Morinoideae

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Morinoideae
Morina persica

Morina persica

Systematics
Nuclear eudicotyledons
Asterids
Euasterids II
Order : Cardigans (Dipsacales)
Family : Honeysuckle Family (Caprifoliaceae)
Subfamily : Morinoideae
Scientific name
Morinoideae
Burnett

The Morinoideae are a subfamily within the family of Honeysuckle (Caprifoliaceae).

description

Inflorescence of Morina nepalensis with flowers with long corolla tubes

Vegetative characteristics

They are hardy, perennial herbaceous plants that look like thistles or acanthus . The hollow stems are often covered with withered leaves. The opposite or whirling arranged leaves are divided into petioles and leaf blades; the base of the petiole is often around the stem and forms a leaf sheath . The leaf blades often have a thorny edge.

Generative characteristics

The flowers are in pseudo whorls or more or less cephalic inflorescences together with (mostly) thorny bracts . The secondary calyx is formed from four intergrown bracts and has a few to many thorny teeth or bristles and twelve nerves.

The hermaphrodite flowers are strongly zygomorphic and five-fold. The five sepals are fused and the calyx is split or two-lipped on the underside. The five petals are fused together to form a tube that hardly or clearly has two lips, but always ends with five corolla lobes. There is only one circle of dust and only two or all four stamens are fertile . The dust bags have two counters and open along the long side. Two or three carpels are one under permanent, single-chamber ovary grown. The ovules are hanging. The slender style ends in a simple scar.

They form achena-like fruits, which are covered by the secondary cup and durable cup.

Systematics

The Morinoideae were established by Gilbert Thomas Burnett . The type genus is Morina L. . The generic name Morina honors the French doctor and botanist Louis Pierre Morin (approx. 1635–1715).

The subheadings genera according to the latest editions of the Angiosperm Phylogeny Group to the family of Honeysuckle asked (Caprifoliaceae). According to another opinion, they are classified in a separate family Morinaceae. The two (or three) genera with around 13 species occur from southeast Europe to the Himalayas and southwest China (two genera and about ten species).

The subfamily Morinoideae includes only two (or three) genera with about 13 species:

  • Acanthocalyx ( DC. ) Tiegh. , (Syn .: Morina sect. Acanthocalyx DC. ): The only two species occur in the Himalaya and in the Hengduan Mountains :
    • Acanthocalyx alba ( Hand.-Mazz. ) MJCannon
    • Acanthocalyx nepalensis ( D.Don ) MJCannon : There are two subspecies:
      • Acanthocalyx nepalensis subsp. delavayi ( Franch. ) DYHong
      • Acanthocalyx nepalensis ( D.Don ) MJCannon subsp. nepalensis
  • Morina L. (Syn .: Asaphes Sprengel 1827 non DC. 1825, Cryptothladia ( Bunge ) MJCannon , Morina sect. Cryptothladia Bunge ): The approximately ten species distributed from the Balkans to Central Asia and the Eastern Himalayas, for example:
  • Cryptothladia ( Bunge ) MJCannon : This genus is listed in the Flora of China 2011 as a synonym of Morina .

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Individual evidence

  1. ↑ a b c d e f Deyuan Hong, Fred R. Barrie: Morinaceae. In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Eds.): Flora of China . Volume 19: Cucurbitaceae through Valerianaceae, with Annonaceae and Berberidaceae . Science Press / Missouri Botanical Garden Press, Beijing / St. Louis 2011, ISBN 978-1-935641-04-9 , pp. 649 ff . (English, PDF file ).
  2. ↑ Yasin J. Nasir: Morinaceae . In: Flora of West Pakistan . tape 67 . Stewart Herbarium, Gordon College (et al.), Rawalpindi 1974, OCLC 223635800 , p. 1 (English, online ).
  3. ↑ a b c Description of the Morinaceae in DELTA by L. Watson & MJ Dallwitz. (English)
  4. ↑ Lotte Burkhardt: Directory of eponymous plant names. Extended Edition. Botanic Garden and Botanical Museum Berlin, Free University Berlin Berlin 2018. online.
  5. ↑ Angiosperm Phylogeny Group : An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III. In: Botanical Journal of the Linnean Society. Volume 161, No. 2, 2009, pp. 105-121, doi: 10.1111 / j.1095-8339.2009.00996.x .
  6. ↑ Morinaceae in APWebsite . (engl.)
  7. ^ A b Morinoideae in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland. Retrieved April 20, 2018.
  8. ↑ Werner Greuter, Hervé-Maurice Burdet, Guy Long (eds.): Med-Checklist. A critical inventory of vascular plants of the circum-Mediterranean countries . Vol. 3: Dicotyledones (Convolvulaceae - Labiatae) . Conservatoire et Jardin Botanique, Genève 1986, ISBN 2-8277-0153-7 , p. 191 ( online ).

Web links

Commons : Morinoideae  - collection of images, videos and audio files