Myrobalans
Myrobalans | ||||||||||||
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Behara tree ( Terminalia bellirica ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Terminalia | ||||||||||||
L. |
The myrobalans ( Terminalia ) are a genus of plants within the winged family (Combretaceae). The 100 to 200 species are distributed almost worldwide in the tropics.
description
Vegetative characteristics
The Terminalia species grow as trees or shrubs , sometimes climbing. Some species are armed. Most alternate and spiral, sometimes almost rarely arranged completely opposite or in whorls leaves are simple and entire. Stipules are missing.
Generative characteristics
The flowers are in terminal or axillary, paniculate , racemose or spiked inflorescences together with bracts . The radially symmetrical or zygomorphic , four or five-fold flowers are mostly hermaphroditic, rarely unisexual. The flower cup (hypanthium) is fused with the ovary. The four or five sepals are fused, rarely eight sepals are present. There are four or five petals or they are absent. There is a discus. There are one or two circles with four or five, rarely more, stamens each. There is only one carpel below with two to six ovules . The simple, thin style ends in a pointed, heady or undefined scar.
Mostly fleshy, stone fruit-like, leathery, two- to five-edged or winged fruits are formed. They contain only one endospermless seed. The oily cotyledons ( cotyledons ) are curled or spirally rolled.
Systematics
Taxonomy
The genus Terminalia was established in 1767 by Carl von Linné in Systema Naturae , 12th edition, 2, pages 665 and 674. The type species is Terminalia catappa L. The generic name Terminalia is derived from the Latin word terminalis = end, tip, border and refers to the accumulation of leaves at the branch ends. Synonyms for Terminalia L. nom. cons. are: Badamia Gaertn. , Catappa Gaertn. , Chicharronia A.Rich. , Chuncoa Pav. Ex Juss. , Fatrea Juss. , Gimbernatea Ruiz & Pav. , Hudsonia A.Rob. ex Lunan , Kniphofia Scop. , Myrobalanifera Houtt. , Myrobalanus Gaertn. , Pentaptera Roxb. , Ramatuela Kunth , Ramatuella Kunth, orth. Var. , Resinaria Comm. ex Lam. , Tanibouca Aubl. , Vicentia Allemão .
External system
The genus Terminalia belongs to subtribe Terminaliinae the tribe Combreteae in the subfamily Combretoideae within the family of Combretaceae .
Internal system
In the genus Terminalia 100 to 200 species are distinguished depending on the author. A distinction is made between several sub-genera; here is a breakdown with a partial list of species from the flora brasiliensis :
- Subgenus Catappa :
- Terminalia australis Cambess.
- Indian myrobalans , katappenbaum ( Terminalia catappa L. )
- Terminalia lucida Hoffmanns.
- Terminalia tanibouca Sm.
- Subgenus Chuncoa :
- Terminalia actinophylla Mart.
- Terminalia brasiliensis Cambess.
- Terminalia glabrescens Mart.
- Terminalia oblonga Poir.
- Terminalia obovata Poir.
- Subgenus Diptera :
- Terminalia argentea Mart. & Zucc.
- Terminalia biscutella Eichler
- Terminalia fagifolia Mart. & Zucc.
- Terminalia guyanensis Eichler
- Terminalia januariensis DC.
- Terminalia macroptera Mart.
- Terminalia modesta Eichler
- Terminalia phaeocarpa Eichler
- Terminalia punctata Eichler
- Subgenus Vicentia :
- Terminalia acuminata Allemão
- Terminalia adamantium Cambess.
- Terminalia riedelii Eichler
Here is a selection of species without considering the sub-genera:
- Terminalia amazonia (JFGmel.) Exell , Syn .: Chuncoa amazonia J.F.Gmel. , Terminalia hayesii Pittier : It is distributed from Mexico through Central America to northern and western South America .
- Terminalia arenicola Byrnes : It occurs only in the Australian state of Queensland .
- Terminalia argentea Mart. : It occurs in Bolivia , Ecuador , Brazil and Paraguay .
