Microchilus (orchids)
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C. Presl |
Microchilus is a genus from the family of the orchid (Orchidaceae). It consists of about 164 species of herbaceous plants native to tropical America.
description
The species of the genus Microchilus form a creeping rhizome that is rooted at the nodes . The rising shoots carry the leaves evenly distributed or in a loose rosette, roughly in the lower half of the upright shoot. The leaves are broad to narrowly oval in shape, often somewhat asymmetrical, with short stalk. The tubular leaf base includes the shoot. The color of the leaves is green to brownish, sometimes with light green or silver markings.
The stem axis continues above the leaves as a multi-flowered, racemose inflorescence . The inflorescence axis is hairy and is surrounded by a few bracts . The bracts are about as long as ovary and pedicel together. The ovary is hairy and twisted, the flowers are resupinated . The petals are free. The three sepals are roughly the same shape and size, hairy on the outside of some species. The lateral sepals are spread apart, the upper sepal forms a tube with the lateral petals attached to it . The lateral petals are narrowly linear to oblong-lanceolate, broadest above the middle. The lip forms a long, cylindrical to oval spur at the base, rarely bilobed at the end. In the interior of the spur there are usually low lamellae. The lip is divided into two parts: the basal part, the hypochil, is U-shaped in cross-section and only rarely has lamellae inside. The front part, the epichil, has entire margins or ends with two lobes. The column is spindle-shaped, its lower part is fused with the edges of the lip. The stamen contains two club-shaped pollinia , each of which is connected to the adhesive disc (Viscidium) by a small stalk. The stigma consists of an undivided area that lies beneath the separating tissue between the stigma and the stamen (rostellum). The rostellum is long triangular with a two-part tip.
distribution
Microchilus is common in tropical America. The area extends from Mexico and the Bahamas in the north over Central America and the Caribbean over all of South America to about the level of the Río de la Plata in the south. The species grow in evergreen forests, occasionally also epiphytically . They occur at altitudes up to 2000 meters.
Systematics and botanical history
The genus Microchilus was established in 1827 by Karl Bořiwog Presl . Type species is Microchilus minor . The name Microchilus is derived from the Greek μικρός mikros , "small", and χεῖλος cheilos , "lip", it refers to the small petal called lip. The species have long been listed in the genus Erythrodes together with very similar plants from Southeast Asia . Microchilus is classified within the tribe Cranichideae in the subtribe Goodyerinae . According to Dressler, this can be further divided into two groups; Dressler puts it under the name Erythrodes together with the majority of the genera that do not have two clearly separated areas of scarring.
While 40 species in the genus Microchilus were counted in the work "Genera Orchidacearum", according to studies by Paul Ormerod , the number increased considerably between 2000 and 2010 and is now around 164 species:
- Microchilus aguindae Ormerod : Ecuador
- Microchilus alzatei Ormerod : Colombia
- Microchilus anchorifer (Schltr.) Ormerod : Ecuador, Bolivia, Peru
- Microchilus anderssonii Ormerod : Ecuador
- Microchilus andrei Ormerod : Colombia
- Microchilus antioquiensis Szlach. & Kolan. : The species first described in 2017 occurs in Colombia.
- Microchilus arevaloi Szlach. & Kolan. : The species first described in 2017 occurs in Colombia.
- Microchilus arietinus (Rchb.f. & Warm.) Ormerod : Tropical South America to Argentina
- Microchilus aspidogynoides Ormerod : Colombia
- Microchilus astilleroensis Ormerod : Peru
- Microchilus atalayae Ormerod : Peru
- Microchilus atwoodii Szlach. & Kolan. : The species first described in 2016 occurs in Costa Rica.
