Systematics of monitor lizards

from Wikipedia, the free encyclopedia
States of Australia

There are currently over 80 species in the genus of monitor lizards ( Varanus ). The list can be sorted according to all criteria. If sorting is done according to images, then it is sorted primarily according to the presence of an image and secondarily according to scientific name . Scientific names are sorted alphabetically. If you sort by German name, then it is sorted primarily by the presence of a German name and secondarily by alphabetical order. In case of doubt, only the scientific name was listed. If the species are arranged according to distribution, they are sorted primarily according to the categories Africa, Asia and Australia, and secondarily according to alphabetical order of the scientific names. The sorting according to subgenus follows the analyzes of the hemipenis morphology by Ziegler & Böhme (1997) and Böhme (2003). The sorting in clades follows the studies of mtDNA by Ast (2001) and Fitch et al. (2006). Even if the affiliation to a cladus is usually more or less clear, this is only included in the case of an explicit DNA analysis.

For species native to Australia, abbreviations for the states were used to specify the area of ​​distribution more precisely, NT = Northern Territory , QLD = Queensland , WA = Western Australia , SA = South Australia , NSW = New South Wales , ACT = Australian Capital Territory , VIC = Victoria .

Species list

image scientific name German name distribution Subgenus Kladus
Barbed tail monitor (Varanus acanthurus) Varanus acanthurus
Boulenger , 1885
 Spiny tail monitor Australia:
N-WA, NT, NW-QLD 		Odatria :
V. acanthurus group 		Indo-Australia:
Odatria :
V. acanthurus group
White-throated monitor (Varanus albigularis) Varanus albigularis
Daudin , 1802 		White-throated monitor Africa:
S-Africa, W-Africa 		Polydaedalus :
V. exanthematicus group 		Africa
Auffenberg's monitor lizard (Varanus auffenbergi) Varanus auffenbergi
Sprackland , 1993 		Auffenberg's monitor lizard Asia:
Lesser Sunda Islands ( Roti ) 		Odatria :
V. timorensis group 		-
- Varanus bangonorum
Welton , Travers , Siler & Brown , 2014 		- Asia:
Philippines 		Soterosaurus Indo-Asia A:
V. Salvator group
- Varanus baritji
King & Horner , 1987 		- Australia:
N-NT 		Odatria :
V. acanthurus group 		Indo-Australia:
Odatria :
V. acanthurus group
Black tree monitor (Varanus beccarii) Varanus beccarii
( Doria , 1874) 		Black tree monitor Asia:
Moluccas ( Aru Islands ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
Bengal monitor (Varanus bengalensis) Varanus bengalensis
( Daudin , 1802) 		Bengal monitor Asia:
From SE-Iran to SE to Java 		Empagusia :
V. bengalensis group 		Indo-Asia A:
V. Salvator group
Varanus bennetti
Weijola el al., 2020 		Oceania:
Palau , western Carolines a . Sarigan 		Euprepiosaurus :
Varanus bitatawa Varanus bitatawa
Welton et al., 2010 		- Asia:
Philippines 		Philippinosaurus -
Varanus boehmei Varanus boehmei
Jacobs , 2003 		- Australia (Oceania):
New Guinea Raja Ampat ( Waigeo ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
- Varanus bogerti
Mertens , 1950 		- Australia (Oceania):
SE New Guinea ( D'Entrecasteaux Islands , Trobriand Islands ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
- Varanus brevicauda
Boulenger , 1898 		Short-tailed monitor Australia:
WA, S-NT, N-SA, W-QLD 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
- Varanus bushi
Aplin , Fitch & King , 2006 		Bush's dwarf monitor Australia:
Pilbara 		Odatria :
V. acanthurus group 		Indo-Australia:
Odatria :
V. acanthurus group
- Varanus caerulivirens
Ziegler , Böhme & Philipp , 1999 		Turquoise monitor Asia:
Moluccas ( Halmahera ) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group
