Vertigo (genus)
Vertigo | ||||||||||||
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Left-handed diaper snail ( Vertigo pusilla ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Vertigo | ||||||||||||
OV Müller , 1773 |
The genus Vertigo is a species-rich genus of the family of diaper snails (Vertiginidae) from the suborder of land snails (Stylommatophora).
features
The representatives of the genus Vertigo have right and left wound, cylindrical, oblong-cylindrical to egg-shaped cases with 5 to 8 turns. The housings are up to 4 mm high and 2.3 mm wide. They are mostly smooth, rarely ribbed and mostly dark in color. The whorls are convex or almost flattened. The outer edge of the mouth is often flattened or slightly curved inward. The mouth edge is simply with a sharp edge, or thickened lipped and / or bent outwards. The mouth can be reinforced with up to seven teeth, but it can also be unreinforced.
In the genital system, the penis is comparatively long with an epiphallus. The penile appendix is missing, the penile retractor muscle consists of one muscle cord. It starts at the epiphallus. The spermatic duct is long and the vagina is comparatively long. The stem of the seminal vesicle is comparatively very long and several times longer than the elongated, egg-shaped seminal vesicle itself.
Geographical distribution
The species of the genus occur in the Holarctic .
Taxonomy
The taxon was drawn up in 1773 by Otto Friedrich Müller . The type is Vertigo pusilla Müller, 1774 by subsequent monotype . Currently over 70 species are included in the genus Vertigo. However, the scope is quite uncertain as there is no comprehensive revision of the genre. The vertigo species of North America accepted as valid taxa by Jeffrey Nekola and Brian Jones in 2010 are marked by individual records .
- Vertigo alabamensis Clapp, 1915 ( syn.Vertigo conecuhensis Clapp, 1915)
- Vertigo allyniana Berry, 1919
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Alpine diaper snail ( Vertigo alpestris Alder, 1838) with the subspecies
- Vertigo alpestris alpestris
- Vertigo alpestris uturyotoensis Kuroda & Fukuda, 1944
- Vertigo andrusiana Pilsbry, 1899
- † Vertigo angulifera Boettger, 1884, Badenian , Sarmatian ( Miocene )
- Narrow diaper snail ( Vertigo angustior (Jeffreys, 1830))
- Marsh diaper snail ( Vertigo antivertigo (Draparnaud, 1801))
- Vertigo arctica (Wallenberg, 1858)
- Vertigo arizonensis (Pilsbry & Vanatta, 1900) (= Vertigo gouldi arizonensis Pilsbry & Vanatta, 1900)
- Vertigo arthuri by Martens, 1882 ( syn.Vertigo gouldi basidens Pilsbry & Vanatta, 1900)
- Vertigo berryi Pilsbry, 1919
- † Vertigo bifida (Deshayes, 1863), Chattium
- Vertigo binneyana Sterki, 1890
- † Vertigo bipalatata Schütt, 1994
- Vertigo bollesiana (Morse, 1865)
- Vertigo botanicorum Horsak & Pokryszko, 2010
- † Vertigo brusinai Stefani, 1880, Pliocene
- Vertigo californica Rowell, 1861
- Vertigo callosa (Reuss, 1852) (with numerous subspecies and varieties), Middle to Upper Miocene
- † Vertigo capellinii Sacco, 1886, Pliocene
- Vertigo clappi Brooks & Hunt, 1936
- Vertigo columbiana Pilsbry & Vanatta, 1900
- Vertigo concinnula Cockerell, 1897
- Vertigo cristata Sterki, 1919
- Vertigo dalliana Sterki, 1890
- † Vertigo defrancei (Brogniart, 1810), Chattium
- Vertigo elatior Sterki, 1894
- † Vertigo elsheimensis Boettger, 1889, Rupelium
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Vertigo eogia Pilsbry, 1919 with the subspecies:
- Vertigo eogia eogia
- Vertigo eogia stagnalis (Kuroda, 1941)
- Vertigo extima (Westerlund, 1877)
- † Vertigo flexidens (Reuss, 1860), Aquitanium
- Bare diaper snail ( Vertigo genesii (Gredler, 1856))
- Four-toothed diaper snail ( Vertigo geyeri Lindholm, 1925)
- † Vertigo globosa Sacco, 1886, Pliocene
