Ptilostemon gnaphaloides

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Ptilostemon gnaphaloides
Systematics
Family : Daisy family (Asteraceae)
Subfamily : Carduoideae
Tribe : Cynareae
Genre : Ivory Thistles ( Ptilostemon )
Type : Ptilostemon gnaphaloides
Scientific name
Ptilostemon gnaphaloides
( Cirillo ) Soják

Ptilostemon gnaphaloides is a plant from the family of the daisy family (Asteraceae). It grows on rocky locations in the central and eastern Mediterranean.

description

Ptilostemon gnaphaloides is an unreinforced, small shrub with a rounded outline and heights between 50 and 120 centimeters. The linear, 10 to 15 cm long, usually only 1 to 2 millimeter wide leaves have entire margins and slightly rolled up at the edge and gradually taper off into an awl. The leaves on the flowering shoots are slightly widened, prickly and shorter than the rest of the leaves. On both sides they usually have one or two stipule-like, narrow fringes with thorns on the base, which in the subspecies gnaphaloides are 0.3 to 2, rarely up to 4 millimeters, in the subspecies pseudofruticosus 1 to 5, rarely up to 9 millimeters long.

The flowering shoots reach lengths of 30 to 60 centimeters. Two to ten flower heads are arranged in an irregular umbrella shape. The fluffy and felty basket shell has at the subspecies gnaphaloides a diameter 16 to 22 millimeters, wherein the subspecies pseudofruticosus 18 to 26 millimeters. The mean husks have a slender, upright-projecting appendage, whose length in the subspecies gnaphaloides 2 to 4 millimeters, at the subspecies pseudofruticosus to 8 millimeters and 4 only in this subspecies in a 1 to 3 mm long mandrel expires. The 18 to 23 millimeter long flowers are purple, pink or whitish in color. The appendages of the anthers are evenly banded. The cardboard hairs of the inner flowers have bearded tips.

Ptilostemon gnaphaloides subsp. gnaphaloides flowers from April to June, the subspecies pseudofruticosus from May to June.

Occurrence

This species has a very fragmented range in the central and eastern Mediterranean. Both subspecies grow in crevices of vertical limestone cliffs, more rarely in less steep locations on dry gravel or in dry Mediterranean scrubland at altitudes from sea level to 750 m.

Systematics

There are two subspecies:

According to the classic monograph by Werner Greuter , Ptilostemon gnaphaloides forms together with Ptilostemon chamaepeuce the section Ptilostemon in the subgenus Ptilostemon . A third species, Ptilostemon greuteri , newly described in 2006 and only populating a tiny area in the mountains near Trapani (Sicily), was initially connected here due to the shrubby growth common to all three species. In a phylogenomic study (analysis of the relationships based on the comparison of homologous DNA sequences) the close relationship of Ptilostemon gnaphaloides and Ptilostemon chamaepeuce (as sister species ) was confirmed, but Ptilostemon greuteri was not closely related to them.

supporting documents

  1. ↑ a b c d e f Werner Greuter: Ptilostemon. In: Peter Hadland Davis (Ed.): Flora of Turkey and the East Aegean Islands. Vol. 5 (Compositae) . Edinburgh University Press, Edinburgh 1975, ISBN 0-85224-280-8 , pp. 415-419 .
  2. ↑ a b c d e f g h Werner Greuter: Monograph of the genus Ptilostemon (Compositae). In: Boissiera. Volume 22, 1973, pp. 1-215.
  3. ↑ a b Klaus Werner: 121. Ptilostemon. In: TG Tutin, VH Heywood, NA Burges, DM Moore, DH Valentine, SM Walters, DA Webb (eds.): Flora Europaea . Volume 4: Plantaginaceae to Compositae (and Rubiaceae) . Cambridge University Press, Cambridge 1976, ISBN 0-521-08717-1 , pp. 242 (English, limited preview in Google Book search).
  4. ↑ Ralf Jahn, Peter Schönfelder: Excursion flora for Crete . With contributions by Alfred Mayer and Martin Scheuerer. Eugen Ulmer, Stuttgart (Hohenheim) 1995, ISBN 3-8001-3478-0 , p. 323 .
  5. ^ A b Werner Greuter: Compositae (pro parte majore). In: Werner Greuter, Eckhard von Raab-Straube (ed.): Compositae. Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity. Ptilostemon gnaphaloides. Berlin 2006+.
  6. ↑ Tarek A. Mukassabi, Gousn Ahmidat, Imhamed M. Sherif, Abdusslam Elmogasapi, Peter A. Thomas: Checklist and lifeforms of plant species in contrasting climatic zones of Libya. In: Biological Diversity and Conservation. Volume 5, No. 3, 2012, pp. 1-12 [( http://www.biodicon.com/YayinlananMakaleler/1.%20250-0612.%20Peter.pdf PDF file)].
  7. ↑ Fabio Conti, Giovanna Abbate, Alessandro Alessandrini, Carlo Blasi (Eds.): An annotated checklist of the Italian vascular flora. Palombi, Roma 2005, ISBN 88-7621-458-5 , p. 149 (PDF file; 9 MB) .
  8. ^ Salvatore Brullo, Anna Guglielmo, Pietro Pavone, Maria Carmen Terrasi: Numeri cromosomici per la flora italiana: 1251-1266. In: Informatore Botanico Italiano. Volume 23, No. 1, 1991 (publ. 1992), pp. 39-47 (quoted from: Gianni Bedini, Fabio Garbari, Lorenzo Peruzzi (eds.): Chrobase.it - ​​Chromosome numbers for the Italian flora. ( Memento des Originals from January 6, 2013 in the Internet Archive ) Info: The @1@ 2Template: Webachiv / IABot / www.biologia.unipi.it archive link has been inserted automatically and has not yet been checked. Please check the original and archive link according to the instructions and then remove this note. Pisa 2010+, accessed on June 21, 2016; Origin of the plants : Calabria).
  9. ↑ Neriman Özhatay, Şükran Kültür, Serdar Aalan: Check-list of Additional Taxa to the Supplement Flora of Turkey IV. In: Turkish Journal of Botany. Volume 33, No. 3, 2009, pp. 191-226, DOI: 10.3906 / bot-0805-12 .
  10. ↑ Francesco M. Raimondo, Gianni Antonio Domination: Ptilostemon greuteri (Compositae), a new species from Sicily. In: Willdenowia. Volume 36, No. 1, 2006, pp. 169-175, DOI: 10.3372 / wi.36.36114 .
  11. ↑ Roser Vilatersana, Núria Garcia-Jacas, Teresa Garnatje, Julian Molero, Gabriella Sonnante, Alfonso Susanna: Molecular Phylogeny of the Genus Ptilostemon (Compositae: Cardueae) and Its Relationships with Cynara and Lamyropsis. In: Systematic Botany. Volume 35, No. 4, 2010, 907-917, DOI: 10.1600 / 036364410X539952 .

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