Striped-tail monitor (Varanus caudolineatus) Varanus caudolineatus
Boulenger , 1885 		Striped-tail monitor Australia:
WA, S-NT, SA 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
- Varanus cerambonensis
Philipp , Böhme & Ziegler , 1999 		- Asia:
Z- Moluccas (Ambon, Banda, Buru, Obi & Seram) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group
Cuming's monitor lizard (Varanus cumingi) Varanus cumingi
Martin , 1838 		Cumings monitor lizard Asia:
Philippines 		Soterosaurus Indo-Asia A:
V. Salvator group
- Varanus dalubhasa
Welton , Travers , Siler & Brown , 2014 		- Asia:
Philippines 		Soterosaurus Indo-Asia A:
V. Salvator group
- Varanus doreanus
( Meyer , 1874) 		Blue-tailed monitor Australia (Oceania):
New Guinea 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
- Varanus douarrha
Lesson , 1830 		New Ireland white monitor monitor Australia (Oceania):
New Ireland 		- -
Duméril's monitor lizard (Varanus dumerilii) Varanus dumerilii
Schlegel , 1839 		Dumérils monitor lizard Asia:
South Indian Peninsula, Borneo, Sumatra 		Empagusia Indo-Asia A:
V. Salvator group
- Varanus eremius
Lucas & Frost , 1895 		Hermit monitor Australia:
WA, S-NT, N-SA, SW-QLD 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
Steppe monitor (Varanus exanthematicus) Varanus exanthematicus
( Bosc , 1792) 		Steppe monitor Africa:
Sub-Saharan Africa from N to the Equator 		Polydaedalus :
V. exanthematicus group 		Africa
- Varanus finschi
Böhme , Horn & Ziegler , 1994 		Finsch's monitor lizard Australia (Oceania):
New Guinea & surrounding islands ( New Ireland , Numfor ), N-QLD 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
Yellow monitor (Varanus flavescens) Varanus flavescens
( Hardwicke & Gray , 1827) 		Yellow monitor Asia:
E-Pakistan, N-India, Nepal & Bangladesh 		Empagusia Indo-Asia A:
V. Salvator group
Giant monitor (Varanus giganteus) Varanus giganteus
( Gray , 1845) 		Giant monitor Australia:
W-QLD via Z-Australia to O-WA 		Varanus Indo-Australia:
V. gouldii group
Gillens monitor lizard (Varanus gilleni) Varanus gilleni
Lucas & Frost , 1895
 Gillens monitor lizard Australia:
Kimberley , N-NT, NW-QLD 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
Glauert's rock monitor (Varanus glauerti) Varanus glauerti
Mertens , 1957 		Glauert's rock monitor Australia:
N-WA, N-NT 		Odatria Indo-Australia:
Odatria :
V. tristis group
- Varanus glebopalma
Mitchell , 1955 		Long-tailed rock monitor Australia:
Kimberley , N-NT, NW-QLD 		Odatria Indo-Australia:
Odatria :
V. tristis group
Gould's monitor lizard (Varanus gouldii) Varanus gouldii
Gray , 1838 		Gould's monitor lizard Australia:
Complete excl. S-VIC, Z-QLD & S-WA 		Varanus :
V. gouldii group 		Indo-Australia:
V. gouldii group
Desert monitor (Varanus griseus) Varanus griseus
( Daudin , 1803) 		Desert monitor Africa & Asia:
N-Africa to NW-India and Pakistan 		Psammosaurus Africa
- Varanus hamersleyensis
Maryan , Oliver , Fitch & O'Connell , 2014 		Southern Pilbara rock monitor Australia:
southern Pilbara (WA) 		Odatria Indo-Australia:
Odatria :
V. tristis group
Pacific monitor (Varanus indicus) Varanus indicus
( Daudin , 1802) 		Pacific monitor Asia:
SE-Southeast Asia, New Guinea, N-Australia 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
Sepik monitor lizard (Varanus jobiensis) Varanus jobiensis
Ahl , 1932 		Sepik monitor lizard Australia (Oceania):
New Guinea 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
- Varanus juxtindicus
Böhme , Philipp & Ziegler , 2002 		Renell Island Monitor Lizard Australia (Oceania):
Solomon Islands ( Rennell ) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group
- Varanus keithhornei
Wells & Wellington , 1985 		Australian tree monitor Australia:
N-QLD 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
Kings monitor lizard (Varanus kingorum) Varanus kingorum
Storr , 1980 		Kings monitor lizard Australia:
NO-WA, NW-NT 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
Komodo dragon (Varanus komodoensis) Varanus komodoensis
Ouwens , 1912 		Komodo dragon Asia:
Lesser Sunda Islands ( Komodo , Rinca , Flores , Gili Motang , Padar ) 		Varanus Indo-Australia:
V. varius group
- Varanus kordensis
( Meyer , 1874) 		Biak tree monitor Australia (Oceania):
New Guinea ( Biak ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
- Varanus lirungensis
Koch , Arida , Schmitz , Böhme & Ziegler , 2009 		- Asia:
Talaud Islands ( Salibabu ) 		Euprepiosaurus :
V. indicus group 		-
- Varanus mabitang
Gaulke & Curio , 2001 		Panay monitor lizard Asia:
Philippines 		Philippinosaurus -
Blue tree monitor (Varanus macraei) Varanus macraei
Böhme & Jacobs , 2001 		Blue tree monitor Australia (Oceania):
New Guinea Raja Ampat ( Batanta ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
Varanus marmoratus Varanus marmoratus
( Wiegmann , 1834) 		- Asia:
Philippines 		Soterosaurus -
Quince monitor (Varanus melinus) Varanus melinus
Böhme & Ziegler , 1997 		Quince monitor Asia:
Moluccas ( Sula Islands ) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
Mertens water monitor (Varanus mertensi) Varanus mertensi
Glauert , 1951 		Mertens water monitor Australia:
N-WA, N-NT, N-QLD 		Varanus Indo-Australia:
V. gouldii group
- Varanus mitchelli
Mertens , 1958 		Mitchell's monitor lizard Australia:
NO-WA, N-NT 		Odatria Indo-Australia:
Odatria :
V. tristis group
Fog monitor (Varanus nebulosus) Varanus nebulosus
( Gray , 1831) 		Fog monitor Asia:
From SE-Iran to SE to Java 		Empagusia :
V. bengalensis group 		Indo-Asia A:
V. Salvator group
- Varanus nesterovi
Böhme , Ehrlich , Milto , Orlov & Scholz , 2015 		- Asia:
Zāgros Mountains (Iran and Iraq) 		Psammosaurus -
Nile monitor (Varanus niloticus) Varanus niloticus
( Linnaeus , 1766) 		Nile monitor Africa:
Africa excl. Sahara 		Polydaedalus :
V. nilotics group 		Africa
- Varanus nuchalis
( Günther , 1872) 		Philippines white monitor monitor Asia:
Philippines 		Soterosaurus -
- Varanus obor
Weijola & Sweet , 2009 		- Asia:
Moluccas ( Sanana ) 		Euprepiosaurus :
V. indicus group 		-
Gray's monitor lizard (Varanus olivaceus) Varanus olivaceus
Hallowell , 1857 		Gray's monitor lizard Asia:
Philippines 		Philippinosaurus Indo-Asia B:
V. olivaceus group
Rainforest Nile Monitor (Varanus ornatus) Varanus ornatus
( Daudin , 1803) 		Rainforest Nile Monitor Africa:
Z-Africa 		Polydaedalus :
V. nilotics group 		Africa
- Varanus palawanensis
Koch , Gaulke & Böhme , 2010 		- Asia:
Philippines 		Soterosaurus -
Argus monitor (Varanus panoptes) Varanus panoptes
Storr , 1980 		Argus monitor Australia:
N-WA, W-WA, N-NT, QLD & S-New Guinea 		Varanus :
V. gouldii group 		Indo-Australia:
V. gouldii group
- Varanus pilbarensis
Storr , 1980 		Pilbara rock monitor Australia:
Northern Pilbara (WA) 		Odatria Indo-Australia:
Odatria :
V. tristis group
Emerald monitor (Varanus prasinus) Varanus prasinus
( Schlegel , 1839) 		Emerald monitor Australia (Oceania):
New Guinea 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
- Varanus primordius
Mertens , 1942 		Dwarf monitor Australia:
N-NT 		Odatria :
V. acanthurus group 		Indo-Australia:
Odatria :
V. acanthurus group
- Varanus rainerguentheri
Ziegler , Böhme & Schmitz , 2002 		Halmahera Pacific Monitor Asia:
Moluccas 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group
- Varanus rasmusseni
Koch , Gaulke & Böhme , 2010 		- Asia:
Philippines 		Soterosaurus -
Reisinger's tree monitor (Varanus reisingeri) Varanus reisingeri
Eidenmüller & Wicker , 2005 		Reisinger's tree monitor Australia (Oceania):
