Eschweilera
Eschweilera | ||||||||||||
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Branch of an Eschweilera species (probably Eschweilera ovata (Cambess.) Miers ), Ajuruteua Peninsula, Bragança , Pará , Brazil |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Eschweilera | ||||||||||||
Mart. ex DC. |
Eschweilera is a genus within the potted fruit tree family(Lecythidaceae). The 90 or so species are common in tropical South and Central America. Several species are used as timber , and many are endangered.
Description and ecology
In the genus Eschweilera there are trees of different sizes that grow accordingly on different levels of the forest: in the undergrowth, in the canopy or as emergent. The alternate leaves are not clustered at the ends of the branches, leaf sizes between 5 and 60 centimeters occur in the genus. The leaves are simple, entire and smooth, the leaf veins are reticulate.
The inflorescences arise either directly on the trunk and thicker branches, or are laterally or terminally on the branches. The hermaphrodite flowers are zygomorphic . There are six sepals and six petals . The very numerous stamens are in a ring around the upper half permanent ovary organized the everts on one side to a hood. This hood is placed over the stamens of the ring and rolled up inside, it carries sterile staminodes and produces nectar inside. Pollinating insects have to crawl under this hood, eat the staminodes or nectar and transfer the pollen in the process .
The round to conical and pot-shaped capsule fruits stick to the tree when they ripen, open there with a lid (operculum) and release the seeds. The seeds are wingless, the inside of the fruit is not filled with “ pulp ”. The seeds can have an aril or a sarcotesta .
Systematics and distribution
The genus Eschweilera was established in 1828 by Carl Friedrich Philipp von Martius in Augustin-Pyrame de Candolle : Prodromus Systematis Naturalis Regni Vegetabilis , Pars 3, p. 293. The botanical genus name Eschweilera honors the botanist Franz Gerhard Eschweiler . Synonyms for Eschweilera Mart. ex DC. are: Jugastrum Miers , Noallia Buc'hoz , Neohuberia Ledoux .
About 90 Eschweilera species have been described, and estimates suggest that just as many are still undescribed. According to molecular genetic studies, Eschweilera is paraphyletic (the species assigned to the genus form two clades, which, however, are not sister taxa).
The neotropical genus Eschweilera is widespread from Mexico to eastern Brazil, making it the most widespread genus within the potted fruit tree family. Eschweilera species usually thrive at low altitudes of up to 1000 meters, dry or flooded areas are avoided.
There are around 90 species of Eschweilera :
- Eschweilera aguilarii S.A.Mori : The home is Costa Rica .
- Eschweilera alata A.C.Sm. : It iswidespreadfrom Guyana to southeastern Venezuela .
- Eschweilera albiflora ( DC.) Miers (Syn .: Eschweilera inaequisepala Cuatrec. , Eschweilera pachysepala (Spruce ex O.Berg) Miers , Eschweilera turbinata (O.Berg) Nied. , Eschweilera verruculosa (O.Berg) Miers ): It is Colombia Spreadover Bolivia to Peru and Brazil .
- Eschweilera alvimii S.A. Mori : The home is northeastern Brazil.
- Eschweilera amazonica R.Knuth : The home is northern Brazil.
- Eschweilera amazoniciformis S.A. Mori : The home is northern Brazil.
- Eschweilera amplexifolia S.A. Mori : It is distributed from Costa Rica, Panama and Colombia.
- Eschweilera andina (Rusby) JFMacbr. (Syn .: Eschweilera knuthii J.F. Macbr. , Eschweilera loretensis R.Knuth ): It is distributed from western Venezuela through Bolivia, Ecuador to Peru and northern Brazil.
- Eschweilera antioquensis Dugand & Daniel (Syn .: Eschweilera montana Cuatrec. , Eschweilera neomontana Cuatrec. , Eschweilera violacea Cuatrec. ): Panama to northwestern Venezuela and Ecuador.
- Eschweilera apiculata (Miers) ACSm. : French Guiana to northern Brazil.
- Eschweilera atropetiolata S.A.Mori : Northern Brazil.
- Eschweilera baguensis S.A.Mori : Northern Peru.
- Eschweilera beebei Pittier ex SAMori : Venezuela.
- Eschweilera biflava S.A.Mori : Costa Rica.
