Rhynchosia
Rhynchosia | ||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
Rhynchosia acuminatifolia , habit, three-part leaves, inflorescences and young fruits |
||||||||||||
Systematics | ||||||||||||
|
||||||||||||
Scientific name | ||||||||||||
Rhynchosia | ||||||||||||
Lour. |
Rhynchosia is a genus in the subfamily Schmetterlingsblütler (Faboideae) within the family of the Leguminosae (Fabaceae). There are 200 to 300 species in thegenus Rhynchosia ,which is found almost worldwide. It is one of the genera of the Faboideae, the species ofwhich are called beans .
description
Appearance and leaves
Rhynchosia TYPES grow as crawling, climbing or self-erect, perennial herbaceous plants , or rarely woody plants: Half bushes , shrubs or vines . They can be evergreen or deciduous. The climbing species wind their way up counterclockwise.
The arranged and alternate spiral or two lines distributed to the stem axis leaves are divided into petiole and leaf blade. The herbaceous to leathery leaf blades are pinnate unpaired. There are usually three pinna flake present, the leaf blades are therefore usually consist of three or is it just the terminal leaflet available. There is a pulvinus on the leaflets . The flat pinnate leaves are rhombic, egg-shaped, obovate, or circular with a wedge-shaped or rounded base and smooth, flat edge with a length of 1 to 5.5 cm. The underside of the leaflets is usually dotted with glands, with sessile glands. The two lateral leaflets are asymmetrical and the petioles of the terminal leaflets are more than four times as long as those of the lateral ones. Network nerves are present. If there are minor leaflets of the leaflets, they are tiny and bristle-shaped. The two durable or early falling stipules are often linear-lanceolate, free from each other and not fused with the petiole.
Inflorescences and flowers
The flowers are rarely single or usually in several in the lateral, simple or compound racemose inflorescences . The bracts usually fall off early. Cover sheets are missing.
The stalked, hermaphrodite flowers are zygomorphic and five-fold with a double flower envelope . The five unequal sepals are fused bell-shaped; of the five calyx teeth, the lowest one is only very short with the others and the two upper ones are mostly fused over almost their entire length. The corollas are longer than the calyx or short. The five petals can be nailed. The flower crowns have the typical structure of the butterfly flowers. The colors of the petals range from yellow or yellow with red to yellow with brown. The two lower petals have grown together to form a boat, which is curved inward. The two wings are curved inward. The two wings can be fused with the shuttle. The normally developed, circular or obovate flag is not spurred, but can have inwardly curved ears at the base. The ten fertile stamens are not fused with the petals. Of the ten fertile stamens, nine stamens have grown together to form a tube. All dust bags are the same and open with a longitudinal slit. The individual, more or less seated, upper carpels rarely contain one or mostly two ovules . The stylus, which is curved above its center, is usually hairy in the lower area and ends with a small scar. The nectar secretion takes place on the disc . The pollination is effected by insects ( Entomophilie ).
Fruits and seeds
The more or less sedentary legumes are more or less flattened and with a length of 1.6 to 2.4 cm oblong, obscure-lanceolate, obovate-ellipsoid, oblique-circular, sickle-shaped or ellipsoid and are always beaked at the top . The legumes, which are flattened or puffed up, often glandular, are not septate or constricted between the seeds. The fruits that open when ripe only rarely contain one or mostly two seeds.
The seeds are usually almost circular in outline or kidney-shaped. The hilum is round and the seed stalk ( funiculus ) is in the middle. If there is a stanza it is relatively small. The unit of dissemination ( diaspore ) is the seed.
Systematics and distribution
The genus Rhynchosia was first published in 1790 by João de Loureiro as Canavali in Flora Cochinchinensis , 2, p. 425, 460. The type species is Rhynchosia volubilis Lour. According to the rules of the ICBN (Vienna ICBN Art. 14.4 & App. III), the generic name Rhynchosia is conserved compared to Dolicholus Medik. nom. rej. and Cylista Aiton nom. rej. Other synonyms for Rhynchosia Lour. nom. cons. are: Baukea Vatke , Copisma E. Mey . , Leycephyllum Piper , Nomismia Wight & Arn. , Orthodanum E. Mey . , Pitcheria Nutt.
