Norops
Norops | ||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
Norops auratus , the type species of the genus, is widespread in southern Central and northern South America. |
||||||||||||
Systematics | ||||||||||||
|
||||||||||||
Scientific name | ||||||||||||
Norops | ||||||||||||
Wagler , 1830 |
Norops is a genus of small, neotropical lizards from the group of iguanas . Because of the close relationship with the genus anole , in which the animals were placed until recently, they are also called beta anoles . The generic name is derived from the Greek and refers to the bright colors of the type species Norops auratus (“norops” = “brilliant” or “shimmering”).
features
Norops species are small lizards and reach head-trunk lengths of 3.5 to 10.8 cm. In addition there is the long tail whose length can be several times the length of the head and torso. Females often stay a little smaller than males. However, many species show no sexual dimorphism . Norops species are usually brownish or greyish in color, less often green.
The monophyly of the genus is supported by 57 apomorphies , 48 of which are molecular biological and 9 morphological .
The morphological include the parietal eye in the parietal bone , the T-shaped interclavicle, a bone that connects the actual clavicle to the breastbone (sternum), 3 or 4 lumbar vertebrae and 6 or 7, less often 8 caudal vertebrae in front of the first vertebra capable of autotomy . Their transverse processes fork in the vertical plane. The nasal bone overlaps the side edge of the premaxillary . The splenial bone, a small bone in the lower jaw of amphibians and reptiles, is usually absent. The lower jaw of the males is usually not ornamented. Palatine teeth are missing. The mean number of ventral scales is increased to 5% of the head-trunk length.
distribution
Species of the genus Norops are found in Mexico, Central America, the Greater and Lesser Antilles , the Bahamas , the Turks and Caicos Islands , the Cayman Islands , Navassa and in South America from Colombia , Venezuela and the three Guayanas across the Amazon basin to Ecuador , in northern Peru , northern Paraguay and south to the state of São Paulo in southeastern Brazil. They also occur on various islands off South America in the north ( Isla Margarita , Curaçao , Bonaire , La Blanquilla ) and on Malpelo and Gorgona in the eastern Pacific.
species
The genus includes 175 species, one of which is only known from fossils . The species are divided into three species groups, the 18 species Norops sagrei species group in Cuba and the Bahamas, the Norops valencienni species group with 7 species in Jamaica and Grand Cayman and the species-rich, 150 species Norops auratus species group throughout the rest of the distribution area .
-
Norops sagrei species group, Cuba and Bahamas.
- Norops ahli (Barbour, 1925)
- Norops allogus (Barbour & Ramsden, 1919)
- Norops birama (Garrido, 1991)
- Norops bremeri (Barbour, 1914)
- Norops confusus (Estrada & Garrido, 1991)
- Norops delafuentei (Garrido, 1982)
- Norops guafe (Estrada & Garrido, 1992)
- Norops homolechis (Cope, 1894)
- Norops imias (Ruibal & Williams, 1961)
- Norops jubar (Schwartz, 1968)
- Norops luteosignifer (Garman, 1887)
- Norops mestrei (Barbour & Ramsden, 1919)
- Norops nelsoni (Barbour, 1914)
- Norops ophiolepis (Cope, 1861)
- Norops ordinatus (Cope, 1864)
- Norops quadriocellifer (Barbour & Ramsden, 1919)
- Norops rubribarbus (Barbour & Ramsden, 1919)
- Bahamaanolis ( Norops sagrei (Cocteau in C. Duméril & Bibron, 1837))
-
Norops valencienni species group, Jamaica and Grand Cayman.
