Systematics of the ferns

The system of ferns follows Smith et al. (2006), who created a classical taxonomy of the recent ferns on a phylogenetic basis.

According to the International Code of Botanical Nomenclature , the scientific names have endings that are indicative of the rank of the corresponding taxa. The names of the families end with "-aceae", those of the orders with "-ales", the subclasses with "-idae" and the classes with "-opsida".

Smith et al. (2006) divide the ferns into four classes.

• Class Psilotopsida
  • Order adder tongue-like (Ophioglossales)
    • Family adder tongue plants (Ophioglossaceae) (incl.Botrychiaceae, Helminthostachyaceae)
      • Botrychium , Helminthostachys , Ophioglossum , Mankyua .
  • Order fork-like (Psilotales)
    • Fork leaf family (Psilotaceae) (including Tmesipteridaceae)
      • Psilotum , Tmesipteris .

• Class Equisetopsida [= Sphenopsida]
  • Order horsetail (Equisetales)
    • Horsetail family (Equisetaceae)
      • Equisetum

• Class Marattiopsida
  • Order marattiales
    • Family Marattiaceae (including Angiopteridaceae, Christenseniaceae, Danaeaceae, Kaulfussiaceae)
      • Angiopteris , Christensenia , Danaea , Marattia .

• Class real ferns (Polypodiopsida) [= Filicopsida, Polypodiopsida]
  • Order royal ferns (Osmundales)
    • Family OSMUNDACEAE (Osmundaceae)
      • Leptopteris , Osmunda , Todea .
  • Order skin ferns (Hymenophyllales)
    • Skin fern family (Hymenophyllaceae) (including Trichomanaceae )
      • Hymenophyllum , Didymoglossum , Crepidomanes , Polyphlebium , Vandenboschia , Abrodictyum , Trichomanes , Cephalomanes , Callistopteris .
  • Order equal
    • Family Gleicheniaceae (including Dicranopteridaceae , Stromatopteridaceae )
      • Dicranopteris , Diplopterygium , Gleichenella , Gleichenia , Sticherus , Stromatopteris .
    • Dipteridaceae family (including Cheiropleuriaceae )
      • Cheiropleuria , Dipteris .
    • Family Matoniaceae
      • Matonia, Phanerosorus.
  • Order Schizaeales
    • Lygodiaceae family
      • Lygodium .
    • Family Anemiaceae (including Mohriaceae )
      • Anemia
    • Schizaeaceae family
      • Actinostachys , Schizaea .
  • Order floating ferns (Salviniales)
    • Family clover family (Marsileaceae) ( incl.Pilulariaceae )
      • Marsilea , Pilularia , Regnellidium .
    • Swimming fern family (Salviniaceae) (including Azollaceae )
      • Salvinia , Azolla .
  • Order tree ferns (Cyatheales)
    • Thyrsopteridaceae family
      • Thyrsopteris ( Thyrsopteris elegans ).
    • Loxomataceae family
      • Loxoma , Loxsomopsis .
    • Family Culcitaceae
      • Culcita .
    • Plagiogyriaceae family
      • Plagiogyria .
    • Cibotiaceae family
      • Cibotium .
    • Family Cyatheaceae (including Alsophilaceae , Hymenophyllopsidaceae )
      • Alsophila (including Nephelea ), Cyathea (including Cnemidaria , Hemitelia , Trichipteris ), Gymnosphaera , Hymenophyllopsis , Sphaeropteris (including Fourniera ).
    • Family Dicksoniaceae (including Lophosoriaceae )
      • Calochlaena , Dicksonia , Lophosoria .
    • Metaxyaceae family
      • Metaxya .
  • Order spotted ferns (Polypodiales)
    • Family Lindsaeaceae ( incl.Cystodiaceae , Lonchitidaceae )
      • Cystodium , Lindsaea , Lonchitis , Odontosoria , Ormoloma , Sphenomeris , Tapeinidium , Xyropteris .
    • Saccolomataceae family
      • Saccoloma (including Orthiopteris )
    • Family dennstaedtiaceae (Dennstaedtiaceae) (incl. Hypolepidaceae , Monachosoraceae , Pteridiaceae )
      • Blotiella , Coptodipteris , Dennstaedtia (incl. Costaricia1 ), Histiopteris, Hypolepis , Leptolepia , Microlepia , Monachosorum , Oenotrichia , Paesia , Pteridium .
    • Family pteridaceae (Pteridaceae) (incl. Acrostichaceae , Actiniopteridaceae , Adiantaceae , Anopteraceae , Antrophyaceae , Ceratopteridaceae , Cheilanthaceae , Cryptogrammaceae , Hemionitidaceae , Negripteridaceae , Parkeriaceae , Platyzomataceae , Sinopteridaceae , Taenitidaceae , Vittariaceae )
      • Acrostichum , Actiniopteris , Adiantopsis , Adiantum , Aleuritopteris , Ananthacorus , Anetium , Anogramma , Antrophyum , Argyrochosma , Aspidotis , Astrolepis , Austro frames , Bommeria , Cassebeera , Ceratopteris , Cerosora , Cheilanthes , Cheiloplecton , Coniogramme , Cosentinia , Cryptogramma , Doryopteris , Eriosorus , Haplopteris , Hecistopteris , Hemionitis , Holcochlaena , Jamesonia , llavea , Mild Ella , monogramma , Nephopteris , Neuro Callis , Notholaena , Ochropteris , Onychium , Paraceterach , Parahemionitis , Pellaea , Pentagramma , Pityrogramma , Platyloma , Platyzoma , Polytaenium , Pteris (incl. Afropteris , Anopteris ) Pterozonium , Radiovittaria , Rheopteris , Scoliosorus , Syngramma , Taenitis , Trachypteris , Vittaria .
