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{{Short description|Extinct genus of dinosaurs}}
{{Speciesbox
{{Speciesbox
| fossil_range = [[Early Cretaceous]], {{fossilrange|112}}
| fossil_range = [[Early Cretaceous]], {{fossilrange|112}}
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'''''Theiophytalia''''' is a [[genus]] of [[herbivorous]] [[iguanodont]]ian [[dinosaur]] from the lower [[Cretaceous]] [[period (geology)|period]] ([[Aptian]]-[[Albian]] stage, about 112 million years ago<ref name="iguanodonts2010carpenter">{{Cite journal|author1=Carpenter, K. |author2=Ishida, Y. |name-list-style=amp |year=2010 |url=http://www.ucm.es/info/estratig/JIG/vol_content/vol_36_2/36_2_145_164_Carpenter.pdf |title=Early and "Middle" Cretaceous Iguanodonts in Time and Space |journal=[[Journal of Iberian Geology]] |volume=36 |issue=2 |pages=145–164 |doi=10.5209/rev_JIGE.2010.v36.n2.3}}</ref>) of [[Colorado]], USA. It contains a single species, '''''T. kerri'''''.<ref name=BrillCarpenter>{{cite book |author1=Brill, K. |author2=K. Carpenter |name-list-style=amp |year=2006 |chapter=A Description of a New Ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a Reassessment of the Skull of ''Camptosaurus'' |editor=Carpenter, Kenneth |title=Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs |publisher=Indiana University Press |location=Bloomington |pages=49–67 }}</ref>
'''''Theiophytalia''''' is a [[genus]] of [[herbivorous]] [[iguanodont]]ian [[dinosaur]] from the lower [[Cretaceous]] [[period (geology)|period]] ([[Aptian]]-[[Albian]] stage, about 112 million years ago<ref name="iguanodonts2010carpenter">{{Cite journal|author1=Carpenter, K. |author2=Ishida, Y. |name-list-style=amp |year=2010 |url=http://www.ucm.es/info/estratig/JIG/vol_content/vol_36_2/36_2_145_164_Carpenter.pdf |title=Early and "Middle" Cretaceous Iguanodonts in Time and Space |journal=[[Journal of Iberian Geology]] |volume=36 |issue=2 |pages=145–164 |doi=10.5209/rev_JIGE.2010.v36.n2.3|doi-access=free }}</ref>) of [[Colorado]], USA. It contains a single species, '''''T. kerri'''''.<ref name=BrillCarpenter>{{cite book |author1=Brill, K. |author2=K. Carpenter |name-list-style=amp |year=2006 |chapter=A Description of a New Ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a Reassessment of the Skull of ''Camptosaurus'' |editor=Carpenter, Kenneth |title=Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs |publisher=Indiana University Press |location=Bloomington |pages=49–67 }}</ref>


==Description==
==Description==
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==Discovery==
==Discovery==
[[File:Marsh Camptosaurus.jpg|thumb|Historical skeletal restoration of ''Camptosaurus'' by O.C. Marsh, with skull based on remains now referred to ''Theiophytalia''.]]
[[File:Marsh Camptosaurus.jpg|thumb|Historical skeletal restoration of ''Camptosaurus'' by O.C. Marsh, with skull based on remains now referred to ''Theiophytalia''.]]
The [[holotype]] and only known specimen, YPM 1887, is a partial [[skull]] that was referred by [[Othniel Charles Marsh|O.C. Marsh]] (affirmed by [[Charles W. Gilmore|Gilmore]], 1909), to whom the skull was given in 1886, as that of ''[[Camptosaurus]] amplus''. Gilmore used the skull to reconstruct the skull of ''Camptosaurus'' assuming that it came from the [[Morrison Formation]].<ref name=Gilmore09>{{Cite journal|author=Gilmore, C.W. |year=1909 |title=Osteology of the Jurassic reptile ''Camptosaurus'', with a revision of the species of the genus, and descriptions of two new species |journal=Proceedings of the United States National Museum |volume=36 |issue=1666 |pages=197–332 |doi=10.5479/si.00963801.36-1666.197}}</ref> However, microscopic comparisons of thin-sections of the Mesozoic formations in the [[Garden of the Gods]] showed that the specimen actually came from the Lytle Member of the [[Purgatoire Formation]]; therefore, the skull was Early Cretaceous in age.
In 1878, a student of [[Colorado College]] named James Kerr found a partial ornithopod skull at [[Garden of the Gods]] park in [[El Paso County, Colorado|El Paso County]], [[Colorado]].<ref name="BrillCarpenter" /> The skull, YPM 1887, was referred by [[Othniel Charles Marsh|O.C. Marsh]] (affirmed by [[Charles W. Gilmore|Gilmore]], 1909), to whom the skull was given in 1886, as that of ''[[Camptosaurus]] amplus''. Gilmore used the skull to reconstruct the skull of ''Camptosaurus'' assuming that it came from the [[Morrison Formation]].<ref name=Gilmore09>{{Cite journal|author=Gilmore, C.W. |year=1909 |title=Osteology of the Jurassic reptile ''Camptosaurus'', with a revision of the species of the genus, and descriptions of two new species |journal=Proceedings of the United States National Museum |volume=36 |issue=1666 |pages=197–332 |doi=10.5479/si.00963801.36-1666.197|url=https://www.biodiversitylibrary.org/part/52542 }}</ref> However, microscopic comparisons of thin-sections of the Mesozoic formations in the [[Garden of the Gods]] showed that the specimen actually came from the Lytle Member of the [[Purgatoire Formation]]; therefore, the skull was Early Cretaceous in age.


