Papilio (genus)
Papilio | ||||||||||||
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Swallowtail ( Papilio machaon ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Papilio | ||||||||||||
Linnaeus , 1758 |
Papilio is a genus of butterflies ( butterflies ) from the family of knight butterflies (Papilionidae).
features
The adults have long antennae with a slightly curved, club-shaped end that is not compressed. The palpi are very short and do not protrude beyond the eyes. The third, terminal link is very small. The legs are walking legs . The tibia of the hind legs have two thorns. The claw links are not divided. The hind wings are arched and in many species with a tail-like flap. In some species this tail appears only in one sex, in other species it is completely absent. The inner edge of the wings is concave or grooved.
The caterpillars are smooth and bare, often decorated in bright colors. Some species form a fleshy horn. The first two links are slimmer and can be pulled back under the third and fourth links.
The dolls are angular and fixed about in the middle with a silk ribbon.
distribution
The genus Papilio is distributed almost all over the world. However, most species are found in the tropics . There are more species in the new world than in the old world . Between 40 and 50 species are native to Brazil alone , and the main areas of distribution are also on islands in the Indian Ocean , especially Java .
There are only three species native to Europe, these are Papilio alexanor , the Corsican swallowtail ( Papilio hospiton ) and the swallowtail ( Papilio machaon ).
Systematics
The internal system of the genus is still unclear, there are several different proposals to divide the genus into sub-genres and groups. Well-known classifications are those of Munroe from 1961, that of Hancock (1983), that of Igarashi (1984) and that of Hauser et al. From 2002. A molecular genetic study from 2004 comes to still different results.
Types (selection)
Sorted alphabetically within the groups:
Subgenus Papilio Linnaeus, 1758:
-
machaon group:
- Papilio alexanor Esper, 1800
- Papilio brevicauda Saunders, 1869
- Corsican swallowtail ( Papilio hospiton ) Géné, 1839
- Papilio indra Reakirt, 1866
- Swallowtail ( Papilio machaon ) Linnaeus, 1758
- Papilio saharae Oberthür, 1879
- Papilio polyxenes Fabricius, 1775
- Papilio zelicaon Lucas, 1852
Subgenus Princeps Hübner, 1807:
-
antimachus group:
- Papilio antimachus Drury, 1782
-
zalmoxis group:
- Papilio zalmoxis Hewitson, 1864
-
nireus group:
- Papilio aristophontes Oberthür, 1897
- Papilio nireus Linnaeus, 1758
- Papilio charopus Westwood, 1843
- Papilio chrapkowskii Suffert, 1904
- Papilio chrapkowskoides Storace, 1952
- Papilio desmondi van Someren, 1939
- Papilio hornimani Distant, 1879
- Papilio interjectana Vane-Wright, 1995
- Papilio nerminae Koçak, 1983
- Papilio sosia Rothschild & Jordan , 1903
- Papilio thuraui Karsch, 1900
- Papilio ufipa Carcasson, 1961
- Papilio wilsoni Rothschild, 1926
-
cynorta group:
- Papilio arnoldiana Vane-Wright, 1995