The genus Rhynchosia belongs to the subtribe Cajaninae of the tribe Phaseoleae in the subfamily Faboideae within the family of the Fabaceae .
The genus Rhynchosia has an almost worldwide distribution; there are species in the tropics and subtropics .
There are 200 to 300 species of Rhynchosia :
- Rhynchosia acuminatifolia Makino
- Rhynchosia acuminatissima Miq.
- Rhynchosia acutissima Thwaites
- Rhynchosia adenodes Eckl. & Zeyh.
- Rhynchosia affinis De Wild.
- Rhynchosia albae-pauli Berhaut
- Rhynchosia albissima Gand.
- Rhynchosia alluaudii Sacleux
- Rhynchosia amabilis Grear
- Rhynchosia ambacensis (Hiern) K.Schum.
- Rhynchosia americana (Mill.) Metz
- Rhynchosia androyensis Du Puy & Labat
- Rhynchosia angulosa Schinz
- Rhynchosia angustifolia DC.
- Rhynchosia arenicola Hassl.
- Rhynchosia argentea Harv.
- Rhynchosia arida CHStirt.
- Rhynchosia aurea (Willd.) DC.
- Rhynchosia avensis Baker
- Rhynchosia axilliflora Hauman
- Rhynchosia bakeri Schinz
- Rhynchosia balansae Micheli
- Rhynchosia barbertonensis C.H. Stirt .
- Rhynchosia baukea Du Puy & Labat
- Rhynchosia baumii Harms
- Rhynchosia beddomei Baker
- Rhynchosia biballensis Torre
- Rhynchosia bracteata Baker
- Rhynchosia braunii Harms
- Rhynchosia brunnea Baker f.
- Rhynchosia buchananii Harms
- Rhynchosia buettneri Harms
- Rhynchosia bullata Harv.
- Rhynchosia burkartii Fortunato
- Rhynchosia burkei Burtt Davy & Baker f.
- Rhynchosia caaguazuensis Hassl.
- Rhynchosia calcicola Craib
- Rhynchosia calobotrya Harms
- Rhynchosia calosperma Warb.
- Rhynchosia calvescens Meikle
- Rhynchosia calycina E. Mey.
- Rhynchosia calycosa Hemsl.
- Rhynchosia cana (Willd.) DC.
- Rhynchosia candida (Hiern) Torre
- Rhynchosia capensis (Burm.f.) Schinz
- Rhynchosia capitata (Roth) DC.
- Rhynchosia caribaea (Jacq.) DC.
- Rhynchosia castroi Baker f.
- Rhynchosia chevalieri Harms
- Rhynchosia chinensis Y.T.Wei & SKLee
- Rhynchosia chrysantha numberbr.
- Rhynchosia chrysoscias Harv. & Sond.
- Rhynchosia ciliata (Thunb.) Druce
- Rhynchosia cinerea Nash
- Rhynchosia clausseni Benth.
- Rhynchosia cliffordii Hutch. & EABruce
- Rhynchosia clivorum S. Moore
- Rhynchosia confusa Burtt Davy
- Rhynchosia congensis Baker
- Rhynchosia connata Baker f.
- Rhynchosia cooperi (Baker f.) Burtt Davy
- Rhynchosia corylifolia Benth.
- Rhynchosia courtallensis Maesen
- Rhynchosia crassifolia Harv.
- Rhynchosia crispa Verdc.
- Rhynchosia cytisoides (Bertol.) Wilbur
- Rhynchosia dekindtii Harms
- Rhynchosia delicatula O.Téllez & M.Sousa
- Rhynchosia densiflora (Roth) DC.