The [[name of a biological genus|generic name]] is, from Greek, θειος, ''theios'': "divine" + Greek φυταλία, ''phytalia'': "garden", or "garden of the gods". [[Garden of the Gods]] is a park in [[Colorado Springs, Colorado|Colorado Springs]], Colorado, where a skull, the only fossil of the genus yet to be discovered, was found in 1878. The [[specific name (zoology)|specific name]] ''kerri'' honors James Hutchinson Kerr, who found the specimen.<ref name=BrillCarpenter/>
The [[name of a biological genus|generic name]] is, from Greek, θειος, ''theios'': "divine" + Greek φυταλία, ''phytalia'': "garden", or "garden of the gods". [[Garden of the Gods]] is a park in [[Colorado Springs, Colorado|Colorado Springs]], Colorado, where a skull, the only fossil of the genus yet to be discovered, was found in 1878. The [[specific name (zoology)|specific name]] ''kerri'' honors James Hutchinson Kerr, who found the specimen.<ref name=BrillCarpenter/>


==Classification==
==Classification==
The article describing the find classified ''Theiophytalia'' as intermediate in derivation between ''Camptosaurus'' and ''[[Iguanodon]]''. The [[type species]] is ''Theiophytalia kerri''.<ref name=BrillCarpenter/> In 2010 and 2011 [[cladistic]] analyses of McDonald and colleagues, ''Theiophytalia'' has been recovered as a basal member of the [[Styracosterna]] and its closest relative was ''[[Hippodraco]]''.<ref name=hippoiguana>{{Cite journal|author=McDonald, A.T.|author2=Kirkland, J.I.|author3=DeBlieux, D.D.|author4=Madsen, S.K.|author5=Cavin, J.|author6=Milner, A.R.C.|author7=Panzarin, L.|name-list-style=amp |year=2010 |title=New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs |journal=PLOS ONE |volume=5 |issue=11 |pages=e14075 |doi=10.1371/journal.pone.0014075 |pmid=21124919 |pmc=2989904}}</ref><ref name="McDonald">{{Cite journal|author=Andrew T. McDonald |year=2011 |title=The taxonomy of species assigned to ''Camptosaurus'' (Dinosauria: Ornithopoda) |url=http://www.mapress.com/zootaxa/2011/f/z02783p068f.pdf |journal=Zootaxa |volume=2783 |pages=52–68 |doi= 10.11646/zootaxa.2783.1.4}}</ref>
The article describing the find classified ''Theiophytalia'' as intermediate in derivation between ''Camptosaurus'' and ''[[Iguanodon]]''. The [[type species]] is ''Theiophytalia kerri''.<ref name=BrillCarpenter/> In 2010 and 2011 [[cladistic]] analyses of McDonald and colleagues, ''Theiophytalia'' has been recovered as a basal member of the [[Styracosterna]] and its closest relative was ''[[Hippodraco]]''.<ref name=hippoiguana>{{Cite journal|author=McDonald, A.T.|author2=Kirkland, J.I.|author3=DeBlieux, D.D.|author4=Madsen, S.K.|author5=Cavin, J.|author6=Milner, A.R.C.|author7=Panzarin, L.|name-list-style=amp |year=2010 |title=New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs |journal=PLOS ONE |volume=5 |issue=11 |pages=e14075 |doi=10.1371/journal.pone.0014075 |pmid=21124919 |pmc=2989904|bibcode=2010PLoSO...514075M|doi-access=free}}</ref><ref name="McDonald">{{Cite journal|author=Andrew T. McDonald |year=2011 |title=The taxonomy of species assigned to ''Camptosaurus'' (Dinosauria: Ornithopoda) |url=http://www.mapress.com/zootaxa/2011/f/z02783p068f.pdf |journal=Zootaxa |volume=2783 |pages=52–68 |doi= 10.11646/zootaxa.2783.1.4}}</ref>