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Vertigo gouldii (Binney, 1843) with the subspecies:
- Vertigo gouldii gouldii ,
- Vertigo gouldi coloradoensis (Cockerell, 1891)
- Vertigo gouldi inserta (Pilsbry, 1919)
- Vertigo hannai (Pilsbry, 1919)
- † Vertigo hauchecornei Klebs, 1886, Eocene
- Vertigo hebardi Vanatta, 1912
- Slender diaper snail ( Vertigo heldi (Clessin, 1877))
- Vertigo hinkleyi Pilsbry, 1921
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Vertigo hirasei Pilsbry, 1901 with the subspecies:
- Vertigo hirasei hirasei
- Vertigo hirasei glans Pilsbry & Hirase , 1904
- Vertigo hubrichti (Pilsbry, 1934) ( syn.Vertigo brierensis Leonard, 1972, Vertigo hubrichti variabilis Frest, 1991, Vertigo iowaensis Frest, 1991)
- Vertigo hydrophila Reinhardt, 1877
- Vertigo idahoensis Pilsbry, 1934
- Vertigo japonica Pilsbry & Hirase, 1904
- † Vertigo kochi Boettger, 1889, Chattium
- † Vertigo kuenowii Klebs, 1886, Eocene
- Vertigo kushiroensis Pilsbry & Hirase, 1905
- Moor diaper snail ( Vertigo lilljeborgi (Westerlund, 1871))
- Vertigo malleata Coles & Nekola, 2007
- † Vertigo maresi Bourguignat, 1862, Pliocene
- † Vertigo marki Gulick, 1904, Pliocene
- † Vertigo maxillosa Schütt, 1994
- Vertigo meramecensis Van Hasselt, 1835
- Vertigo microsphaera Schileyko, 1984
- Vertigo milium (Gould, 1840)
- † Vertigo minor Boettger, 1870, Burdigalium
- Vertigo miyakejimana Pilsbry & Hirase, 1919
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Arctic diaper snail ( Vertigo modesta (Say, 1824)) with the subspecies:
- Vertigo modesta modesta
- Vertigo modesta castanea Brooks & Brooks, 1940
- Vertigo modesta corpulenta Morse, 1865
- Vertigo modesta hoppii (Möller, 1842)
- Vertigo modesta ingersolli Cockerell, 1891
- Vertigo modesta ultima (Pilsbry, 1948)
- † Vertigo moeniana Zinndorf, 1901 Chattium
- Vertigo morsei Sterki, 1894
- Bellied diaper snail ( Vertigo moulinsiana (Dupuy, 1849))
- † Vertigo myrmido Michaud, 1855, Pliocene
- Vertigo nangaparbatensis Pokryszko & Hlaváč, 2009
- Vertigo neglecta Arango at Poey, 1856
- Vertigo nitens Pease, 1861
- Vertigo nitidula (Mousson, 1876) ( syn.Vertigo sieversi Boettger, 1879)
- † Vertigo nouleti Michaud, 1862, Pliocene
- † Vertigo numellata Gulick, 1904
- Vertigo nylanderi Sterki, 1909
- Vertigo occidentalis Sterki, 1907
- Vertigo okinoerabuensis Pilsbry & Hirase, 1904
- Vertigo oralis Sterki, 1898
- Vertigo oscariana Sterki, 1890
- Vertigo oughtoni Pilsbry, 1948
- Vertigo ovata (Say, 1822)
- † Vertigo ovatula (Sandberger, 1874) (with several subspecies), Chattium
- Vertigo paradoxa Sterki in Pilsbry, 1900
- Vertigo parcedentata (Al. Braun, 1847)
- Vertigo parvula Sterki, 1890
- Vertigo perryi Sterki, 1905
- † Vertigo protracta (Sandberger, 1874), Chattium
- † Vertigo pseudoantivertigo Paladilhe, 1873, Pliocene
- Vertigo pseudosubstriata Ložek, 1954
- Left-handed diaper snail ( Vertigo pusilla (OF Müller, 1774)), recent and? Pliocene
- Common diaper snail ( Vertigo pygmaea (Draparnaud, 1801)), recent and? Pliocene
- Vertigo rowelli (Newcomb, 1860)
- Vertigo rugosula Sterki, 1890 ( syn.Vertigo wheeleri Pilsbry, 1928)
- Nordic diaper snail ( Vertigo ronnebyensis (Westerlund, 1871))
- † Vertigo sandbergeri Wenz, 1923, Tortonium
- Vertigo sterkii Pilsbry, 1919
- Striped diaper snail ( Vertigo substriata (Jeffreys, 1833))
- Vertigo superstriata Pokryszko & Auffenberg, 2009
- Vertigo teskeyae Hubricht, 1961
- Vertigo torrei Aguayo & Jaume, 1934
- Vertigo tosana Pilsbry, 1919
- Vertigo tridentata Wolf, 1870
- † Vertigo tuchoricensis Pilsbry. 1919
- Vertigo ultimathule by Proschwitz, 2007
- Vertigo ventricosa (ES Morse, 1865)
Vertigo shimochii Kuroda & Amano, 1960 was synonymous with Gastrocopta servilis (Gould, 1843) in 2012 .
