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→‎Ecology: Chilean hazel (''Gevuina avellana''), despite its name, is not related to this genus.
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|taxon = Corylus
|taxon = Corylus
|authority = [[Carl Linnaeus|L.]]
|authority = [[Carl Linnaeus|L.]]
|type_species = [[Corylus avellana]]
|type_species = ''[[Corylus avellana]]''
|type_species_authority = [[L.]]
|type_species_authority = [[L.]]
|subdivision_ranks = Species
|subdivision_ranks = Species
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[[File:Catkins Corylus avellana-Mont Bart-5124~2015 12 26.JPG|thumb|right|Young male catkins of ''[[Corylus avellana]]'']]
[[File:Catkins Corylus avellana-Mont Bart-5124~2015 12 26.JPG|thumb|right|Young male catkins of ''[[Corylus avellana]]'']]


The '''hasel''' (''Corylus'') is a [[genus]] of [[deciduous]] [[tree]]s and large [[shrub]]s native to the [[temperateness|temperate]] Northern Hemisphere. The genus is usually placed in the [[birch]] family [[Betulaceae]],<ref name=grin>Germplasm Resources Information Network: [http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?2962 ''Corylus''] {{Webarchive|url=https://web.archive.org/web/20090114213945/http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?2962 |date=2009-01-14 }}</ref><ref>{{Cite journal|last1=Chen|first1=Zhi-Duan|last2=Manchester|first2=Steven R|last3=Sun|first3=Hai-Ying|date=August 1999|title=Phylogeny and evolution of the Betulaceae as inferred from DNA sequences, morphology, and paleobotany|url=http://www.amjbot.org/cgi/content/full/86/8/1168?ck=nck#F5|journal=American Journal of Botany|volume=86|issue=8|pages=1168–1181|doi=10.2307/2656981|issn=0002-9122|jstor=2656981|pmid=10449397|doi-access=free}}</ref><ref name=rushforth>Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins {{ISBN|0-00-220013-9}}.</ref><ref name=rhs>Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan {{ISBN|0-333-47494-5}}.</ref> though some botanists split the hazels (with the [[hornbeam]]s and allied genera) into a separate family Corylaceae.<ref>{{Cite book|title=Trees and Shrubs Hardy in the British Isles|last=Bean|first=William Jackson|date=1976|publisher=J. Murray|others=Taylor, George|isbn=0719517907|edition=8th|volume=1|location=London|oclc=103403}}</ref><ref>{{cite journal|last1=Erdogan |first1=V. |last2=Mehlenbacher |first2=S. A. |year=2002 |title=Phylogenetic analysis of hazelnut species (Corylus, Corylacae) based on morphology and phenology |journal=Sist. Bot. Dergisi |volume=9 |pages=83–100}}</ref> The fruit of the hazel is the [[hazelnut]].
'''Hazels''' are plants of the genus '''''Corylus''''' of [[deciduous]] [[tree]]s and large [[shrub]]s native to the [[temperateness|temperate]] Northern Hemisphere. The genus is usually placed in the [[birch]] family [[Betulaceae]],<ref name=grin>Germplasmgobills Information Network: [http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?2962 ''Corylus''] {{Webarchive|url=https://web.archive.org/web/20090114213945/http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?2962 |date=2009-01-14 }}</ref><ref>{{Cite journal|last1=Chen|first1=Zhi-Duan|last2=Manchester|first2=Steven R|last3=Sun|first3=Hai-Ying|date=August 1999|title=Phylogeny and evolution of the Betulaceae as inferred from DNA sequences, morphology, and paleobotany|journal=American Journal of Botany|volume=86|issue=8|pages=1168–1181|doi=10.2307/2656981|issn=0002-9122|jstor=2656981|pmid=10449397|doi-access=free}}</ref><ref name=rushforth>Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins {{ISBN|0-00-220013-9}}.</ref><ref name=rhs>Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan {{ISBN|0-333-47494-5}}.</ref> though some botanists split the hazels (with the [[hornbeam]]s and allied genera) into a separate family Corylaceae.<ref>{{Cite book|title=Trees and Shrubs Hardy in the British Isles|last=Bean|first=William Jackson|date=1976|publisher=J. Murray|others=Taylor, George|isbn=0719517907|edition=8th|volume=1|location=London|oclc=103403}}</ref><ref>{{cite journal|last1=Erdogan |first1=V. |last2=Mehlenbacher |first2=S. A. |year=2002 |title=Phylogenetic analysis of hazelnut species (Corylus, Corylacae) based on morphology and phenology |journal=Sist. Bot. Dergisi |volume=9 |pages=83–100}}</ref> The fruit of the hazel is the [[hazelnut]].