{{clade| style=font-size:90%; line-height:90%
{{clade| style=font-size:90%; line-height:90%
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[[Category:Taxa named by Kenneth Carpenter]]
[[Category:Taxa named by Kenneth Carpenter]]
[[Category:Paleontology in Colorado]]
[[Category:Paleontology in Colorado]]
[[Category:Ornithischian genera]]

Latest revision as of 17:00, 13 January 2023

Theiophytalia
Temporal range: Early Cretaceous, 112 Ma
Holotype skull, white parts are reconstructed
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Ornithopoda
Clade: Styracosterna
Genus: Theiophytalia
Brill & Carpenter, 2006
Species:
T. kerri
Binomial name
Theiophytalia kerri
Brill & Carpenter, 2006

Theiophytalia is a genus of herbivorous iguanodontian dinosaur from the lower Cretaceous period (Aptian-Albian stage, about 112 million years ago[1]) of Colorado, USA. It contains a single species, T. kerri.[2]

Description[edit]

Reconstructed Theiophytalia skull mounted on a Camptosaurus skeleton cast, Museo Nacional de Ciencias Naturales, Madrid

Detailed comparisons by Brill and Carpenter (2006) also showed that the skull differed in a number of key features from that of Camptosaurus, namely: a longer, heavier, and more rugose snout; a wider dorsal process on the maxilla; a proportionally smaller antorbital fenestra; and stouter quadrate, with a bulbous articulation for the lower jaw. Compare the skull image with that of Camptosaurus. Therefore, they put it into its own genus and species.[2]

Discovery[edit]

Historical skeletal restoration of Camptosaurus by O.C. Marsh, with skull based on remains now referred to Theiophytalia.

In 1878, a student of Colorado College named James Kerr found a partial ornithopod skull at Garden of the Gods park in El Paso County, Colorado.[2] The skull, YPM 1887, was referred by O.C. Marsh (affirmed by Gilmore, 1909), to whom the skull was given in 1886, as that of Camptosaurus amplus. Gilmore used the skull to reconstruct the skull of Camptosaurus assuming that it came from the Morrison Formation.[3] However, microscopic comparisons of thin-sections of the Mesozoic formations in the Garden of the Gods showed that the specimen actually came from the Lytle Member of the Purgatoire Formation; therefore, the skull was Early Cretaceous in age.

The generic name is, from Greek, θειος, theios: "divine" + Greek φυταλία, phytalia: "garden", or "garden of the gods". Garden of the Gods is a park in Colorado Springs, Colorado, where a skull, the only fossil of the genus yet to be discovered, was found in 1878. The specific name kerri honors James Hutchinson Kerr, who found the specimen.[2]

Classification[edit]

The article describing the find classified Theiophytalia as intermediate in derivation between Camptosaurus and Iguanodon. The type species is Theiophytalia kerri.[2] In 2010 and 2011 cladistic analyses of McDonald and colleagues, Theiophytalia has been recovered as a basal member of the Styracosterna and its closest relative was Hippodraco.[4][5]

Ankylopollexia

References[edit]

  1. ^ Carpenter, K. & Ishida, Y. (2010). "Early and "Middle" Cretaceous Iguanodonts in Time and Space" (PDF). Journal of Iberian Geology. 36 (2): 145–164. doi:10.5209/rev_JIGE.2010.v36.n2.3.
  2. ^ a b c d e Brill, K. & K. Carpenter (2006). "A Description of a New Ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a Reassessment of the Skull of Camptosaurus". In Carpenter, Kenneth (ed.). Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Bloomington: Indiana University Press. pp. 49–67.
  3. ^ Gilmore, C.W. (1909). "Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species". Proceedings of the United States National Museum. 36 (1666): 197–332. doi:10.5479/si.00963801.36-1666.197.
  4. ^ McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C. & Panzarin, L. (2010). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLOS ONE. 5 (11): e14075. Bibcode:2010PLoSO...514075M. doi:10.1371/journal.pone.0014075. PMC 2989904. PMID 21124919.
  5. ^ Andrew T. McDonald (2011). "The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" (PDF). Zootaxa. 2783: 52–68. doi:10.11646/zootaxa.2783.1.4.

External links[edit]