- Rhynchosia dielsii Harms
- Rhynchosia dieterlenae Baker f.
- Rhynchosia difformis (Elliott) DC.
- Rhynchosia discolor M. Martens & Galeotti
- Rhynchosia distans Craib
- Rhynchosia divaricata Baker
- Rhynchosia diversifolia Micheli
- Rhynchosia edulis Griseb.
- Rhynchosia elegans A.Rich.
- Rhynchosia emarginata Germish.
- Rhynchosia erecta Thunb.
- Rhynchosia Erlangeri Harms
- Rhynchosia erythraeae Schweinf.
- Rhynchosia erythrinoides Schltdl. & Cham.
- Rhynchosia exellii Torre
- Rhynchosia falconeri Baker
- Rhynchosia ferruginea A.Rich.
- Rhynchosia ferulifolia Harv.
- Rhynchosia filipes Benth.
- Rhynchosia fischeri P.Satyanar. & Thoth.
- Rhynchosia flava (Forssk.) Thulin
- Rhynchosia fleckii Schinz
- Rhynchosia foliosa marrow viper
- Rhynchosia galpinii Baker f.
- Rhynchosia gandensis Torre
- Rhynchosia gansole Chiov.
- Rhynchosia genistoides Burtt Davy
- Rhynchosia goetzei Harms
- Rhynchosia gossweileri Baker f.
- Rhynchosia grandiflora Steud.
- Rhynchosia hainesiana P. Satyanar. & Thoth.
- Rhynchosia harae H.Ohashi & Tateishi
- Rhynchosia harmsiana Zahlbr.
- Rhynchosia harveyi Eckl. & Zeyh.
- Rhynchosia hauthalii Kuntze
- Rhynchosia heterophylla Hauman
- Rhynchosia heynei Wight & Arn.
- Rhynchosia himalensis Baker
- Rhynchosia hirsuta Eckl. & Zeyh.
- Rhynchosia hirta (Andrews) Meikle & Verdc.
- Rhynchosia holosericea Schinz
- Rhynchosia holstii Harms
- Rhynchosia holtzii Harms
- Rhynchosia huillensis (Hiern) K.Schum.
- Rhynchosia insignis (O.Hoffm.) REFr.
- Rhynchosia jacobii Chandrab. & BVShetty
- Rhynchosia jacottetii Schinz
- Rhynchosia kilimanjarica Harms
- Rhynchosia komatiensis Harms
- Rhynchosia kunmingensis Y.T.Wei & SKLee
- Rhynchosia laetissima Baker
- Rhynchosia lateritia Burkart
- Rhynchosia latifolia Torr. & A.Gray
- Rhynchosia leandrii Du Puy & Labat
- Rhynchosia ledermannii Harms
- Rhynchosia leucophylla (Benth.) Benth.
- Rhynchosia leucoscias Harv.
- Rhynchosia lineata Benth.
- Rhynchosia longeracemosa M. Martens & Galeotti
- Rhynchosia longiflora Schinz
- Rhynchosia longipetiolata Hosseus
- Rhynchosia longissima Hauman
- Rhynchosia lukafuensis Baker f.
- Rhynchosia lutea Dunn
- Rhynchosia luteola (Hiern) K.Schum.
- Rhynchosia macrantha Hauman
- Rhynchosia macrocarpa Benth.
- Rhynchosia madagascariensis R.Vig.
- Rhynchosia malacophylla (Spreng.) Bojer
- Rhynchosia malacotricha Harms
- Rhynchosia mannii Baker
- Rhynchosia mantaroensis J.F. Macbr .
- Rhynchosia marcanii Craib
- Rhynchosia meeboldii P. Satyanar. & Thoth.
- Rhynchosia megalocalyx Thulin
- Rhynchosia melanocarpa Grear
- Rhynchosia mensensis Schweinf.