Hazels have simple, not rounded [[leaf|leaves]] with double-serrate margins. The [[flower]]s are produced very early in spring before the leaves, and are [[monoecious]], with single-sex [[catkin]]s. The male catkins are pale yellow and {{Convert|5–12|cm|abbr=off|frac=4}} long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3&nbsp;mm-long [[Carpel|styles]] visible. The fruits are [[Nut (fruit)|nuts]] {{Convert|1–2.5|cm|abbr=on|frac=4}} long and 1–2&nbsp;cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut.<ref name=rushforth/>
Hazels have simple, rounded [[leaf|leaves]] with double-serrate margins. The [[flower]]s are produced very early in spring before the leaves, and are [[monoecious]], with single-sex [[catkin]]s. The male catkins are pale yellow and {{Convert|5–12|cm|abbr=off|frac=4}} long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3&nbsp;mm-long [[Carpel|styles]] visible. The fruits are [[Nut (fruit)|nuts]] {{Convert|1–2.5|cm|abbr=on|frac=4}} long and 1–2&nbsp;cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut.<ref name=rushforth/>


The shape and structure of the [[involucre]], and also hazel the growth habit (whether a tree or a suckering shrub), are important in the identification of the different species of hazel.<ref name=rushforth/>
The shape and structure of the [[involucre]], and also the growth habit (whether a tree or a suckering shrub), are important in the identification of the different species of hazel.<ref name=rushforth/>


The pollen of hazel species, which are often the cause for allergies in late winter or early spring, can be identified under magnification (600×) by their characteristic granular [[exine]]s bearing three conspicuous pores.<ref>{{Cite journal | doi=10.1038/010355b0| title=Pollen-grains in the Air| journal=Nature| volume=10| issue=253| pages=355| year=1874| last1=Airy| first1=Hubert| bibcode=1874Natur..10..355A| s2cid=4077214| url=https://zenodo.org/record/1429213| doi-access=free}}</ref>
The pollen of hazel species, which are often the cause for allergies in late winter or early spring, can be identified under magnification (600×) by their characteristic granular [[exine]]s bearing three conspicuous pores.<ref>{{Cite journal | doi=10.1038/010355b0| title=Pollen-grains in the Air| journal=Nature| volume=10| issue=253| pages=355| year=1874| last1=Airy| first1=Hubert| bibcode=1874Natur..10..355A| s2cid=4077214| url=https://zenodo.org/record/1429213| doi-access=free}}</ref>


==Species==
==Species==
''Corylus'' has around 14–18 species. The circumscription of species in [[eastern Asia]] is disputed, with [[World Flora Online]] <!-- replaced ''WCSP''--> and the ''Flora of China'' differing in which taxa are accepted, within this region. WFO accepts 17 species while Flora of China accepts 20 species (including ''Corylus mandshurica'').
Only those taxa accepted by both sources are listed below.<ref name="WorldFloraOnline">{{cite web |title=''Corylus'' L. |url=https://www.worldfloraonline.org/taxon/wfo-4000009390 |access-date=19 October 2023}}</ref><ref name=foc>{{cite web |title=''Corylus'' in Flora of China @ efloras.org |url=http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=108088 |website=www.efloras.org |access-date=19 October 2023}}</ref><ref name=fna>Flora of North America: [http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=108088 ''Corylus'']</ref>