- Norops conspersus (Garman, 1887)
- Jamaica anole ( Norops garmani (Stejneger, 1899))
- Norops grahami (Garman, 1888)
- Norops lineatopus (Gray, 1840)
- Norops opalinus (Goose, 1850)
- Norops reconditus (Underwood & Williams, 1859)
- Norops valencienni (C. Duméril & Bibron, 1837)
-
Norops pentaprion species group sensu Koehler (2010):
- Norops beckeri (Boulenger, 1881)
- Norops charlesmyersi (Koehler, 2010)
- Norops cristifer (Smith, 1968)
- Norops fungosus (Myers, 1971)
- Norops ibague (Williams, 1975)
- Norops ortonii (Cope, 1868)
- Norops pentaprion (Cope, 1862)
- Norops salvini (Boulenger, 1885)
- Norops sulcifrons (Cope, 1899)
- Norops triumphalis Kirsten E. Nicholson & Gunther Koehler, 2014
- Norops utilensis Koehler, 1996
-
Norops auratus species group
- Norops adleri (H. Smith, 1972)
- Norops altae (Dunn, 1930)
- Norops alvarezdeltoroi Nieto Montes de Oca, 1996
- Norops amplisquamosus McCranie, Wilson, & K. Williams, 1992
- Norops anisolepis (H. Smith, Burley, & Fritts, 1968)
- Norops annectens (Williams, 1974)
- Norops antonii (Boulenger, 1908)
- Norops apletophallus (Köhler & Sunyer, 2008)
- Norops aquaticus (Taylor, 1956)
- Norops auratus (Daudin, 1802) ( type species )
- Norops barkeri (Schmidt, 1939)
- Norops benedikti (Lotzkat, Bienentreu, Hertz, & Köhler, 2011)
- Norops bicaorum Koehler, 1996
- Norops biporcatus (Wiegmann, 1834)
- Norops bitectus (Cope, 1864)
- Norops bocourtii (Cope, 1875)
- Norops bombiceps (Cope, 1875)
- Norops brasiliensis (Vanzolini & Williams, 1970)
- Norops breedlovei (H. Smith & Paulson, 1968)
- Norops campbelli Koehler & E. Smith, 2008
- Norops capito (W. Peters, 1863)
- Norops carpenteri (Echelle, Echelle, & Fitch, 1971)
- Norops chrysolepis (C. Duméril & Bibron, 1837)
- Norops cobanensis (Stuart, 1942)
- Norops compressicauda (H. Smith & Kester, 1955)
- Norops concolor (Cope, 1863)
- Norops crassulus (Cope, 1864)
- Norops cryptolimifrons (Köhler & Sunyer, 2008)
- Norops cupreus (Hallowell, “1860”, 1861)
- Norops cuprinus (H. Smith, 1964)
- Norops cusuco McCranie, Koehler, & Wilson, 2000
- Norops cymbops (Cope, 1864)
- Norops dariense (Fitch & Siegel, 1984)
- Norops datzorum (Köhler, Ponce, Sunyer, & Batista, 2007)
- Norops dollfusianus (Bocourt, 1873)
- Norops duellmani (Fitch & Henderson, 1973)
- Norops dunni (H. Smith, 1936)
- Norops eewi (Roze, 1958)
- Norops electrum (Lazell, 1965) fossil
- Norops forbesi (H. Smith & Van Gelder, 1955)
- Norops fortunensis (Arosemana & Ibáñez, 1993)
- Norops fuscoauratus (D'Orbigny, 1817)
- Norops gadovii (Boulenger, 1905)
- Norops gibbiceps (Cope, 1864)
- Norops godmani (Boulenger, 1885)
- Norops gracilipes (Boulenger, 1898)
- Norops granuliceps (Boulenger, 1898)
- Norops gruuo (Köhler, Ponce, Sunyer, & Batista, 2007)
- Norops Haguei (Stuart, 1942)
- Norops hobartsmithi (Nieto-Montes de Oca, 2001)
- Norops humilis (W. Peters, 1863)
- Norops intermedius (W. Peters, 1863)
- Norops isthmicus (Fitch, 1978)
- Norops johnmeyeri (Wilson & McCranie, 1982)
- Norops kemptoni (Dunn, 1940)
- Norops kreutzi McCranie, Koehler, & Wilson, 2000
- Norops laeviventris (Weigmann, 1834)
- Norops lemniscatus (Boulenger, 1898)
- Norops lemurinus (Cope, 1861)
- Norops limifrons (Cope, 1862)
- Norops lineatus (Daudin, 1802)
- Norops liogaster (Boulenger, 1905)
- Norops lionotus (Cope, 1861)
- Norops loveridgei (Schmidt, 1936)
- Norops Lynchi (Miyata, 1985)
- Norops lyra (Poe, Velasco, Miyata, & Williams, 2009)
- Norops macrinii (H. Smith, 1968)
- Norops macrolepis (Boulenger, 1898)