- Rhynchosia michauxii Vail
- Rhynchosia micrantha Harms
- Rhynchosia microscias Harv.
- Rhynchosia minima (L.) DC.
- Rhynchosia mollis Burtt Davy
- Rhynchosia monophylla Schltr.
- Rhynchosia monticola Grear
- Rhynchosia naineckensis Fortunato
- Rhynchosia namaensis Schinz
- Rhynchosia nelsonii (Rose) Grear
- Rhynchosia nepalensis H.Ohashi & Tateishi
- Rhynchosia nervosa Harv.
- Rhynchosia nipensis Urb.
- Rhynchosia nitens Harv.
- Rhynchosia nitida Harv.
- Rhynchosia nummularia (L.) DC.
- Rhynchosia nyasica Baker
- Rhynchosia nyikensis Baker
- Rhynchosia oblatifoliolata Verdc.
- Rhynchosia oblongifoliolata Hauman
- Rhynchosia orthobotrya Harms
- Rhynchosia ovata J.M.Wood & MSEvans
- Rhynchosia ovatifoliolata Torre
- Rhynchosia pallida Micheli
- Rhynchosia parviflora E. Mey.
- Rhynchosia parvifolia DC.
- Rhynchosia pauciflora Bolus
- Rhynchosia peglerae Baker f.
- Rhynchosia pentheri numberbr.
- Rhynchosia phaseoloides (Sw.) DC.
- Rhynchosia picta (E. Mey.) Burtt Davy
- Rhynchosia pinnata Harv.
- Rhynchosia platyphylla Benth.
- Rhynchosia potosina Brandegee
- Rhynchosia precatoria (Willd.) DC.
- Rhynchosia preussii (Harms) Harms
- Rhynchosia pringlei rose
- Rhynchosia procurrens (Hiern) K.Schum.
- Rhynchosia prostrata Brandegee
- Rhynchosia pseudo-cajan Cambess.
- Rhynchosia pseudoteramnoides Hauman
- Rhynchosia pseudoviscosa Harms
- Rhynchosia pubescens DC.
- Rhynchosia pulchra (Vatke) Harms
- Rhynchosia pulverulenta stocks
- Rhynchosia pycnostachya (DC.) Meikle
- Rhynchosia pyramidalis (Lam.) Urb.
- Rhynchosia quadrata Harv.
- Rhynchosia quercetorum Standl.
- Rhynchosia ramosa Verdc.
- Rhynchosia reniformis (Pursh) DC.
- Rhynchosia reptabunda N.E.Br.
- Rhynchosia resinosa (A.Rich.) Baker
- Rhynchosia reticulata (Sw.) DC.
- Rhynchosia rhomboidea Benth.
- Rhynchosia rojasii Hassl.
- Rhynchosia rostrata Benth.
- Rhynchosia rothii Aitch.
- Rhynchosia rotundifolia Walp.
- Rhynchosia rudolfii Harms
- Rhynchosia rufescens (Willd.) DC.
- Rhynchosia salicifolia Hauman
- Rhynchosia schimperi Boiss.
- Rhynchosia schlechteri Baker f.
- Rhynchosia schomburgkii Benth.
- Rhynchosia scutulaefolia Baker f.
- Rhynchosia secunda Eckl. & Zeyh.
- Rhynchosia senna Hook.
- Rhynchosia sordida (E. Mey.) Schinz
- Rhynchosia speciosa Verdc.
- Rhynchosia spectabilis Schinz
- Rhynchosia splendens Schweinf.
- Rhynchosia stenodon Baker f.
- Rhynchosia stenophylla Urb. & Ekman
- Rhynchosia stipata Meikle
- Rhynchosia suaveolens (Lf) DC.
- Rhynchosia sublobata (Schum.) Meikle
- Rhynchosia swartzii (Vail) Urb.
- Rhynchosia tamaulipensis Grear
- Rhynchosia tarphantha Standl.