The species are grouped as follows:
''Corylus'' has 14–18 species. The circumscription of species in [[eastern Asia]] is disputed, with ''WCSP'' and the ''Flora of China'' differing in which taxa are accepted; within this region, only those taxa accepted by both sources are listed below.<ref name=rushforth/><ref name=kew>WCSP: [http://www.kew.org/wcsp/qsearch.do?plantName=Corylus&page=quickSearch ''Corylus'']{{Dead link|date=July 2019 |bot=InternetArchiveBot |fix-attempted=yes }}</ref><ref name=foc>Flora of China: [http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=108088 ''Corylus'']</ref><ref name=fna>Flora of North America: [http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=108088 ''Corylus'']</ref> The species are grouped as follows:
* Nut surrounded by a soft, leafy [[Involucral bract|involucre]], multiple-stemmed, suckering [[shrub]]s to 12 m tall
* Nut surrounded by a soft, leafy [[Involucral bract|involucre]], multiple-stemmed, suckering [[shrub]]s to 12 m tall
** Involucre short, about the same length as the nut
** Involucre short, about the same length as the nut
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*** ''[[Corylus fargesii]]'' – Farges' hazel, western China
*** ''[[Corylus fargesii]]'' – Farges' hazel, western China
*** ''[[Corylus jacquemontii]]'' – Jacquemont's hazel, [[Himalaya]]
*** ''[[Corylus jacquemontii]]'' – Jacquemont's hazel, [[Himalaya]]
*** ''[[Corylus wangii]]'' – Wang's hazel, southwest [[China]]
*** ''[[Corylus wangii]]'' – Wang's hazel, [[southwest China]]
** Involucre densely spiny, resembling a chestnut burr
** Involucre densely spiny, resembling a chestnut burr
*** ''[[Corylus ferox]]'' – Himalayan hazel, Himalaya, [[Tibet]] and southwest China (syn. ''C. tibetica'').
*** ''[[Corylus ferox]]'' – Himalayan hazel, Himalaya, [[Tibet]] and southwest China (syn. ''C. tibetica'').


Several [[Hybrid (biology)|hybrids]] exist, and can occur between species in different sections of the genus, e.g. ''[[Corylus × colurnoides]]'' (''C. avellana'' × ''C. colurna''). The oldest confirmed hazel species is ''[[Corylus johnsonii]]'' found as [[fossil]]s in the [[Ypresian]]-age rocks of [[Ferry County, Washington]].<ref name=Pigg2003>{{cite journal |last1=Pigg |first1=K.B. |author2=Manchester S.R. |author3=Wehr W.C. |year=2003 |title=''Corylus'', ''Carpinus'', and ''Palaeocarpinus'' (Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America |journal= International Journal of Plant Sciences |volume=164 |issue= 5 |pages=807–822 |doi=10.1086/376816|s2cid=19802370 }}</ref>
Several [[Hybrid (biology)|hybrids]] exist, and they can occur between species in different sections of the genus, e.g. ''[[Corylus × colurnoides]]'' (''C. avellana'' × ''C. colurna''). The oldest confirmed hazel species is ''[[Corylus johnsonii]]'' found as [[fossil]]s in the [[Ypresian]]-age rocks of [[Ferry County, Washington]].<ref name=Pigg2003>{{cite journal |last1=Pigg |first1=K.B. |author2=Manchester S.R. |author3=Wehr W.C. |year=2003 |title=''Corylus'', ''Carpinus'', and ''Palaeocarpinus'' (Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America |journal= International Journal of Plant Sciences |volume=164 |issue= 5 |pages=807–822 |doi=10.1086/376816|s2cid=19802370 }}</ref>

Chilean hazel (''[[Gevuina avellana]]''), despite its name, is not related to this genus.<ref>{{cite book |last1=Crawford |first1=Martin |title=Shrubs for Gardens, Agroforestry and Permaculture |date=2020 |publisher=Permanent Publications |isbn=978-1-85623-342-2 |language=en}}</ref>



