cercozoa: Granofilosea cl.n . and Proteomyxidea Revised . Protist, 2008 doi : 10.1016 / j.protis.2008.07.002