- Papilio cynorta Fabricius, 1793
- Papilio plagiatus Aurivillius, 1898
-
dardanus group:
- Papilio dardanus Brown, 1776
- Papilio Constantinus Ward, 1871
- Papilio delalandei Godart, 1824
- Papilio phorcas Cramer, 1775
- Papilio rex Oberthür, 1886
-
zenobia group:
- Papilio cyproeofila Butler, 1868
- Papilio fernandus Fruhstorfer, 1903
- Papilio filaprae Suffert, 1904
- Papilio gallienus Distant, 1879
- Papilio mechowi Dewitz, 1881
- Papilio mechowianus Dewitz, 1885
- Papilio nobicea Suffert, 1904
- Papilio zenobia Fabricius, 1775
-
demodocus group:
- Papilio demodocus Esper, 1799
- Papilio demoleus Linnaeus, 1758
- Papilio erithonioides Grose-Smith, 1891
- Papilio grosesmithi Rothschild, 1926
- Papilio morondavana Grose-Smith, 1891
-
echerioides group:
- Papilio echerioides Trimen, 1868
- Papilio fuelleborni Karsch, 1900
- Papilio jacksoni Sharpe, 1891
- Papilio sjoestedti Aurivillius, 1908
-
oribazus group:
- Papilio oribazus Boisduval, 1836
- Papilio epiphorbas Boisduval, 1833
- Papilio nobilis Rogenhofer, 1891
-
hesperus group:
- Papilio hesperus Westwood, 1843
- Papilio euphranor Trimen, 1868
- Papilio horribilis Butler, 1874
- Papilio pelodurus Butler, 1896
-
menestheus group:
- Papilio menestheus Drury, 1773
- Papilio lormieri Distant, 1874
- Papilio ophidicephalus Oberthür, 1878
-
incertae sedis group
- Papilio andronicus Ward, 1871
- Papilio chitondensis Bivar de Sousa & Fernandes, 1966
- Papilio leucotaenia Rothschild, 1908
- Papilio mackinnoni Sharpe, 1891
- Papilio mangoura Hewitson, 1875
- Papilio manlius Fabricius, 1798
- Papilio microps Storace, 1951
- Papilio phorbanta Linnaeus, 1771
-
demolion group:
- Papilio demolion Cramer, 1776
- Papilio liomedon Moore, 1875
- Papilio gigon C. & R. Felder, 1864
- Papilio euchenor Guérin-Méneville , 1829
- Not assigned:
- Papilio luzviae Schröder & Treadaway, 1991
-
anactus group:
- Papilio anactus MacLeay, 1826
-
aegeus group:
- Papilio aegeus Donovan, 1805
- Papilio bridgei Mathew, 1886
- ? Papilio erskinei Mathew, 1886
- Papilio gambrisius Cramer, 1777
- Papilio inopinatus Butler, 1883
- Papilio ptolychus Godman & Salvin, 1888
- Papilio tydeus C. & R. Felder, 1860
- Papilio weymeri Niepelt, 1914
- Papilio woodfordi Godman & Salvin, 1888
-
godeffroyi group:
- Papilio amynthor Boisduval, 1859
- Papilio godeffroyi Georg Semper , 1866
- Papilio schmeltzi Herrich-Schäffer, 1869
-
polytes group:
- Papilio ambrax Boisduval, 1832
- Papilio polytes Linnaeus, 1758
- Papilio phestus Guérin-Méneville, 1830
-
castor group:
- Papilio castor Westwood, 1842
- Papilio dravidarum Wood-Mason, 1880
- Papilio mahadeva Moore, 1879
-
fuscus group:
- Papilio albinus Wallace, 1865
- Papilio antonio Hewitson, 1875
- Papilio diophantus Grose-Smith, 1883
- Papilio fuscus Goeze, 1779
- Papilio hipponous C. & R. Felder, 1862
- Papilio jordani Fruhstorfer, 1906
- Papilio noblei de Nicéville, 1889
- Papilio pitmani Elwes & de Nicéville, 1887
- Papilio prexaspes C. & R. Felder, 1865
- Papilio sakontala Hewitson, 1864
-
helenus group:
- Papilio helenus Linnaeus, 1758