    • Striped fern family (Aspleniaceae)
      • 1 - 10 genera: Asplenium , Camptosorus , Loxoscaphe , Diellia , Pleurosorus , Phyllitis , Ceterach , Thamnopteris , Antigramma , Holodictyum , Schaffneria , Sinephropteris , Hymenasplenium .
    • Marsh fern family (Thelypteridaceae)
      • 5-30 genera: Cyclosorus (incl. Ampelopteris , Amphineuron , Chingia , Christella , Cyclogramma , Cyclosorus s s.. Glaphyropteridopsis , Goniopteris , Meniscium , Menisorus , Mesophlebion , Pelazoneuron , Plesioneuron , Pneumatopteris , Pronephrium , Pseudocyclosorus , Sphaerostephanos , Stegnogramma , Steiropteris , Trigonospora ) , Macrothelypteris , Phegopteris , Pseudophegopteris , Thelypteris (including Amauropelta , Coryphopteris , Metathelypteris , Oreopteris , Parathelypteris , Thelypteris see str.)
    • Eyelash family (Woodsiaceae) ( incl.Athyriaceae , Cystopteridaceae )
      • Athyrium , Diplazium (incl. Callipteris , Monomelangium ) Acystopteris , Cheilanthopsis , Cornopteris , Cystopteris , Deparia (incl. Lunathyrium , Dryoathyrium , Athyriopsis , Dictyodroma ) Diplaziopsis , Gymnocarpium (incl. Currania ) Hemidictyum , Homalosorus , Protowoodsia , Pseudocystopteris , Rhachidosorus , Woodsia (including Hymenocystis )
    • Rib fern family (Blechnaceae) ( incl.Stenochlaenaceae )
      • Blechnum s. l., Brainea , Doodia , Pteridoblechnum , Sadleria , Salpichlaena , Steenisioblechnum , Stenochlaena , Woodwardia .
    • Pearl fern family (Onocleaceae)
      • Matteuccia , Onoclea , Onocleopsis , Pentarhizidium .
    • Fern family (Dryopteridaceae) ( incl.Aspidiaceae , Bolbitidaceae , Elaphoglossaceae , Hypodematiaceae , Peranemataceae )
      • Acrophorus , Acrorumohra , Adenoderris , Arachniodes , Ataxipteris , Bolbitis (incl. Egenolfia ) Coveniella , Ctenitis , Cyclodium , Cyrtogonellum , Cyrtomidictyum , Cyrtomium , Didymochlaena , Dryopolystichum , Dryopsis , Dryopteris (incl. Nothoperanema ) Elaphoglossum (incl. Microstaphyla ) Hypodematium , Lastreopsis , Leucostegia , Lithostegia , Lomagramma , Maxonia , Megalastrum , Oenotrichia pp, Olfersia , Peranema , Phanerophlebia , Polystichum (incl. Papuapteris , Plecosorus , Polybotrya , Polystichopsis , Revwattsia , Rumohra , Stenolepia , Stigmatopteris , Teratophyllum ).
    • Family Lomariopsidaceae (including Nephrolepidaceae )
      • Cyclopeltis, Lomariopsis, Nephrolepis , Thysanosoria .
    • Family Tectariaceae
      • Aenigmopteris , Arthropteris , Heterogonium , Hypoderris , Pleocnemia , Psammiosorus , Psomiocarpa , Pteridrys , Tectaria s. l. (including Amphiblestra , Camptodium , Chlamydograms , Cionidium , Ctenitopsis , Dictyoxiphium , Fadyenia , Hemigramma , Pleuroderris , Pseudotectaria , Quercifilix ), Triplophyllum .
    • Oleandraceae family
      • Oleandra .
    • Davalliaceae family
      • Araiostegia, Davallia (including Humata , Parasorus , Scyphularia ), Davallodes , Pachypleuria .
    • Family polypodiaceae (Polypodiaceae) (incl. Drynariaceae , Grammitidaceae , Gymnogrammitidaceae , Loxogrammaceae , Platyceriaceae , Pleurisoriopsidaceae )
      • Acrosorus , Adenophorus , Aglaomorpha (incl. Photinopteris , Merinthosorus , Pseudodrynaria , Holostachyum ) Arthromeris , Belvisia , Calymmodon , Campyloneurum , Ceradenia , Christiopteris , Chrysogrammitis , cochlidium , Colysis , Ctenopteris , Dicranoglossum , Dictymia , Drynaria , Enterosora , Goniophlebium s. l., Grammitis , Lecanopteris , Lellingeria , Lemmaphyllum , Lepisorus (incl. Platygyria ) Leptochilus , Loxogramme (incl. Anarthropteris ), Melpomene , Microgramma (incl. Solanopteris ) Micropolypodium , Microsorum , Neocheiropteris (incl. Neolepisorus ) Neurodium , Niphidium , Pecluma , Phlebodium , Phymatosorus , Platycerium , Pleopeltis , Polypodioides , Polypodium , Prosaptia , Pyrrosia (incl. Drymoglossum ) Scleroglossum , Selliguea (incl. Crypsinus , Polypodiopteris ) Serpocaulon , Synammia , Terpsichore , Themelium , Thylacopteris , Zygophlebia . Monotypic genera: Caobangia , Drymotaenium , Gymnogrammitis , Kontumia , Luisma , Pleurosoriopsis , Podosorus .

literature

• Alan R. Smith, Kathleen M. Pryer, Eric Schuettpelz, Petra Korall, Harald Schneider, Paul G. Wolf: A classification for extant ferns. In: Taxon. Volume 55, No. 3, 2006, ISSN  0040-0262 , pp. 705-731, abstract, PDF file .