- Microchilus austrobrasiliensis (Porsch) Ormerod : Southern and southeastern Brazil
- Microchilus bangii Ormerod : Bolivia
- Microchilus bimentatus (Dressler) Ormerod : Costa Rica
- Microchilus borjaquijosae Ormerod : Ecuador
- Microchilus boyacanus Ormerod : Colombia
- Microchilus brachyplectron (Pabst) Ormerod : Brazil
- Microchilus bravocollinus Ormerod : Colombia
- Microchilus brunnescens Ormerod : Ecuador
- Microchilus buchtienii (Schltr.) Ormerod : Bolivia
- Microchilus callejasii Ormerod : Colombia
- Microchilus callophylloides (Garay) Ormerod : Ecuador
- Microchilus calophyllus (Rchb.f.) Ormerod : Costa Rica to Colombia
- Microchilus campanensis Ormerod : Peru
- Microchilus campanulatus Ormerod : Northern South America
- Microchilus canaliculatus Ormerod : Colombia
- Microchilus canarensis (Ormerod) Szlach. & Kolan. : Ecuador
- Microchilus capitatus Ormerod : Peru
- Microchilus caramantae Ormerod : Colombia
- Microchilus carbonerae Ormerod : Venezuela
- Microchilus carchiensis Ormerod : Ecuador
- Microchilus carinatus Ormerod : Colombia
- Microchilus carlos-parrae Szlach. & Kolan. : The species first described in 2014 occurs in Colombia.
- Microchilus caucanus (Schltr.) Ormerod : Colombia
- Microchilus ceratostele Ormerod : Galapagos Islands
- Microchilus chicalensis Ormerod : Ecuador
- Microchilus chingualensis Ormerod : Ecuador
- Microchilus constrictus Ormerod : Colombia, northern Brazil, northern Peru
- Microchilus crassibasis Ormerod : Ecuador
- Microchilus cundinamarcae Ormerod : Colombia
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Microchilus curviflorus Ormerod : Northwestern and northern Venezuela. With the two varieties:
- Microchilus curviflorus var. Azulitae Ormerod
- Microchilus curviflorus var. Curviflorus
- Microchilus cuscoensis Kolan. & Szlach. : The species first described in 2015 occurs in Peru.
- Microchilus dolichostachys (Schltr.) Ormerod : Colombia
- Microchilus dressleri Kolan. : The species first described in 2015 occurs in Panama.
- Microchilus dryanderae Ormerod : Colombia
- Microchilus ecuadorensis (Garay) Ormerod : Ecuador
- Microchilus ensicalcar Ormerod : Peru
- Microchilus epiphyticus (Dressler) Ormerod : Costa Rica and Panama
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Microchilus erythrodoides (Schltr.) Ormerod : Colombia to northwestern Venezuela. With two subspecies:
- Microchilus erythrodoides subsp. erythrodoides
- Microchilus erythrodoides subsp. venadosae Ormerod
- Microchilus familiaris Ormerod : Caribbean
- Microchilus federalensis (Ormerod) Meneguzzo : Brazil
- Microchilus fendleri Ormerod : Venezuela
- Microchilus folsomii Szlach. & Kolan. : The species first described in 2015 occurs in Panama.
- Microchilus forceps (Ormerod) Ormerod : Ecuador
- Microchilus fosbergii Ormerod : Colombia
- Microchilus fosteri Ormerod : Peru
- Microchilus frontinoensis Ormerod : Colombia
- Microchilus fuscatus Ormerod : Panama
- Microchilus gentryi Ormerod : Colombia
- Microchilus giraldo-gensinii Ormerod : Colombia
- Microchilus glacensis (Dod) Ormerod : Haiti
- Microchilus globosicalcar Szlach. & Kolan. : The species first described in 2014 occurs in Colombia.
- Microchilus globosus Ormerod : Colombia
- Microchilus guianensis Ormerod : Guiana and Suriname
- Microchilus haughtii Ormerod : Colombia
- Microchilus hetaerioides (Schltr.) Ormerod : Southwestern Colombia
- Microchilus hirtellus (Sw.) D.Dietr. : Caribbean
- Microchilus huangobioensis Ormerod : Colombia
- Microchilus hughjonesii Ormerod : Colombia
- Microchilus ibaguensis Ormerod : Colombia
- Microchilus integer Ormerod : Ecuador
- Microchilus kuduyariensis Ormerod : Colombia
- Microchilus laegaardii Ormerod : Ecuador
- Microchilus lamprophyllus (Linden & Rchb.f.) Ormerod : Eastern Brazil
- Microchilus laticalcar (Dod) Ormerod : Dominican Republic
- Microchilus lechleri Ormerod : Peru
- Microchilus leucostictus Rchb.f. ex Szlach. & Kolan. : The species first described in 2014 occurs in Peru.