- Norops macrophallus (Werner, 1917)
- Norops maculiventris (Boulenger, 1898)
- Norops magnaphallus (Poe & Ibáñez, 2007)
- Norops mariarum (Barbour, 1932)
- Norops matudai (H. Smith, 1956)
- Norops medemi (Ayala & Williams, 1988)
- Norops megapholidotus (H. Smith, 1933)
- Norops meridionalis (Boettger, 1885)
- Norops microlepidotus (Davis, 1954)
- Norops milleri (H. Smith, 1950)
- Norops monteverde (Koehler, 2009)
- Norops Morazani (Townsend & Wilson, 2009)
- Norops muralla Koehler, McCranie, & Wilson, 1999
- Norops naufragus Campbell, Hillis, & Lamar, 1989
- Norops nebuloides (Bocourt, 1873)
- Norops nebulosus (Wiegmann, 1834)
- Norops notopholis (Boulenger, 1896)
- Norops ocelloscapularis Koehler, McCranie, & Wilson, 2001
- Norops omiltemanus (Davis, 1954)
- Norops onca O'Shaughnessy, 1875
- Norops osa (Köhler, Dehling, & Köhler, 2010)
- Norops oxylophus (Cope, 1875)
- Norops pachypus (Cope, 1875)
- Norpos parvicirculatus (Alvarez del Toro & H. Smith, 1956)
- Brown anole ( Norops petersii (Bocourt, 1873))
- Norops pijolense McCranie, Wilson, & K. Williams, 1993
- Norops pinchoti (Cochran, 1931)
- Norops poecilopus (Cope, 1862)
- Norops polylepis (W. Peters, “1873”, 1874)
- Norops polyrhachis (H. Smith, 1968)
- Norops pseudokemptoni (Köhler, Ponce, Sunyer, & Batista, 2007)
- Norops pseudopachypus (Köhler, Ponce, Sunyer, & Batista, 2007)
- Norops purpurgularis McCranie, Cruz, & Holm, 1993
- Norops pygmaeus (Alvarez del Toro & H. Smith, 1956)
- Norops quaggulus (Cope, 1885)
- Norops quercorum (Fitch, 1978)
- Norops rivalis (Williams, 1984)
- Norops roatanensis Koehler & McCranie, 2001
- Norops rodriguezii (Bocourt, 1873)
- Norops rubribarbaris Koehler, McCranie, & Wilson, 1999
- Norops scapularis (Boulenger, 1908)
- Norops schiedii (Wiegmann, 1834)
- Norops schmidti (Smith, 1939)
- Norops scypheus (Cope, 1864)
- Norops sericeus (Hallowell, 1856)
- Norops serranoi Koehler, 1999
- Norops Simmonsi (Holman, 1964)
- Norops sminthus (Dunn & Emlen, 1932)
- Norops subocularis (Davis, 1954)
- Norops tandai (Avila-Pires, 1995)
- Norops taylori (H. Smith & Players, 1945)
- Norops tenorioensis (Koehler, 2011)
- Norops tolimensis (Werner, 1916)
- Norops townsendi (Stejneger, 1900)
- Norops trachyderma (Cope, 1875)
- Norops tropidogaster (Hallowell, 1856)
- Norops tropidolepis (Boulenger, 1885)
- Norops tropidonotus (Peters, 1863)
- Norops uniformis (Cope, “1884”, 1885)
- Norops unilobatus (Köhler & Vesely, 2010)
- Norops utowanae (Barbour, 1932)
- Norops vicarius (Williams, 1986)
- Norops villai (Fitch & Henderson, 1976)
- Norops wampuensis McCranie & Koehler, 2001
- Norops wellbornae (Ahl, 1940)
- Norops wermuthi Köhler & Obermeier, 1998
- Norops williamsi (Bocourt, 1870)
- Norops woodi (Dunn, 1940)
- Norops yoroensis McCranie, Nicholson, & Koehler, 2001
- Norops zeus Koehler & McCranie, 2001
Individual evidence
- ^ Gunther Köhler: A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae). Zootaxa, 2354, pp. 1-18, 2010
- ↑ Kirsten E. Nicholson & Gunther Köhler: A new species of the genus Norops from Darién, Panama, with comments on N. sulcifrons (Cope 1899) (Reptilia, Squamata, Dactyloidae) . Zootaxa, 3895, 2, pp. 225-237, 2014 doi : 10.11646 / zootaxa.3895.2.5
literature
- Kirsten E. Nicholson, Brian I. Crother, Craig Guyer, Jay M. Savage : It is time for a new classification of anoles (Squamata: Dactyloidae) . Zootaxa , 3477, Magnolia Press, Auckland 2012, ISBN 978-1-77557-010-3 , digital version (PDF; 8.24 MB) .