- Real myrobalans ( Terminalia arjuna (Roxb. Ex DC.) Wight & Arn. , Syn .: Pentaptera arjuna Roxb. Ex DC. , Pentaptera glabra Roxb. )
- Belerian Myrobalan, Bahera tree ( Terminalia bellirica (Gaertn.) Roxb. , Syn .: Myrobalanus bellirica Gaertn. ): It occurs in tropical Asia and in Yunnan .
- Terminalia bentzoe (L.) Pers. , Syn .: Croton bentzoe L. , Terminalia angustifolia Jacq. , Terminalia benzoin L. f. : It occurs on Mauritius and Réunion .
- Terminalia bialata (Roxb.) Steud. , Syn .: Pentaptera bialata Roxb. : It occurs in the Philippines , the Andaman and Nicobar Islands .
- Terminalia brassii Exell : It occurs in Papua New Guinea and the Solomon Islands .
- Terminalia brownii Fresen. : It occurs in Africa and Yemen .
- Terminalia calamansanai (Blanco) Rolfe , Syn .: Gimbernatea calamansanai Blanco : It occurs in Indochina , in Malesia and in Papua New Guinea.
- Indian myrobalans, Katappenbaum ( Terminalia catappa L. , Syn .: Phytolacca javanica Osbeck )
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Chebulic myrobalans ( Terminalia chebula Retz. ): It occurs in Sri Lanka , India , Bangladesh , Nepal , Bhutan , Myanmar , Cambodia , Laos , Vietnam and in Yunnan. There are about two varieties:
- Terminalia chebula Retz. var. chebula
- Chebulic myrobalan ( Terminalia chebula var. Tomentella (short) CBClarke , Syn .: Terminalia tomentella short )
- Terminalia citrina (Gaertn.) Roxb. ex Fleming , Syn .: Embryogonia arborea Teijsm. & Binn. , Myrobalanus citrina Gaertn. , Terminalia arborea (Teijsm. & Binn.) Coord. & Valeton : It occurs in India, Bhutan, Myanmar, Thailand , Indonesia , Malaysia and the Philippines.
- Terminalia elliptica Willd. , Syn .: Pentaptera tomentosa Roxb. ex DC. , Terminalia alata B. Heyne ex Roth , Terminalia tomentosa (Roxb. Ex DC.) Wight & Arn. : It occurs in India, Bangladesh, Myanmar, Thailand, Cambodia, Laos and Vietnam.
- Terminalia ferdinandiana Exell , Syn .: Terminalia latipes Benth. subsp. psilocarpa Pedley : It occurs in Australia.
- Terminalia foetidissima handle. , Syn. Terminalia ovocarpa Merr. : It occurs in Thailand, Myanmar, Malaysia, Indonesia and the Philippines.
- Terminalia franchetii Gagnep. , Syn. Terminalia triptera Franch.
- Terminalia ivorensis A. Chev. : It occurs in Cameroon and in tropical West Africa .
- Terminalia kaernbachii Warb. , Syn .: Terminalia okari C.T.White : It occurs in Indonesia, Papua New Guinea and the Solomon Islands .
- Terminalia latipes Benth. : It occurs in Australia.
- Terminalia macroptera Guill. & Perr. : It occurs in tropical Africa and is also given by the Cote d'Or in France.
- Terminalia microcarpa Decne. : Syn. Terminalia edulis Blanco : It occurs in Indonesia, Malaysia, Papua New Guinea and the Philippines.
- Terminalia muelleri Benth. : It occurs in Queensland.
- Terminalia myriocarpa Van Heurck & Müll. Arg . : It occurs in tropical Asia and in China.
- Terminalia nitens C. Presl : It occurs in the Philippines and Japan .
- Terminalia oblonga (Ruiz & Pav.) Steud. , Syn .: Gimbernatea oblonga Ruiz & Pav .: It is distributed from Mexico via Central America to northern and western South America.
- Terminalia oblongata F. Muell. : It occurs in Australia.
- Terminalia oliveri Brandis : It occurs in Myanmar.
- Terminalia paniculata Roth : It occurs in India.