- Microchilus libanoensis Ormerod : Colombia
- Microchilus longiflorus Ormerod : Ecuador
- Microchilus lunifer (Schltr.) Ormerod : Mexico to Costa Rica
- Microchilus luteynii Szlach. & Kolan. : The species first described in 2017 occurs in Colombia.
- Microchilus maasii Ormerod : Costa Rica to Panama
- Microchilus machalillae Ormerod : Ecuador
- Microchilus madrinanii Ormerod : Northeastern Colombia
- Microchilus major C. Presl : Colombia, Ecuador, Peru
- Microchilus marulandae Ormerod : Colombia
- Microchilus mazarunensis Szlach., S.Nowak & Baranow : The species first described in 2017 occurs in Guiana.
- Microchilus meridanus Ormerod : Venezuela
- Microchilus mexicanus (Ames) Ormerod : Mexico
- Microchilus micayvallis Ormerod : Colombia
- Microchilus microcalcar Ormerod : Colombia
- Microchilus microcaprinus Ormerod : Peru
- Microchilus minor C. Presl : Peru
- Microchilus montanus Szlach. & Kolan. : The species first described in 2014 occurs in Colombia.
- Microchilus moritzii Ormerod : Venezuela
- Microchilus mutisii Szlach. & Kolan. : The species first described in 2014 occurs in Colombia.
- Microchilus nigrescens (Schltr.) Ormerod : Honduras to Ecuador
- Microchilus nugax Ormerod : Colombia
- Microchilus ormerodianus Kolan. : The species first described in 2013 occurs in Colombia.
- Microchilus oroensis (Dodson) Ormerod : Ecuador
- Microchilus ortgiesii (Rchb.f.) Ormerod : Colombia
- Microchilus ovalis Ormerod : Panama
- Microchilus ovatus (Lindl.) D. Dietr. : Colombia, Peru to western Bolivia
- Microchilus pacaizapae Ormerod : Peru
- Microchilus paleaceus (Schltr.) Ormerod : Tropical South America
- Microchilus palmazuensis Ormerod : Peru
- Microchilus panamanicus Ormerod : Panama
- Microchilus paraisoensis Ormerod : Colombia
- Microchilus parvilabrum Ormerod : Ecuador
- Microchilus pauciflorus (Poepp. & Endl.) D.Dietr. : Bolivia to northern Brazil
- Microchilus pedrojuanensis Ormerod : Southeastern Brazil to Paraguay
- Microchilus perijanus Ormerod : Colombia
- Microchilus peruvianus Szlach. & Kolan. : The species first described in 2016 occurs in Peru.
- Microchilus pilosus Szlach. & Kolan. : The species first described in 2016 occurs in Costa Rica.
- Microchilus pimentelii Ormerod : Dominican Republic
- Microchilus plantagineus (L.) D.Dietr. : Caribbean to northern South America
- Microchilus platanilloensis Ormerod : Costa Rica
- Microchilus platysepalus Ormerod : Colombia
- Microchilus plowmanii Ormerod : Peru
- Microchilus preslii Ormerod : Peru
- Microchilus procerus (Schltr.) Ormerod : Colombia to northwestern Venezuela
- Microchilus pseudobrunnescens Ormerod : Ecuador
- Microchilus pseudocaucanus Ormerod : Colombia
- Microchilus pseudominor Ormerod : Colombia
- Microchilus putumayoensis Ormerod : Colombia
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Microchilus puyoensis Ormerod : Ecuador. With three varieties:
- Microchilus puyoensis var. Brachyceras Ormerod
- Microchilus puyoensis var. Brevipedunculatus Ormerod
- Microchilus puyoensis var. Puyoensis
- Microchilus quadratus (Garay) Ormerod : Northwestern Venezuela, Ecuador, Colombia
- Microchilus queremalensis Ormerod : Colombia
- Microchilus quetamensis Ormerod : Colombia
- Microchilus rioesmeraldae Ormerod : Colombia
- Microchilus rioitayanus Ormerod : Peru
- Microchilus riopalenquensis Ormerod : Colombia, Ecuador, Peru
- Microchilus ruizteranii Ormerod : Venezuela
- Microchilus sanpabloensis Ormerod : Ecuador
- Microchilus sarmientoi Szlach. & Kolan. : The species first described in 2014 occurs in Colombia.