== Ecology ==
== Ecology ==
At least 21 species of [[fungus]] have a mutualistic relationship with hazel. ''[[Lactarius pyrogalus]]'' grows almost exclusively on hazel, and hazel is one of two kinds of host for the rare ''[[Hypocreopsis rhododendri]]''. Several rare species of Graphidion [[lichen]] depend on hazel trees. In the UK, five species of [[moth]] are specialised to feed on hazel including ''[[Parornix devoniella]]''. Animals which eat hazelnuts include [[red deer]], [[dormouse]] and [[red squirrel]].<ref>[https://treesforlife.org.uk/forest/species-profiles/hazel/ Trees for Life - Hazel]</ref>
At least 21 species of [[fungus]] have a mutualistic relationship with hazel. ''[[Lactarius pyrogalus]]'' grows almost exclusively on hazel, and hazel is one of two kinds of host for the rare ''[[Hypocreopsis rhododendri]]''. Several rare species of ''Graphidion'' [[lichen]] depend on hazel trees. In the UK, five species of [[moth]] are specialised to feed on hazel including ''[[Parornix devoniella]]''. Animals which eat hazelnuts include [[red deer]], [[dormouse]] and [[red squirrel]].<ref>[https://treesforlife.org.uk/forest/species-profiles/hazel/ Trees for Life - Hazel]</ref>


== Uses ==
== Uses ==
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A number of [[cultivar]]s of the common hazel and filbert are grown as [[ornamental plant]]s in [[garden]]s, including forms with contorted stems (''C. avellana'' 'Contorta', popularly known as "Corkscrew hazel" or "[[Harry Lauder]]'s walking stick" from its gnarled appearance); with weeping branches (''C. avellana'' 'Pendula'); and with purple leaves (''C. maxima'' 'Purpurea').
A number of [[cultivar]]s of the common hazel and filbert are grown as [[ornamental plant]]s in [[garden]]s, including forms with contorted stems (''C. avellana'' 'Contorta', popularly known as "Corkscrew hazel" or "[[Harry Lauder]]'s walking stick" from its gnarled appearance); with weeping branches (''C. avellana'' 'Pendula'); and with purple leaves (''C. maxima'' 'Purpurea').


Hazel is a traditional material used for making [[Wattle (construction)|wattle]], [[withy]] fencing, baskets, and the frames of [[coracle]] boats. The tree can be [[coppicing|coppiced]], and regenerating shoots allow for harvests every few years.
Hazel is a traditional material used for making [[Wattle (construction)|wattle]], [[withy]] fencing, baskets, and the frames of [[coracle]] boats. The tree can be [[coppicing|coppiced]],<ref>{{cite web |title=Coppicing |url=https://www.smallwoods.org.uk/en/advice-and-information/the-economics-of/coppicing/ |website=Small Woods |access-date=19 October 2023}}</ref> and regenerating shoots allow for harvests every few years. There is a 7 year cycle (cut and grow) for [[hurdle]] (fence) making.<ref>{{cite web |title=TYPES OF WOODLAND MANAGEMENT |url=https://www.woodlandtrust.org.uk/plant-trees/managing-trees-and-woods/types-of-woodland-management/ |access-date=19 October 2023}}</ref>


Hazels are used as food plants by the [[larva]]e of various [[list of Lepidoptera that feed on hazels|species]] of [[Lepidoptera]].
Hazels are used as food plants by the [[larva]]e of various species of [[Lepidoptera]]. Including ''[[Eriocrania chrysolepidella]]''.<ref>{{cite web |title=''Paracrania chrysolepidella'' [Lepidoptera: Eriocraniidae] in Leaf and stem mines of British flies and other insects |url=http://www.ukflymines.co.uk/Moths/Eriocrania_chrysolepidella.php |website=www.ukflymines.co.uk |access-date=19 October 2023}}</ref>