New Guinea Raja Ampat ( Misool ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
Rosenberg's monitor lizard (Varanus rosenbergi) Varanus rosenbergi
Mertens , 1957 		Rosenberg's monitor lizard Australia:
S-WA, S-SA, ACT, S-NSW 		Varanus :
V. gouldii group 		Indo-Australia:
V. gouldii group
Rough-naped monitor (Varanus rudicollis) Varanus rudicollis
( Gray , 1845) 		Raunackenwaran Asia:
Malay Peninsula, Borneo, Sumatra 		Empagusia Indo-Asia A:
V. Salvator group
Papuan dragon (Varanus salvadorii) Varanus salvadorii
( Peters & Doria , 1878) 		Papuan dragon Australia (Oceania):
New Guinea 		Papusaurus Indo-Australia:
V. varius group
White monitor monitor (Varanus salvator) Varanus salvator
( Laurenti , 1768) 		Bandage monitor Asia:
All of SE Asia excl. Philippines 		Soterosaurus Indo-Asia A:
V. Salvator group
- Varanus samarensis
Koch , Gaulke & Böhme , 2010 		- Asia:
Philippines 		Soterosaurus Indo-Asia A:
V. Salvator group
Banded tree monitor (Varanus scalaris) Varanus scalaris
Mertens , 1941 		Banded tree monitor Australia:
N-WA, N-NT, N-QLD, S-New Guinea 		Odatria :
V. timorensis group 		Indo-Australia:
Odatria :
V. tristis group
- Varanus semiremex
Peters , 1869 		Rust-headed monitor Australia:
O-QLD (Cape York to Brisbane) 		Odatria Indo-Australia:
Odatria :
V. tristis group
- Varanus semotus
Weijola , Donnellan & Lindqvist , 2016 		- Australia (Oceania):
New Guinea ( Mussau Islands ) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B:
V. indicus group
- Varanus similis
Mertens , 1958
 Spotted tree monitor Australia:
N-Australia, S-New Guinea 		Odatria :
V. timorensis group 		Indo-Australia:
Odatria :
V. tristis group
- Varanus sparnus
Doughty , Kealley , Fitch & Donnellan , 2014 		- Australia:
WA 		Odatria Indo-Australia:
Odatria :
V. acanthurus group
Spencer's monitor lizard (Varanus spenceri) Varanus spenceri
Lucas & Frost , 1903 		Spencer's monitor lizard Australia:
W-QLD, O-NT 		Varanus Indo-Australia:
V. gouldii group
Varanus spinulosus Varanus spinulosus
Mertens , 1941 		- Australia (Oceania):
Solomon Islands 		Incertae sedis -
Storrs dwarf monitor (Varanus storri) Varanus storri
Mertens , 1966
 Storrs dwarf monitor Australia:
WA, NT, QLD 		Odatria :
V. acanthurus group 		Indo-Australia:
Odatria :
V. acanthurus group
- Varanus telenesetes
Sprackland , 1991 		- Australia (Oceania):
New Guinea ( Rossel Island ) 		Euprepiosaurus :
V. prasinus group 		Indo-Asia B:
V. prasinus group
Timor monitor lizard (Varanus timorensis) Varanus timorensis
Gray , 1831 		Timor monitor lizard Asia:
Lesser Sunda Islands ( Timor , Sawu , Semau ) 		Odatria :
V. timorensis group 		Indo-Australia:
Odatria :
V. tristis group
- Varanus togianus
( Peters , 1872) 		- Asia:
Greater Sunda Islands ( Sulawesi & Togian Islands ) 		Soterosaurus Indo-Asia A:
V. Salvator group
Black monitor monitor (Varanus tristis) Varanus tristis
( Schlegel , 1838)
 Black monitor Australia:
All of Australia excl. South (S-SA, S-NSW, ACT, VIC) 		Odatria :
V. timorensis group 		Indo-Australia:
Odatria :
V. tristis group
Varanus tsukamotoi
Kishida , 1929 		Oceania:
Guam , Mariana Islands 		Euprepiosaurus :
Colored monitor (Varanus varius) Varanus varius
( Shaw in White , 1790) 		Colored monitor Australia:
O-QLD, NSW, ACT, VIC & SO-SA 		Varanus Indo-Australia:
V. varius group
- Varanus yemenensis
Böhme , Joger & Schätti , 1989 		Yemen monitor lizard Asia:
Arabian Peninsula 		Polydaedalus :
V. exanthematicus group 		Africa
- Varanus yuwonoi
Harvey & Barker , 1998 		- Asia:
Moluccas (Halmahera) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group
- Varanus zugorum
Böhme & Ziegler , 2002 		- Asia:
Moluccas (Halmahera) 		Euprepiosaurus :
V. indicus group 		Indo-Asia B: V. indicus group

