Alchemilla sect. Erectae
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| Alchemilla sect. Erectae | ||||||||||||
| SEFröhner |
Alchemilla sect. Erectae is an independent base section from the genus lady's mantle ( Alchemilla ) in the family of Rosaceae (Rosaceae).
features
The lady's mantle species in the erectae section are medium-sized to very large subshrubs . Their vascular bundles are strongly thickened with sclerosis . The longest internodes of the seedlings are up to 4 millimeters long. The main axis is woody, up to 20 millimeters thick and has a secondary growth in thickness for up to 20 years (possibly longer) . Their internodes are 1 to 5 millimeters long.
The cotyledons are elliptical to rounded, narrow slightly in the stem and are long-stalked. The primary leaf is shallow three-lobed with the lobes reaching an incision depth of 10 to 25% of the leaf radius, usually longer than wide and has a rounded to wedge-shaped base. The top of the petioles is flat, so they are two-edged. Their vascular bundles are concentric, with the thickness of the abaxial 1.5 to 3 times the thickness of the adaxial . The leaf base is usually three-veined. The nerves are up to 7 millimeters apart and are rapidly converging. The leaf blade has a width of 5 to 22 centimeters, is flat or slightly wavy, light green to light gray-green in color and dull. The upper side of the leaf has raised nerves (at least when dry), 5 to 25 percent of the radius is incised in (7) 9 to 13 lobes. The width of the end lobes is 27 to 40 (45) degrees and they are 13 to 27 teeth. The length of the teeth is 1.5 to 7% of the radius of the blade and they are 0.3 to 1.5 times as long as they are wide.
The stipules are 15 to 30 millimeters long, which corresponds to 2 to 7% of the stem length. They have entire margins or 2 to 4 lobes, and usually poor in nerves. They quickly dry up into an orange-brownish tunic, only the two youngest pairs are still fresh. The ears are fused. The incision has a depth of 0 to 2 millimeters, which corresponds to 0 to 10%.
The stem is 20 to 100 centimeters long, erect and 2 to 6 millimeters thick. 10 to 20 internodes are formed by the end of flowering. The plant usually takes root from the lowest nodes. On the lowest stem leaf, the stipules are fused together opposite the petiole to form a horizontally opposed, richly toothed tongue and mostly adaxially hairy. On the uppermost stem leaf, the stipules are toothed and lobed with up to 20 different lobes.
The inflorescence is very floriferous, it contains up to 2000 flowers. At the bottom of Monochasien -Wirteln sometimes lacking bracts , within the Monochasien are often some available. 2 flowers are seldom approximated to form shark gold.
The flowers are yellowish, not red in color and rarely have five. The goblet is short top-shaped. Upwards it is widened, downwards it narrows like a stalk and in most species it is hairy protruding in the lower part. The sepals are 1.3 to 2 times as long as they are wide and 1.2 to 2 times as long as the sepals. The outer sepals are 3 to 5 annoying. At terminal flowers they are often longer and wider than sepals. They are often serrated, 1 to 3 times as long as the sepal cup and, like the sepals, remain horizontal. The discus bulge is narrower than the opening. The stamens are not spreading, have a length of 0.4 to 0.8 millimeters and gradually narrow from the 0.2 to 0.3 millimeter wide base. There is a carpel . The scar is lenticular.
The fruits are nuts . These are about 1.5 times as long as they are wide, up to just over 1.5 millimeters long and tower over the discus with 33 to 50% of their length.
Flowering time is from May to July, subalpine locations from June to August. The beginning of flowering is 3 to 4 weeks after the peach. Synflorescence takes about 8 weeks to bloom. There is no autumn bloom. The fruits ripen after at least 8 weeks. There are some winter men.
The chromosomes are roughly egg-shaped.
Occurrence
The distribution area of the section includes the Middle East and especially the Caucasus and Anatolia , 3 species can also be found in Iran . To the northwest it extends into the southern Carpathians and the Balkan Peninsula (only individual species). In the Urals there is also said to be an Alchemilla species of this section, which has to be checked more closely. This section was introduced into Central Europe.
Large-leaved subshrubs occur in tall herbaceous and high grass corridors as well as the fringing communities of the montane to subalpine level (sub-) meridional mountains. The more xerophilically built species, which are usually richly hairy, but often have almost unlapped leaf blades, can be found in the mountain steppes of Anatolia and Iran. Here they can assert themselves because of their deep roots that have lived for decades if they had enough superficial moisture available in some of their youth.
Systematics
The Erectae section is one of the four basic sections of the European alchemils, from which the remaining sections have emerged through hybridization.
The Central European species are:
- Soft lady's mantle ( Alchemilla mollis (Buser) Rothm. )
- Handsome lady's mantle ( Alchemilla speciosa Buser )
supporting documents
- Sigurd Fröhner: Alchemilla . In: Hans. J. Conert et al. a. (Ed.): Gustav Hegi. Illustrated flora of Central Europe. Volume 4 Part 2B: Spermatophyta: Angiospermae: Dicotyledones 2 (3). Rosaceae 2 . Blackwell 1995, ISBN 3-8263-2533-8 , pp. 13-242.