==Culture==
==Culture==
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"The Hazel Branch" from ''[[Grimms' Fairy Tales]]'' claims that hazel branches offer the greatest protection from snakes and other things that creep on the earth. In the Grimm tale "Cinderella", a hazel branch is planted by the protagonist at her mother's grave and grows into a tree that is the site where the girl's wishes are granted by birds.<ref>Pitt.edu https://www.pitt.edu/~dash/grimm021.html June 1, 2011</ref>
"The Hazel Branch" from ''[[Grimms' Fairy Tales]]'' claims that hazel branches offer the greatest protection from snakes and other things that creep on the earth. In the Grimm tale "Cinderella", a hazel branch is planted by the protagonist at her mother's grave and grows into a tree that is the site where the girl's wishes are granted by birds.<ref>Pitt.edu https://www.pitt.edu/~dash/grimm021.html June 1, 2011</ref>

This is not true the hazel was fond in a cave under a rock and then the animals grew the hazel into 10 and then 20 and so on we have even more hazels


==Gallery==
==Gallery==

Revision as of 11:16, 11 January 2024

Hazels
Common hazel (Corylus avellana)
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fagales
Family: Betulaceae
Subfamily: Coryloideae
Genus: Corylus
L.
Type species
Corylus avellana
Species

See text for species.

Synonyms[1]

Lopima Dochnahl

Young male catkins of Corylus avellana

Hazels are plants of the genus Corylus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,[2][3][4][5] though some botanists split the hazels (with the hornbeams and allied genera) into a separate family Corylaceae.[6][7] The fruit of the hazel is the hazelnut.

Hazels have simple, rounded leaves with double-serrate margins. The flowers are produced very early in spring before the leaves, and are monoecious, with single-sex catkins. The male catkins are pale yellow and 5–12 centimetres (2–4+34 inches) long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3 mm-long styles visible. The fruits are nuts 1–2.5 cm (12–1 in) long and 1–2 cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut.[4]

The shape and structure of the involucre, and also the growth habit (whether a tree or a suckering shrub), are important in the identification of the different species of hazel.[4]

The pollen of hazel species, which are often the cause for allergies in late winter or early spring, can be identified under magnification (600×) by their characteristic granular exines bearing three conspicuous pores.[8]

Species

Corylus has around 14–18 species. The circumscription of species in eastern Asia is disputed, with World Flora Online and the Flora of China differing in which taxa are accepted, within this region. WFO accepts 17 species while Flora of China accepts 20 species (including Corylus mandshurica).

Only those taxa accepted by both sources are listed below.[9][10][11]

The species are grouped as follows:

Several hybrids exist, and they can occur between species in different sections of the genus, e.g. Corylus × colurnoides (C. avellana × C. colurna). The oldest confirmed hazel species is Corylus johnsonii found as fossils in the Ypresian-age rocks of Ferry County, Washington.[12]

Chilean hazel (Gevuina avellana), despite its name, is not related to this genus.[13]


Ecology

At least 21 species of fungus have a mutualistic relationship with hazel. Lactarius pyrogalus grows almost exclusively on hazel, and hazel is one of two kinds of host for the rare Hypocreopsis rhododendri. Several rare species of Graphidion lichen depend on hazel trees. In the UK, five species of moth are specialised to feed on hazel including Parornix devoniella. Animals which eat hazelnuts include red deer, dormouse and red squirrel.[14]

Uses

Hazel coppice in winter at Bubbenhall in Warwickshire, England

The nuts of all hazels are edible. The common hazel is the species most extensively grown for its nuts, followed in importance by the filbert. Nuts are also harvested from the other species, but apart from the filbert, none is of significant commercial importance.[5]

A number of cultivars of the common hazel and filbert are grown as ornamental plants in gardens, including forms with contorted stems (C. avellana 'Contorta', popularly known as "Corkscrew hazel" or "Harry Lauder's walking stick" from its gnarled appearance); with weeping branches (C. avellana 'Pendula'); and with purple leaves (C. maxima 'Purpurea').