- Eschweilera bogotensis R.Knuth (Syn .: Eschweilera kalbreyeri R.Knuth , Eschweilera oligosperma Cuatrec. , Eschweilera papillata L.Uribe ): Colombia.
- Eschweilera boltenii S.A.Mori : Suriname.
- Eschweilera bracteosa (Poepp. Ex Berg) Miers : Colombia to southern Venezuela and Peru.
- Eschweilera cabrerana Philipson : Colombia.
- Eschweilera calyculata Pittier (Syn .: Eschweilera collinsii Pittier ): Costa Rica to Colombia.
- Eschweilera carinata S.A. Mori : Northern Brazil.
- Eschweilera caudiculata Engl. (Syn .: Eschweilera cauliflora R. Knuth ): Panama to Peru.
- Eschweilera chartaceifolia S.A. Mori : Colombia to northern Brazil and Peru.
- Eschweilera collina Eyma : Northern South America to northern Brazil.
- Eschweilera collinsii Pittier (Syn .: Eschweilera longirachis S.A. Mori ): Costa Rica to Panama.
- Eschweilera complanata S.A. Mori : Brazil.
- Eschweilera compressa (Vell.) Miers (Syn .: Eschweilera angustifolia Mart. , Eschweilera guanabarica Miers ): Brazil.
- Eschweilera congestiflora (Benoist) Eyma : French Guiana and Suriname.
- Eschweilera coriacea (DC.) SAMori (Syn .: Jugastrum coriaceum (DC.) Miers , Lecythis coriacea DC. , Eschweilera grandifolia Mart. Ex DC. , Chytroma grandifolia (Mart. Ex DC.) Miers , Lecythis grandifolia O.Berg , Lecythis odora Poepp. ex O.Berg , Eschweilera odora (Poepp. ex O.Berg) Miers , Lecythis acuminatissima O.Berg , Eschweilera acuminatissima (O.Berg) Miers , Chytroma cincturata Miers , Eschweilera pallida Miers , Eschweilera matamata Huber , Lecythis retroflexa Benoist , Eschweilera retroflexa (Benoist) R.Knuth , Eschweilera vageleri R.Knuth , Eschweilera eymaana R.Knuth , Eschweilera fractiflexa R.Knuth , Neohuberia matamata Ledoux , Lecythis peruviana L.O.Williams ): Tropical America from Honduras to Brazil.
- Eschweilera correae J.E.Bat. & SAMori : It was first described in 2017 from Costa Rica and Panama.
- Eschweilera costaricensis S.A. Mori : Nicaragua to Costa Rica.
- Eschweilera cyathiformis S.A. Mori : Northern Brazil.
- Eschweilera decolorans Sandwith (Syn .: Eschweilera sandwithiana A.C.Sm. & Beard ): Tobago to tropical South America.
- Eschweilera donosoensis J.E.Bat. & SAMori : It was first described from Panama in 2017.
- Eschweilera eperuetorum Sandwith : Guiana.
- Eschweilera fanshawei Sandwith : Guiana.
- Eschweilera foetulenta ined.
- Eschweilera gigantea (R.Knuth) JFMacbr. (Syn .: Eschweilera mexiana (R.Knuth) JFMacbr. ): Colombia, Ecuador, Peru.
- Eschweilera grandiflora (Aubl.) Sandwith (Syn .: Lecythis grandiflora Aubl. , Eschweilera alba R.Knuth , Eschweilera fracta R.Knuth ): French Guiana to Peru.
- Eschweilera harmonii S.A. Mori : Costa Rica.
- Eschweilera hondurensis Standl. : Honduras, Nicaragua, Costa Rica and Panama.
- Eschweilera integricalyx S.A.Mori Colombia.
- Eschweilera integrifolia (Ruiz & Pav. Ex Miers) R.Knuth (Syn .: Eschweilera garagarae Pittier , Eschweilera woodsoniana Dwyer ): Costa Rica to Ecuador.
- Eschweilera itayensis R.Knuth : Colombia, Peru and northern Brazil.
- Eschweilera jacquelyniae S.A.Mori : Panama to Colombia.
- Eschweilera juruensis R.Knuth : Colombia, Bolivia, Peru and northern Brazil.