- Papilio Iswara White, 1842
- Papilio iswaroides Fruhstorfer, 1898
- Papilio nephelus Boisduval, 1836
- Papilio nubilus Staudinger, 1895
- Papilio sataspes C. & R. Felder, 1865
-
memnon group:
- Papilio acheron Grose-Smith, 1887
- Papilio ascalaphus Boisduval, 1836
- Papilio deiphobus Linnaeus, 1758
- Papilio rumanzovia Eschscholtz, 1821
- Papilio forbesi Grose-Smith, 1883
- Papilio lampsacus Boisduval, 1836
- Papilio mayo Atkinson, 1873
- Papilio memnon Linnaeus, 1758
- Papilio oenomaus Godart, 1819
- Papilio polymnestor Cramer, 1775
- Papilio taiwanus Rothschild
-
protenor group:
- Papilio protenor - Spangle
- Papilio alcmenor C. & R. Felder, 1864
- Papilio macilentus Janson, 1877
- Papilio thaiwanus Rothschild, 1898
-
boats group:
- Papilio boats Westwood, 1842
- Papilio elwesi Leech, 1889
- Papilio maraho Shiraki & Sonan, 1934
Subgenus?:
-
agestor group:
- Papilio agestor Gray, 1831
- Papilio epycides Hewitson, 1864
- Papilio slateri Hewitson, 1853
-
clytia group:
- Papilio clytia Linnaeus, 1758
- Papilio paradoxa Zincken, 1831
-
veiovis group:
- Papilio veiovis Hewitson, 1865
-
laglaizei group:
- Papilio laglaizei Depuiset, 1877
- Papilio moerneri Aurivillius, 1919
- Papilio toboroi Ribbe, 1907
- Unnamed group:
- Papilio osmana Jumalon, 1967
- Papilio carolinensis Jumalon, 1967
Subgenus Achillides Hübner, 1819:
-
paris group:
- Papilio arcturus Westwood, 1842
- Papilio bianor Cramer, 1777
- Papilio dialis Leech, 1893
- Papilio doddsi Janet, 1896
- Papilio hoppo Matsumura, 1908
- Papilio karna C. & R. Felder, 1864
- Papilio Krishna Moore, 1857
- Papilio maackii Ménétriés, 1859
- Papilio paris Linnaeus, 1758
- Papilio polyctor Boisduval, 1836
- Papilio elephenor Doubleday, 1845
-
palinurus group:
- Papilio buddha Westwood, 1872
- Papilio crino Fabricius, 1793
- Papilio palinurus Fabricius, 1787
- Papilio blumei Boisduval, 1836
- Unnamed group:
- Papilio anchisiades Esper, 1788
- Papilio chikae Igarashi, 1965
- Papilio peranthus Fabricius, 1787
- Papilio pericles Wallace, 1865
- Papilio lorquinianus C. & R. Felder, 1865
- Papilio neumoegeni Honrath, 1890
-
ulysses group:
- Papilio montrouzieri Boisduval, 1859
- Papilio syfanius Oberthür, 1886
- Papilio ulysses Linnaeus, 1758
Subgenus Heraclides Hübner, 1819:
-
anchisiades group:
- Papilio chiansiades Westwood, 1872
- Papilio epenetus Hewitson, 1861
- Papilio erostratus Westwood, 1847
- Papilio hyppason Cramer, 1775
- Papilio isidorus Doubleday, 1846
- Papilio pelaus Fabricius, 1775
- Papilio oxynius Geyer, 1827
- Papilio rogeri Boisduval, 1836
-
thoas group:
- Papilio andraemon Huebner, 1823
- Papilio androgeus Cramer, 1775
- Papilio aristodemus Esper, 1794
- Papilio aristor Godart, 1819
- Papilio astyalus Godart, 1819
- Papilio caiguanabus Poey, 1852
- Papilio cresphontes Cramer, 1777
- Papilio homothoas Rothschild & Jordan, 1906
- Papilio machaonides Esper, 1796
- Papilio melonius Rothschild & Jordan, 1906
- Papilio ornythion Boisduval, 1836
- Papilio paeon Boisduval, 1836
- Papilio thersites Fabricius, 1775