literature

  • JC Ast (2001): Mitochondrial DNA Evidence and Evolution in Varanoidea (Squamata) . Cladistics 17, pp. 211-226
  • W. Böhme (2003): Checklist of the living monitor lizards of the world (family Varanidae) . Zoologische Verhandelingen 341, pp. 3-43
  • AJ Fitch, AE Goodman & SC Donnellan (2006): A molecular phylogeny of the Australian monitor lizards (Squamata: Varanidae) inferred from mitochondrial DNA sequences . Australian Journal of Zoology 54, pp. 253-269
  • A. Koch, M. Auliya & T. Ziegler (2010): Updated checklist of the living monitor lizards of the world (Squamata: Varanidae) . Bonn Zoological Bulletin 57 (2), pp. 127-136
  • T. Ziegler, A. Schmitz, A. Koch & W. Böhme (2007): A review of the subgenus Euprepiosaurus of Varanus (Squamata: Varanidae): Morphological and molecular phylogeny, distribution and zoogeography, with an identification key for the members of the V. indicus and the V. prasinus species groups . Zootaxa 1472, pp. 1-28
  • T. Ziegler & W. Böhme (1997): Genital structures and mating biology in squamatic reptiles, especially the Platynota, with comments on the system . Mertensiella 8: 3-210
  • Varanus in The Reptile Database ; Retrieved February 5, 2011.