Schileyko (1998) divides the genus into eight sub-genera:
- Vertigo (Vertigo) Müller, 1773
- Vertigo (Alloptyx) Pilsbry, 1953
- Vertigo (Augustula) Sterki, 1888
- Vertigo (Isthmia) Gray, 1821
- Vertigo (Nearctula) Sterki, 1892
- Vertigo (Ptychalaea) O. Boettger, 1889
- Vertigo (Staurodon) Lowe, 1852
- Vertigo (Vertillaria) Pilsbry, 1920
This subdivision has hardly found any resonance. Some sub-genera were also understood as independent genera. Other authors also rate Vertilla Moquin-Tandon, 1856, as a subgenus of Vertigo .
supporting documents
literature
- Beata M. Pokryszko: The Vertiginidae of Poland (Gastropoda: Pulmonata: Pupilloidea) - a systematic monograph. Annales Zoologici, 43 (8): 133-257, Warsaw 1990.
- Anatolij A. Schileyko: Treatise on Recent Terrestrial Pulmonate Molluscs. Part 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica, Supplement 2 (2): 129-261, Moscow 1998 ISSN 0136-0027
- Francisco W. Welter-Schultes: European non-marine molluscs, a guide for species identification = identification book for European land and freshwater mollusks. A1-A3 S., 679 S., Q1-Q78 S., Göttingen, Planet Poster Ed., 2012 ISBN 3-933922-75-5 , ISBN 978-3-933922-75-5
- Wilhelm Wenz: Gastropoda extramarina tertiaria. In: Carl Diener (Ed.), Fossilium catalogus, 1 Animalium , 20: 737-1068, Berlin 1923 Online at www.biodiversitylibrary.org (p. 981ff.)
Individual evidence
- ↑ Otto Friedrich Müller: Vermivm terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. Voluminis Imi pars Ima. S. (1-33), pp. 1–135, Havniæ / Copenhagen & Lipsiæ / Leipzig, Heineck & Faber, 1773.
- ↑ Michal Horsák & Beata M. Pokryszko: Vertigo botanicorum sp. nov. (Gastropoda: Pulmonata: Vertiginidae) - a new whorl-snail from the Russian Altai Mountains. Zootaxa, 2634: 57-62, 2010 PDF ( page no longer available , search in web archives ) Info: The link was automatically marked as defective. Please check the link according to the instructions and then remove this notice.
- ↑ World Mollusc Species Data Base by Bagni Liggia
- ^ Integrated Taxonomic Information System - Genus Vertigo
- ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad Jeffrey C. Nekola and Brian F. Coles: Pupillid Land Snails of Eastern North America. American Malacological Bulletin, 28 (2): 29-57, 2010 doi : 10.4003 / 006.028.0221
- ^ A b Mathias Harzhauser, Martin Gross, Herbert Binder: Biostratigraphy of Middle Miocene (Sarmatian) wetland systems in an Eastern Alpine intramontane basin (Gratkorn Basin, Austria): the terrestrial gastropod approach. Geologica Carpathica, 59 (1): 45-58, 2008
- ↑ a b H. Schütt: Neogene mollusks from the tertiary basin near Afyon, Turkey. Geologica et Palaeontologica, 28: 167-183, Marburg 1994 ISSN 0072-1018
- ↑ a b Beata M. Pokryszko, Kurt Auffenberg, Jaroslav Č. Hlaváč, Fred Naggs: Pupilloidea of Pakistan (Gastropoda: Pulmonata): Truncatellininae, Vertigininae, Gastrocoptinae, Pupillinae (In Part). Annales Zoologici, 59 (4): 423-458, 2009 doi : 10.3161 / 000345409X484847 .
- ^ Frank Walther, Pavel V. Kijashko, Bernhard Hausdorf: Redescription of Vertigo (Vertigo) nitidula (Mousson, 1876) (Gastropoda: Vertiginidae) from the Caucasus region. Zootaxa, 3872, 1, pages 75-82, October 2014
- ↑ Jeffrey C. Nekola, Alex Jones, Gabe Martinez, Sarah Martinez, Khayman Mondragon, Terrence Lebeck, John Slapcinsky, Satoshi China: Vertigo shimochii Kuroda & Amano 1960 synonymized with Gastrocopta servilis (Gould, 1843) based on conchological and DNA sequence data. Zootaxa, 3161: 48-52, 2012 PDF
- ^ Molluscs of Central Europe