- Rhynchosia teixeirae Torre
- Rhynchosia teramnoides Harms
- Rhynchosia thorncroftii (Baker f.) Burtt Davy
- Rhynchosia tomentosa (L.) Hook. & Arn.
- Rhynchosia totta (Thunb.) DC.
- Rhynchosia tricuspidata Baker f.
- Rhynchosia usambarensis Deaf.
- Rhynchosia velutina Wight & Arn.
- Rhynchosia vendae C.H. Stirt.
- Rhynchosia verdcourtii Thulin
- Rhynchosia versicolor Baker
- Rhynchosia villosa (Meissner) Druce
- Rhynchosia villosula Burtt Davy
- Rhynchosia viscidula Steud.
- Rhynchosia viscosa (Roth) DC.
- Rhynchosia volubilis Lour.
- Rhynchosia wellmaniana Harms
- Rhynchosia woodii Schinz
- Rhynchosia yucatanensis Grear
- Rhynchosia yunnanensis Franch.
- Rhynchosia zernyi Harms
use
Some species are cultivated for the production of food. For example, plant parts of Rhynchosia diversifolia , Rhynchosia malacophylla , Rhynchosia minima , Rhynchosia memnonia , Rhynchosia senna , Rhynchosia sennaarensis , Rhynchosia sublobata are eaten.
Since they fix nitrogen ( Rhizobium root nodules ), they are important for soil improvement.
Some species are called piule in Mexico; its beautiful red and black seeds were probably used as narcotics by the Mexican civilizations . Some Rhynchosia species have tested positive for alkaloid levels, but their effects have not yet been characterized. The seeds of some species are used in folk medicine in some countries.
See also
swell
- Ren Sa, Delin Wu, Dezhao Chen, Dianxiang Zhang , Hang Sun, Puhua Huang, Michael G. Gilbert, Mats Thulin, C. Melanie Wilmot-Dear & Hiroyoshi Ohashi: Phaseoleae : Ren Sa & Michael G. Gilbert: Rhynchosia , S. 223 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven & Deyuan Hong (eds.): Flora of China , Volume 10 - Fabaceae , Science Press and Missouri Botanical Garden Press, Beijing and St. Louis 2010 . ISBN 978-1-930723-91-7 (sections description, distribution, use and systematics)
- Syed Irtifaq Ali: Papilionaceae : Rhynchosia , p. 226 - Online. in the Flora of West Pakistan , Volume 100, Stewart Herbarium, Rawalpindi 1977.
- Leslie Watson: Papilionaceae in the Western Australian Flora , 2008: Rhynchosia - Online. (Sections Description and Distribution)
- C. Gardner & TA James: Rhynchosia in the New South Wales Flora Online . (Sections Description and Distribution)
- G. Lewis et al .: Legumes of the world. (Leg World), 2005, p. 410.
Individual evidence
- ↑ a b c d e Ren Sa & Michael G. Gilbert: p. 223 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven & Deyuan Hong (eds.): Flora of China , Volume 10 - Fabaceae , Science Press and Missouri Botanical Garden Press, Beijing and St. Louis 2010. ISBN 978-1-930723-91-7
- ↑ a b c d e f Leslie Watson: Papilionaceae in the Western Australian Flora , 2008: Rhynchosia - Online.
- ^ A b c d C. Gardner & TA James: Rhynchosia in the New South Wales Flora Online .
- ↑ First publication scanned at biodiversitylibrary.org .
- ↑ a b Rhynchosia at Tropicos.org. Missouri Botanical Garden, St. Louis, accessed June 29, 2013.
- ↑ a b c d Rhynchosia in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland. Retrieved June 29, 2013.
- ↑ Enter the taxon in the search mask at ILDIS = International Legume Database & Information Service - World Database of Legumes .
- ↑ Some Rhynchosia species at the FAO .
- ↑ data sheet at erowid.org .
Web links
- Entry in the Flora of Zimbabwe . (engl.)