- Eschweilera klugii R.Knuth : Northern Peru.
- Eschweilera laevicarpa S.A.Mori : Tropical South America.
- Eschweilera longipedicellata S.A. Mori : Panama to Colombia.
- Eschweilera macrocarpa Pittier : Northern Venezuela.
- Eschweilera mattos-silvae S.A.Mori Brazil.
- Eschweilera mexicana T. Wendt, SAMori & Prance : Mexico.
- Eschweilera micrantha (O.Berg) Miers (Syn .: Eschweilera floribunda Eyma , Eschweilera gracilipes (Sagot) R.Knuth , Eschweilera polyantha A.C.Sm. ): Northern South America to western Brazil.
- Eschweilera microcalyx S.A. Mori : Colombia.
- Eschweilera nana (O.Berg) Miers (Syn .: Eschweilera trochiformis Miers ): Brazil.
- Eschweilera neblinensis S.A. Mori : Venezuela.
- Eschweilera neei S.A. Mori : Panama to Colombia.
- Eschweilera nitida ined.
- Eschweilera obversa (O.Berg) Miers (Syn .: Eschweilera obidosensis R.Knuth ): Brazil.
- Eschweilera ovalifolia (DC.) Nied. (Syn .: Eschweilera ucayalensis R.Knuth ): Southeast Colombia to Bolivia and northern Brazil.
- Eschweilera ovata (Cambess.) Mart. ex Miers (Syn .: Eschweilera acuminata (O.Berg) Miers , Eschweilera blanchetiana (O.Berg) Miers , Eschweilera gracilis Miers , Eschweilera luschnathii (O.Berg) Miers , Eschweilera siberiana (O.Berg) Miers ): Brazil.
- Eschweilera pachyderma Cuatrec. : Panama to Ecuador.
- Eschweilera panamensis Pittier : Costa Rica to Ecuador.
- Eschweilera paniculata (O.Berg) Miers : Colombia to Venezuela.
- Eschweilera parviflora (Aubl.) Miers (Syn .: Lecythis parviflora Aubl. , Lecythis acuminata DC. , Eschweilera grata Sandwith , Eschweilera montana A.C.Sm. ): Northern South America to Northern Brazil.
- Eschweilera parvifolia Mart. ex DC. (Syn .: Lecythis elegans O.Berg , Lecythis caffanjonii Benoist , Eschweilera caffanjonii (Benoist) R.Knuth , Eschweilera elegans (O.Berg) Miers , Eschweilera iquitosensis R.Knuth , Eschweilera krukovii A.C.Sm. , Eschweilera lancifolia Cuatrec. , Eschweilera timbuchensis R.Knuth ): Tropical South America.
- Eschweilera pedicellata (Rich.) SAMori (Syn .: Eschweilera flaccida Miers , Eschweilera idatimonoides (O.Berg) Miers , Eschweilera longipes (Poit.) Miers , Eschweilera macrophylla (O.Berg) Miers , Eschweilera pilosa (Poepp. Ex O.Berg ) Miers , Eschweilera platycarpa (Poit.) Miers ): Southeast Colombia to northern South America and western Brazil.
- Eschweilera perumbonata Pittier : Venezuela.
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Eschweilera piresii S.A.Mori : With two subspecies:
- Eschweilera piresii S.A.Mori subsp. piresii : Brazil.
- Eschweilera piresii subsp. viridipetala S.A. Mori : French Guiana.
- Eschweilera potaroensis Sandwith : Guiana.
- Eschweilera praealta (Sprague) Sandwith : Colombia.
- Eschweilera pseudodecolorans S.A. Mori : Northern Brazil.
- Eschweilera punctata S.A. Mori : Colombia to northern Brazil.
- Eschweilera raven liana S.A.Mori Brazil.
- Eschweilera rankiniae S.A.Mori Brazil.
- Eschweilera reversa Pittier (Syn .: Eschweilera pittieri R.Knuth , Eschweilera verruculosa Pittier ): Costa Rica to Ecuador.
- Eschweilera revoluta S.A. Mori : Venezuela.
- Eschweilera rhododendrifolia (R.Knuth) ACSm. : Northern Brazil.
- Eschweilera rimbachii Standl. : Colombia to Ecuador.