- King swallowtail ( Papilio thoas ) Linnaeus, 1771
-
torquatus group:
- Papilio garleppi Staudinger, 1892
- Papilio hectorides Esper, 1794
- Papilio himeros Hopffer, 1865
- Papilio lamarchei Staudinger, 1892
- Papilio torquatus Cramer, 1777
- Unnamed group:
- Papilio peleides Esper, 1793
- Papilio okinawensis Fruhstorfer, 1898
- Papilio tros Huebner, 1825
Subgenus Pterourus Scopoli, 1777:
-
troilus group:
- Papilio palamedes Drury, 1773
- Papilio troilus Linnaeus, 1758
-
glaucus group:
- Papilio appalachiensis Pavulaan & Wright, 2002
- Papilio canadensis Rothschild & Jordan, 1906
- Papilio esperanza Beutelspacher, 1975
- Papilio eurymedon Lucas, 1852
- Papilio glaucus Linnaeus, 1758
- Papilio multicaudatus Kirby, 1884
- Papilio pilumnus Boisduval, 1836
- Papilio rutulus Lucas, 1852
-
zagreus group:
- Papilio neyi Niepelt, 1909
- Papilio zagreus Doubleday, 1847
- ( Papilio bachus ) C. & R. Felder, 1865
-
scamander group:
- Papilio birchallii Hewitson, 1863
- Papilio hellanichus Hewitson, 1868
- Papilio scamander Boisduval, 1836
- Papilio xanthopleura Godman & Salvin, 1868
-
homerus group:
- Papilio cacicus Lucas, 1852
- Papilio euterpinus Salvin & Godman, 1868
- Papilio garamas Geyer, 1829
- Papilio homerus Fabricius, 1793
- Papilio menatius Huebner, 1819
- Papilio warscewiczii Hopffer, 1865
Subgenus Sinoprinceps Hancock, 1983:
- Papilio benguetanus Joicey & Talbot, 1923
-
xuthus group Hancock, 1983:
- Papilio xuthus Linnaeus, 1767
swell
literature
- James Duncan: British Butterflies . In: Sir William Jardine (Ed.): The Naturalists Library, Entomology . tape III . Edinburgh 1835, p. 92 ff . (English, online in the Google book search).
- James Duncan: Foreign Butterflies . In: Sir William Jardine (Ed.): The Naturalists Library, Entomology . tape V . Edinburgh 1837, p. 93 ff . (English, online in the Google book search).
Individual evidence
- ^ Duncan, 1837.
- ^ Duncan, 1835.
- ↑ Duncan 1937.
- ↑ Papilio. Fauna Europaea, accessed December 8, 2014 .
- ^ EG Munroe: The classification of the Papilionidae (Lepidoptera) . In: Canadian Entomology Supplies . tape 17 , 1961, pp. 1-51 (English).
- ^ DL Hancock: Classification of the Papilionidae (Lepidoptera): A phylogenetic approach . In: Smithersia . tape 2 , 1983, p. 1-48 (English).
- ^ S. Igarashi: The classification of the Papilionidae mainly based on the morphology of their immature stages . In: Tyo to Ga . tape 34 , 1984, pp. 41-96 (English).
- ↑ CL Hauser, R. de Jong, G. Lamas, K. Robbins, C. Smith, RI Vane-Wright: Papilionidae – Revised GloBIS / GART species checklist, 2nd draft . In: Tex. Mem. Mus. tape 29 , 2002, pp. 1–25 (English, online ).
- ^ Evgueni V. Zakharov, Michael S. Caterino, Felix AH S Perling: Molecular Phylogeny, Historical Biogeography, and Divergence Time Estimates for Swallowtail Butterflies of the Genus Papilio (Lepidoptera: Papilionidae) . In: Syst. Biol. Band 53 , no. 2 , 2004, p. 193-215 , doi : 10.1080 / 10635150490423403 (English).
- ↑ nic.funet.fi, 2010.
Web links
- Papilio Linnaeus, 1758. www.nic.funet.fi,accessed on December 17, 2010(English).