- Terminalia pellucida C. Presl : It occurs in the Philippines.
- Terminalia phanerophlebia Engl. & Diels : It occurs in Mozambique and southern Africa .
- Terminalia procera Roxb. , Syn .: Terminalia copelandii Elmer : It occurs on the Andaman and Nicobar Islands, in Indonesia, Papua New Guinea and the Philippines.
- Terminalia prunioides M.A.Lawson : It occurs in tropical and southern Africa.
- Terminalia richii A.Gray : It occurs on islands in the southwestern Pacific.
- Terminalia saffordii Merr. : It occurs in Guam and Micronesia .
- Terminalia sericea Burch. ex DC. : It occurs in tropical and southern Africa.
- White myrobalans, limba tree ( Terminalia superba Engl. & Diels )
- Terminalia triflora (Griseb.) Lillo , Syn .: Chuncoa triflora Griseb. : It occurs in Bolivia, Brazil, Argentina and Paraguay .
- Terminalia trifoliata Spreng. : It occurs in Brazil.
- Terminalia tripteroides Craib : It occurs in Thailand.
The following described species are assigned to other genera:
- Terminalia erythrophylla Burch. → Combretum erythrophyllum (Burch.) Sond.
- Terminalia molinetii M.Gómez → Bucida molinetii (M.Gómez) Alwan & Stace
use
Several types of Terminalia are used to extract substances such as cyclic triterpenes , flavonoids and tannins . Some of these substances are effective agents against bacteria and fungi ; cancer-fighting agents are also included.
Terminalia citrina and Terminali chebula as well as Terminalia bellirica were mainly used medicinally from antiquity to the early modern period .
photos
Indian myrobalans ( Terminalia catappa ):
Trunk and bark
swell
- Saida Qaiser, Mohammad Qaiser: Combretaceae. : Terminalia at Tropicos.org. In: Flora of Pakistan . Missouri Botanical Garden, St. Louis
- Jie Chen, Nicholas J. Turland: Combretaceae. : Terminalia , p. 310 - online with the same text as the printed work , In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Ed.): Flora of China. Volume 13: Clusiaceae through Araliaceae. Science Press and Missouri Botanical Garden Press, Beijing and St. Louis, 2007, ISBN 978-1-930723-59-7 .
Individual evidence
- ↑ a b Terminalia at Tropicos.org. Missouri Botanical Garden, St. Louis, Retrieved May 27, 2017.
- ↑ GENAUST: etymological dictionary of the botanical name of the plant . Springer-Verlag, 2013, ISBN 3-0348-7650-5 ( Terminalia on p. 361 in the Google book search).
- ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an Terminalia in the Germplasm Resources Information Network (GRIN) , USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland. Retrieved May 26, 2017.
- ↑ List of species in Flora brasiliensis .
- ↑ Werner Dressendörfer: Late medieval medicine taxes of the Munich city doctor Sigmund Gotzkircher from the Grazer Codex 311. A contribution to the early history of the southern German pharmacy. Königshausen and Neumann Würzburg 1978 (= Würzburg medical-historical research , 15), p. 240 f.
Web links
- Terminalia . In: U. Brunken, M. Schmidt, S. Dressler, T. Janssen, A. Thiombiano, G. Zizka: West African plants - A Photo Guide. Senckenberg Research Institute, Frankfurt am Main 2008.
- Terminalia at Tropicos.org. In: IPCN Chromosome Reports . Missouri Botanical Garden, St. Louis
- Terminalia at Tropicos.org. In: Flora Mesoamericana . Missouri Botanical Garden, St. Louis
- Terminalia at Tropicos.org. In: Catalog of the Vascular Plants of Madagascar . Missouri Botanical Garden, St. Louis
- Terminalia at Tropicos.org. In: Flora de Nicaragua . Missouri Botanical Garden, St. Louis
- Terminalia at Tropicos.org. In: Flora of Panama (WFO) . Missouri Botanical Garden, St. Louis
- Terminalia at Tropicos.org. In: 66 . Missouri Botanical Garden, St. Louis