supporting documents

  1. ↑ Varanus in The Reptile Database , accessed May 13, 2020.
  2. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da W. Böhme (2003): Checklist of the living monitor lizards of the world (family Varanidae) . Zoologische Verhandelingen 341, pp. 3-43
  3. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw Information from the respective species chapter in ER Pianka & DR King (Ed.): Varanoid Lizards of the World . Indiana University Press, Bloomington & Indianapolis. ISBN 0253343666
  4. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab A. J. Fitch, AE Goodman & SC Donnellan (2006): A molecular phylogeny of the Australian monitor lizards ( Squamata: Varanidae) inferred from mitochondrial DNA sequences . Australian Journal of Zoology 54, pp. 253-269
  5. ↑ a b c African Varanid Species . In: ER Pianka & DR King (Eds.): Varanoid Lizards of the World , p. 90. Indiana University Press, Bloomington & Indianapolis. ISBN 0253343666
  6. ↑ a b c d e f g Luke J. Welton, Scott L. Travers, Caeron D. Siler & Rafe M. Brown: Integrative taxonomy and phylogeny-based species delimitation of Philippine water monitor lizards (Varanus salvator Complex) with descriptions of two new cryptic species. Zootaxa 3881 (3): 201-227 (5 Nov. 2014), doi: 10.11646 / zootaxa.3881.3.1
  7. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am Ziegler, T .; A. Schmitz, A. Koch & W. Böhme (2007): A review of the subgenus Euprepiosaurus of Varanus (Squamata: Varanidae): Morphological and molecular phylogeny, distribution and zoogeography, with an identification key for the members of the V. indicus and the V. prasinus species groups . Zootaxa 1472, pp. 1-28
  8. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai A. Koch, M. Auliya & T. Ziegler (2010) : Updated checklist of the living monitor lizards of the world (Squamata: Varanidae) . Bonn Zoological Bulletin 57 (2), pp. 127-136
  9. ↑ a b c d e f g h i j k l m J. C. Ast (2001): Mitochondrial DNA Evidence and Evolution in Varanoidea (Squamata) . Cladistics 17, pp. 211-226
  10. ↑ a b c d e f Valter Weijola, Varpu Vahtera, André Koch, Andreas Schmitz and Fred Kraus. 2020. Taxonomy of Micronesian monitors (Reptilia: Squamata: Varanus ): Endemic Status of New Species argues for Caution in Pursuing Eradication Plans. Royal Society Open Science. 7 (5) DOI: 10.1098 / rsos.200092
  11. ↑ Varanus bushi in The Reptile Database ; Retrieved January 20, 2011.
  12. ↑ A. Koch (2010): Underestimated and exploited: systematics, diversity and endemism of Southeast Asian black monitors . Koenigiana 4 (1), pp. 27-41
  13. ^ A b c d e A. Koch, M. Auliya, A. Schmitz, U. Kuch & W. Böhme (2007): Morphological Studies on the Systematics of South East Asian Water Monitors (Varanus salvator Complex): Nominotypic Populations and Taxonomic Overview . Mertensiella 16 ( Advances in Monitor Research III ), pp. 109-180
  14. ↑ a b c d A. Koch, M. Gaulke & W. Böhme (2010): Unraveling the underestimated diversity of Philippine water monitor lizards (Squamata: Varanus salvator complex), with the description of two new species and a new subspecies . Zootaxa 2446, pp. 1-54
  15. ↑ a b Valter Weijola, Fred Kraus, Varpu Vahtera, Christer Lindqvist and Stephen C. Donnellan. 2017. Reinstatement of Varanus douarrha Lesson, 1830 As A Valid Species with Comments on the Zoogeography of Monitor Lizards (Squamata: Varanidae) in the Bismarck Archipelago, Papua New Guinea. Australian Journal of Zoology. DOI: 10.1071 / ZO16038
  16. ^ German name according to Spektrum.de
  17. ^ A b Maryan B, Oliver PM, Fitch AJ & O'Connell M. 2014: Molecular and Morphological Assessment of Varanus pilbarensis (Squamata: Varanidae), with a description of a new species from the southern Pilbara, Western Australia. Zootaxa. 3768 (2): 139-158.
  18. ↑ a b c Wolfgang Böhme, Klaus Ehrlich, Konstantin D. Milto, Nikolay Orlov and Sebastian Scholz. 2015. A New Species of Desert Monitor Lizard (Varanidae: Varanus: Psammosaurus) from the Western Zagros region (Iraq, Iran). Russian Journal of Herpetology. 22 (1): 41-52.
  19. ↑ Varanus rainerguentheri in The Reptile Database ; Retrieved February 23, 2011.
  20. ↑ H.-G. Horn, SS Sweet & KM Philipp (2007): On the distribution of the Papuan monitor (Varanus salvadorii Peters & Doria , 1878) in New Guinea . Mertensiella 16 ( Advances in Monitor Research III ), pp. 25-43
  21. ↑ a b Valter Weijola, Stephen Donnellan and Christer Lindqvist. 2016. A New Blue-tailed Monitor Lizard (Reptilia, Squamata, Varanus) of the Varanus indicus group from Mussau Island, Papua New Guinea. ZooKeys. 568: 129-154. DOI: 10.3897 / zookeys.568.6872
  22. ↑ Paul Doughty, Luke Kealley, Alison Fitch and Stephen C. Donnellan. 2014. A New Diminutive Species of Varanus from the Dampier Peninsula, western Kimberley region, Western Australia. Records of the Western Australian Museum. 29 128-140.
  23. ↑ a b W. Böhme & T. Ziegler (2007): Notes on the Distribution, Diet, Hemipenis Morphology and Systematifcs of Varanus spinulosus Mertens , 1941 . Mertensiella 16 ( Advances in Monitor Research III ), pp. 100-108
  24. ↑ Varanus zugorum in The Reptile Database ; Retrieved February 23, 2011.

Remarks

  1. ↑ According to Fitch et al. (2006) paraphyletic, since the subspecies V. a. insulanicus appears to be more closely related to V. baritji than to other subspecies of V. acanthurus .
  2. ↑ a b c According to Valter Weijola & Samuel Sweet, only a local form of the Pacific monitor ( Varanus indicus ). see: Weijola V, Sweet SS. 2015. A Single Species of Mangrove Monitor (Varanus) occupies Ambon, Seram, Buru and Saparua, Moluccas, Indonesia. Amphibian & Reptile Conservation. 9 (1) [General Section]: 14–23 (e95).
  3. ↑ According to Fitch et al. (2006) V. gilleni is paraphyletic.
  4. ↑ According to Smith, Sweet & King in Pianka & King (2004) it is seen as a subspecies of V. scalaris , but in the Checklist by Böhme (2003), in the Reptile Database and also according to Fitch et al. (2006) listed as a species.
  5. ↑ According to Fitch et al. (2006) a species complex.
  6. ↑ According to Fitch et al. (2006) a species complex.