- Microchilus schneideri Ormerod : Colombia
- Microchilus schultesianus (Garay) Ormerod : Trinidad
- Microchilus scrotiformis ( C. Pig ) Ormerod : Colombia to northwestern Venezuela
- Microchilus sparreorum (Garay) Ormerod : Western Colombia and Ecuador
- Microchilus sprucei (Garay) Ormerod : Ecuador
- Microchilus steyermarkii Szlach. & Kolan. : The species first described in 2016 occurs in Venezuela.
- Microchilus subquadratus Ormerod : Panama to northwestern Colombia
- Microchilus tequendamae Ormerod : Colombia
- Microchilus tessellatus Ormerod : Costa Rica and Panama
- Microchilus trianae Ormerod : Colombia
- Microchilus tridax (Rchb.f.) Ormerod : Guatemala to Panama
- Microchilus trifasciatus Ormerod : Northern Peru
- Microchilus tunquianus Ormerod : Peru
- Microchilus uribei Szlach. & Kolan. : The species first described in 2014 occurs in Venezuela.
- Microchilus valdivianus Ormerod : Colombia
- Microchilus valverdei Ormerod : Costa Rica
- Microchilus venezuelanus (Garay & Dunst.) Ormerod : Venezuela
- Microchilus ventosus Ormerod : Colombia
- Microchilus vesicifer (Rchb.f.) Ormerod : Mexico to Panama, Peru
- Microchilus vilnerae Ormerod : Colombia
- Microchilus viridissimus Ormerod : Ecuador
- Microchilus weberianus (Garay) Ormerod : Ecuador, Galapagos Islands
- Microchilus whitefoordiae Ormerod : Panama
- Microchilus xeranthum Ormerod : Colombia
- Microchilus xystophyllus (Rchb.f.) Ormerod : Colombia, Venezuela to Brazil
- Microchilus zeuxinoides (Schltr.) Ormerod : Colombia
- Microchilus zingarae Ormerod : Colombia
See also
literature
- Alec M. Pridgeon, Phillip Cribb, Mark W. Chase, Finn Rasmussen (Eds.): Genera Orchidacearum. Orchidoideae (Part 2). Vanilloideae . tape 3 . Oxford University Press, New York / Oxford 2003, ISBN 0-19-850711-9 , pp. 121-124 .
- Paul Ormerod: Studies of neotropical Goodyerinae (Orchidaceae) . In: Harvard Papers in Botany . tape 9 , no. 2 , 2005, p. 391-423 .
- Paul Ormerod: Studies of neotropical Goodyerinae (Orchidaceae) 2 . In: Harvard Papers in Botany . tape 11 , no. 2 , 2007, p. 145-177 .
- Paul Ormerod: Studies of neotropical Goodyerinae (Orchidaceae) 3 . In: Harvard Papers in Botany . tape 13 , no. 1 , 2008, p. 55-87 .
- Paul Ormerod: Studies of neotropical Goodyerinae (Orchidaceae) 4 . In: Harvard Papers in Botany . tape 14 , no. 2 , 2009, p. 111-128 .
- Paul Ormerod: Notulae Goodyerinae (IV) . In: Taiwania . tape 54 , no. 1 , 2009, p. 45-51 .
Individual evidence
- ↑ a b c d e Alec M. Pridgeon, Phillip Cribb, Mark W. Chase, Finn Rasmussen (eds.): Genera Orchidacearum. Orchidoideae (Part 2). Vanilloideae . tape 3 . Oxford University Press, New York / Oxford 2003, ISBN 0-19-850711-9 , pp. 121-124 .
- ^ Robert L. Dressler: Phylogeny and Classification of the Orchid Family . Cambridge University Press, Cambridge 1993, ISBN 0-521-45058-6 , pp. 118 .
- ↑ a b c d e f g h i j k l m n o p q r s t Rafaël Govaerts (Ed.): Microchilus. In: World Checklist of Selected Plant Families (WCSP) - The Board of Trustees of the Royal Botanic Gardens, Kew . Retrieved March 27, 2020.