Casuarina family

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Casuarina family
Horsetail-leaved casuarines (Casuarina equisetifolia), branches with inflorescences.

Horsetail-leaved casuarines ( Casuarina equisetifolia ), branches with inflorescences.

Nuclear eudicotyledons
Eurosiden I
Order : Beech-like (Fagales)
Family : Casuarina family
Scientific name

Casuarina plants (Casuarinaceae) are a family of plants from the order of the beech-like (Fagales). This family includes four genera with a total of about 90 to 95 species. Previously, all species belonging to this family had been assigned to the genus Casuarina ( Casuarina ).

The plant species of this family are common in the tropics of the Old World from the Malay Archipelago to India , Australia, and Pacific islands. Many of the species do not have an established German name and are uniformly referred to as kangaroo trees, casuarinas or club trees.


Allocasuarina littoralis , illustration: 2) male inflorescence, 3) branches with male and female inflorescences, 4) infructescence with open fruits, 5) seed.

All species are evergreen trees or shrubs . They look very similar to some conifers, but are flowering plants . Many species have rod-like small branches that act as leaves. The real leaves, on the other hand, are reduced to small triangular scales. The scale-shaped leaves are four to 16 in whorls . There are no stipules.

The plant species are single- sexed ( monoecious ) or dioecious ( dioecious ). The inflorescences often look like small cones . The unisexual flowers are greatly reduced. Bracts are missing. The male flowers consist of only one stamen . In the female flowers are two carpels to a top permanent ovary grown; the style ends in two scars. Pollination takes place by the wind ( anemophilia ).

There are walnut fruits formed, they are usually winged (Samara).


The roots have a symbiosis with the nitrogen-fixing (N-fixing) Frankia .


Since LAS Johnson divided the genus Casuarina in 1980 , there have been four genera in the family with around 95 species:

  • Allocasuarina L.ASJohnson : The approximately 61 species are common in Australia.
  • Ceuthostoma L.ASJohnson : Of the only two species, one occurs in the Philippines and the other in northern Borneo and western New Guinea .
  • Gymnostoma L.ASJohnson (Syn .: Quadrangula Baum.-Bod. ): The approximately 14 species are distributed from Malesia to islands in the southwestern Pacific.
  • Casuarina ( Casuarina L. ): The 14 species are distributed in Australia and on the islands to the north of it.


Individual evidence

  1. Jump up ↑ LAS Johnson: Notes on Casuarinaceae , In: Telopea , Volume 2 (1), 1980, pp. 83-84
  2. Casuarinaceae in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
  3. a b c d Rafaël Govaerts (ed.): Casuarinaceae. In: World Checklist of Selected Plant Families (WCSP) - The Board of Trustees of the Royal Botanic Gardens, Kew . Retrieved January 13, 2015.

Web links

Commons : Casuarinaceae  - Collection of images, videos and audio files