Rhamneae
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Illustration of the Purgier Buckthorn ( Rhamnus cathartica ) |
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Rhamneae | ||||||||||||
Hook f. |
Rhamneae are a tribe in the family of buckthorn plants (Rhamnaceae), to which 13 genera are assigned. The genera of the tribe are distributed almost worldwide.
description
The representatives of the tribe grow as trees, shrubs or climbing plants. Some species are armed. The leaves are opposite, almost opposite or alternate. The leaf margin is serrated or entire. The leaf blade is pinnate. Stipules may be absent or often fall off early. Individual, axillary or terminal clusters, umbels , grapes or cymes are formed as inflorescences . Petals may be missing. The ovary is usually above, rarely below, free and usually two, rarely one or four-way. The discus releases nectar and is free or attached to the flower cup . Two permanent styles are formed, which can then be seen at the tip of the fruit. The fruits are fleshy, one to fourfold drupes . The seeds have no or only a thin, fleshy nutrient tissue . The pollen - exine is reticulate, wrinkled, perforated or grooved.
The chromosome numbers are 2n = 12, 20, 24 and 26.
distribution
The representatives of the tribe grow in the entire distribution area of the buckthorn family with the exception of southern South America.
Systematics and research history
Rhamneae is one of the eleven tribes of the family of the cross thorn plants (Rhamnaceae) with Rhamnus as type - species . The tribe is assigned to 13 genera:
- Auerodendron Urb.
- Berchemia Neck. ex DC.
- Berchemiella Nakai
- Condalia Cav.
- Dallachya F. Muell.
- Karwinskia Zucc.
- Krugiodendron Urb.
- Reynosia Griseb.
- Rhamnella Miq.
- Rhamnidium rice sec
- Buckthorn ( Rhamnus L. )
- Sageretia Brongn.
- Scutia (Comm. Ex DC.) Brongn.
The delimitation between the genera Reynosia , Rhamnidium , Auerodendron and Karwinskia is not clear, further studies are pending.
Flowers and branches of Berchemia racemosa
Branches with fruits of Condalia globosa
Branches with fruits of Rhamnela franguloides
Branch with leaves of Sageretia thea
Individual evidence
- ^ Illustration from Franz Eugen Köhler: Köhler's Medizinal-Pflanzen . 1897.
- ↑ a b c D. Medan, C. Schiarend: Rhamnaceae. In: Klaus Kubitzki: The Families and Genera of Vascular Plants . Volume VI, p. 327.
- ^ A b c JE Richardson, MF Fay, QCB Cronk, MW Chase: A Revision of the Tribal Classification of Rhamnaceae . In: Kew Bulletin . Vol. 55, No. 2 . Springer, 2000, p. 332 , doi : 10.2307 / 4115645 .
- ^ Rhamneae in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
literature
- Klaus Kubitzki (Ed.): The Families and Genera of Vascular Plants . Volume 6: Flowering Plants, Dicotyledons: Celastrales, Oxalidales, Rosales, Cornales, Ericales . Springer, Berlin / Heidelberg / New York 2004, ISBN 3-540-06512-1 , pp. 327 (English).
- JE Richardson, MF Fay, QCB Cronk, MW Chase: A Revision of the Tribal Classification of Rhamnaceae . In: Kew Bulletin . Vol. 55, No. 2 . Springer, 2000, p. 311-340 , doi : 10.2307 / 4115645 .