- Eschweilera rionegrense S.A.Mori : Southern Venezuela to northern Brazil.
- Eschweilera rodriguesiana S.A. Mori : Northern Brazil.
- Eschweilera roraimensis S.A. Mori : Southern Venezuela to Northern Brazil.
- Eschweilera rotundicarpa J.E.Bat. & SAMori : This species was first described from Panama in 2017.
- Eschweilera rufifolia S.A.Mori : Southeastern Colombia to northern Peru.
- Eschweilera sagotiana Miers (Syn .: Eschweilera jucunda (Benoist) R.Knuth , Eschweilera melinonis (Sagot) R.Knuth , Eschweilera tapuya (Benoist) R.Knuth ): Northern South America to Northern Brazil.
- Eschweilera sclerophylla Cuatrec. : Colombia.
- Eschweilera sessilis A.C.Sm. (Syn .: Eschweilera ciroana R.Knuth ): Costa Rica to Colombia.
- Eschweilera simiorum (Benoist) Eyma (Syn .: Eschweilera simiorum var. Latifolia Eyma ): Northern South America to northern Brazil.
- Eschweilera sphaerocarpa M. Ribeiro & SAMori : It was first described from Brazil in 2015.
- Eschweilera squamata S.A. Mori : French Guiana.
- Eschweilera subcordata S.A. Mori : Brazil.
- Eschweilera subglandulosa (Steud. Ex O.Berg) Miers (Syn .: Eschweilera laevifolia Miers ): Trinidad to northern Brazil.
- Eschweilera tenax (Moritz ex O.Berg) Miers (Syn .: Eschweilera fendleriana Miers , Eschweilera monosperma Pittier , Eschweilera trinitensis A.C.Sm. & Beard ): Trinida to Venezuela.
- Eschweilera tenuifolia (O.Berg) Miers (Syn .: Eschweilera obtecta (Miers) ACSm. , Eschweilera riedeliana (O.Berg) Miers ): Southeastern Colombia to Venezuela and northern Brazil.
- Eschweilera tessmannii R.Knuth : It occurs in northern Brazil and Peru.
- Eschweilera tetrapetala S.A. Mori : It occurs in Brazil.
- Eschweilera truncata A.C.Sm. (Syn .: Eschweilera romeu-cardosoi S.A.Mori ): It occurs in northwestern Brazil.
- Eschweilera venezuelica S.A. Mori : It occurs in northern Venezuela.
- Eschweilera wachenheimii (Benoist) Sandwith : It occurs in northern South America.
The following is no longer included in this category:
- Eschweilera rhodogonoclada Rizzini & A.Mattos => Lecythis lurida (Miers) SAMori
swell
- SAMori, GTPrance, 2006 onward: The Lecythidaceae Pages. The New York Botanical Garden, Bronx, New York. online , accessed August 15, 2007.
Individual evidence
- ↑ First publication scanned at biodiversitylibrary.org .
- ^ Frans A. Stafleu, Richard S. Cowan: Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types. Volume 1 AG, Bohn, Scheltema & Holkema, Utrecht, 2nd edition, 1976. P. 801.
- ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv Rafaël Govaerts (Ed.): Eschweilera. In: World Checklist of Selected Plant Families (WCSP) - The Board of Trustees of the Royal Botanic Gardens, Kew . Retrieved October 28, 2018.
- ↑ Complete species list from Lecythidaceae Pages .
- ↑ Eschweilera in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
Web links
- Photos: Wood from Eschweilera "black" ( memento from March 26, 2016 in the Internet Archive ), wood from Eschweilera confertifolia ( Memento from March 4, 2016 in the Internet Archive ), wood from Eschweilera conjugata ( Memento from March 4, 2016 in the Internet Archive ).
- Eschweilera alvimii in the endangered Red List species the IUCN 2006 Posted by: J. Pires O'Brien, 1998. Retrieved on 14 August, 2007.
- Eschweilera cyathiformis in the endangered Red List species the IUCN 2006. Posted by: World Conservation Monitoring Center, 1998. Retrieved on 14 August, 2007.
- Eschweilera mexicana in the endangered Red List species the IUCN 2006. Posted by: World Conservation Monitoring Center, 1998. Retrieved on 14 August, 2007.