Hazel is a traditional material used for making wattle, withy fencing, baskets, and the frames of coracle boats. The tree can be coppiced,[15] and regenerating shoots allow for harvests every few years. There is a 7 year cycle (cut and grow) for hurdle (fence) making.[16]

Hazels are used as food plants by the larvae of various species of Lepidoptera. Including Eriocrania chrysolepidella.[17]

Culture

The Celts believed hazelnuts gave one wisdom and inspiration. There are numerous variations on an ancient tale that nine hazel trees grew around a sacred pool, dropping into the water nuts that were eaten by salmon (a fish sacred to Druids), which absorbed the wisdom. A Druid teacher, in his bid to become omniscient, caught one of these special salmon and asked a student to cook the fish, but not to eat it. While he was cooking it, a blister formed and the pupil used his thumb to burst it, which he naturally sucked to cool, thereby absorbing the fish's wisdom. This boy was called Fionn Mac Cumhail (Fin McCool) and went on to become one of the most heroic leaders in Gaelic mythology.[18]

"The Hazel Branch" from Grimms' Fairy Tales claims that hazel branches offer the greatest protection from snakes and other things that creep on the earth. In the Grimm tale "Cinderella", a hazel branch is planted by the protagonist at her mother's grave and grows into a tree that is the site where the girl's wishes are granted by birds.[19]

Gallery

References

  1. ^ Kew World Checklist of Selected Plant Families
  2. ^ Germplasmgobills Information Network: Corylus Archived 2009-01-14 at the Wayback Machine
  3. ^ Chen, Zhi-Duan; Manchester, Steven R; Sun, Hai-Ying (August 1999). "Phylogeny and evolution of the Betulaceae as inferred from DNA sequences, morphology, and paleobotany". American Journal of Botany. 86 (8): 1168–1181. doi:10.2307/2656981. ISSN 0002-9122. JSTOR 2656981. PMID 10449397.
  4. ^ a b c Rushforth, K. (1999). Trees of Britain and Europe. Collins ISBN 0-00-220013-9.
  5. ^ a b Huxley, A., ed. (1992). New RHS Dictionary of Gardening. Macmillan ISBN 0-333-47494-5.
  6. ^ Bean, William Jackson (1976). Trees and Shrubs Hardy in the British Isles. Vol. 1. Taylor, George (8th ed.). London: J. Murray. ISBN 0719517907. OCLC 103403.
  7. ^ Erdogan, V.; Mehlenbacher, S. A. (2002). "Phylogenetic analysis of hazelnut species (Corylus, Corylacae) based on morphology and phenology". Sist. Bot. Dergisi. 9: 83–100.
  8. ^ Airy, Hubert (1874). "Pollen-grains in the Air". Nature. 10 (253): 355. Bibcode:1874Natur..10..355A. doi:10.1038/010355b0. S2CID 4077214.
  9. ^ "Corylus L." Retrieved 19 October 2023.
  10. ^ "Corylus in Flora of China @ efloras.org". www.efloras.org. Retrieved 19 October 2023.
  11. ^ Flora of North America: Corylus
  12. ^ Pigg, K.B.; Manchester S.R.; Wehr W.C. (2003). "Corylus, Carpinus, and Palaeocarpinus (Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America". International Journal of Plant Sciences. 164 (5): 807–822. doi:10.1086/376816. S2CID 19802370.
  13. ^ Crawford, Martin (2020). Shrubs for Gardens, Agroforestry and Permaculture. Permanent Publications. ISBN 978-1-85623-342-2.
  14. ^ Trees for Life - Hazel
  15. ^ "Coppicing". Small Woods. Retrieved 19 October 2023.
  16. ^ "TYPES OF WOODLAND MANAGEMENT". Retrieved 19 October 2023.
  17. ^ "Paracrania chrysolepidella [Lepidoptera: Eriocraniidae] in Leaf and stem mines of British flies and other insects". www.ukflymines.co.uk. Retrieved 19 October 2023.
  18. ^ Floriz: Mythology and Folklore of the Hazel Tree Archived 2012-03-31 at the Wayback Machine
  19. ^ Pitt.edu https://www.pitt.edu/~dash/grimm021.html June 1, 2011

External links