Catocalinae
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Catocalinae | ||||||||||||
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Red ribbon ( Catocalina nupta ) |
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Systematics | ||||||||||||
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Scientific name | ||||||||||||
Catocalinae | ||||||||||||
Boisduval , 1828 |
The catocalinae are a subfamily of butterflies ( moths ) from the family of Erebidae . Over 10,000 species are known worldwide. In Europe there are only about 96 species, in Central Europe a little over 30 species. They are mostly relatively large, mostly nocturnal moths with mostly very colored hind wings and mostly inconspicuous forewings.
features
The species in this subfamily are usually relatively large. The European species are on the order of 22 to 98 mm wingspan. The hind wings are often prominently colored, while the forewings are mostly inconspicuous black, gray or brown. However, there are also a few exceptions (e.g. Ophiusa tirhaca with striking yellow forewings ). Goater et al. (2003) list three autapomorphies that justify this group as a monophylum:
- Hind wing with a terminal black spot and fringes between the M3 and Cu1 veins of the wing veins
- In the female genital apparatus, the posterior apophyses are rod-shaped or basal only slightly wider, without the usually flat diamond-shaped basal structure near the ovipositor.
There are also a few features that are not present in all groups:
- the wings are kept flat in the rest position
- colored and / or patterned hind wings
- a quadrifid veining of the hind wings
- a flake-free field directly above the proboscis
- as well as five other characteristics in the male and female genital apparatus
Geographical distribution and habitat
The subfamily Catocalinae is distributed worldwide; the main focus of the distribution is in the tropics and subtropics . Some species migrate over longer distances, and therefore subtropical species can sometimes be found in Europe. Due to the large geographical distribution and the huge number of species, a wide range of habitats are populated, ranging from dry deserts and semi-deserts to tropical rainforests.
Way of life
Due to the large geographical distribution, some species form only one generation per year, other species several generations. The moths are mostly nocturnal. However, there are also some species that are also diurnal, such as the brown day owl ( Euclidia glyphica ) and the check day owl ( Callistege mi ). The moths ingest food and can therefore be baited . As a rule, you also come across artificial light sources. There is also a very wide range of host plants due to their large geographical distribution, diverse habitats and large number of species. However, this does not apply to the individual species, which are sometimes even monophagous .
Systematics
The subfamily Catocalinae originally comprised all "quadrifine" Noctuids. However, this characteristic is original and cannot be used to define the group. Only Goater, Ronkay and Fibiger managed to find three characteristics in 2003 that are to be assessed as apomorphic and can therefore be used to detect monophyly. However, some groups had to be excluded in this version. The Calpinae, which were previously often brought here, are regarded as an independent subfamily. The following list of the genera and species occurring in Europe is Goater et al. (2003) taken. Some non-European species are added.
- Tribus Acantholipini Fibiger & Lafontaine, 2005
- Genus Acantholipes Lederer, 1857
- Acantholipes regularis (Huebner, 1813)
- Genus Acantholipes Lederer, 1857
- Tribus Arytrurini Fibiger & Lafontaine, 2005
- Genus Arytrura John, 1912
- Arytrura musculus (Ménétriés, 1859)
- Genus Arytrura John, 1912
- Tribus Catocalini Boisduval, 1828
- Genus Catocala cabinet, 1802
- Eurasian species
- Catocala actaea Felder & Rogenhofer, 1874
- Catocala adultera Ménétriés, 1856
- Catocala brandti Hacker & Kautt, 1999
- Catocala conjuncta (Esper, 1787)
- Catocala conversa (Esper, 1783)
- Catocala deducta Eversmann, 1843
- Catocala detrita Warren, 1913
- Catocala dilecta (Huebner, 1808)
- Catocala disjuncta (Geyer, 1828)
- Catocala diversa (Geyer, 1828)
- Willow carmine ( Catocala electa (Vieweg, 1790))
- Poplar carmine ( Catocala elocata ) (Esper, 1787)
- Catocala eutychea Treitschke, 1835
- Blue ribbon ( Catocala fraxini (Linné, 1758))
- Yellow ribbon ( Catocala fulminea (Scopoli, 1763))
- Catocala hymenaea (Denis & Schiffermüller, 1775)
- Catocala lara Bremer, 1861
- Catocala lupina Herrich-Schäffer, 1851
- Catocala mariana Rambur, 1858
- Catocala neonympha (Esper, 1805)
- Red ribbon ( Catocala nupta (Linné, 1767))
- Catocala nymphaea (Esper, 1787)
- Catocala nymphagoga (Esper, 1787)
- Catocala oberthueri Austaut, 1879
- Catocala optata (Godart, 1825)
- Catocala orientalis Staudinger, 1877
- Broken willow carmine ( Catocala pacta (Linné, 1758))
- Small oak carmine ( Catocala promissa (Denis & Schiffermüller, 1775))
- Catocala puerpera (Giorna, 1791)
- Catocala separata Freyer, 1848
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Large oak carmine ( Catocala sponsa (Linné, 1767))
- North American species
- Catocala amica Huebner, 1818
- Catocala angusi Grote, 1876
- Catocala antinympha (Huebner, 1823)
- Catocala cerogama Guenée, 1852
- Catocala coccinata Grote, 1852
- Catocala connubialis Guenée, 1852
- Catocala epione (Drury, 1773)
- Catocala grynea Cramer, 1780
- Catocala innubens Guenée, 1852
- Catocala lacrymosa Guenée, 1852
- Catocala micronympha Guenée, 1852
- Catocala nebulosa Edwards, 1864
- Catocala palaeogama Guenée, 1852
- Catocala relicta Walker, 1858
- Catocala ultronia (Huebner, 1823)
- Genus Catocala cabinet, 1802
- Tribus Euclidiini Fibiger & Lafontaine, 2005
- Genus Euclidia Ochsenheimer, 1816 (with the subgenera E. (Euclidia) , E. Callistege) Huebner, 1823 and E. Gonospileia) Huebner, 1823)
- Euclidia fortalicium (Tauscher, 1809)
- Brown day owl ( Euclidia glyphica (Linné, 1758))
- Check day owl ( Euclidia mi (Clerck, 1759)
- Euclidia munita (Huebner, 1813)
- Euclidia triquetra (Denis & Schiffermüller, 1775)
- Genus Euclidia Ochsenheimer, 1816 (with the subgenera E. (Euclidia) , E. Callistege) Huebner, 1823 and E. Gonospileia) Huebner, 1823)
- Tribus Melipotini Richards, 1933
- Genus Drasteria Huebner, 1818
- Drasteria flexuosa (Ménétriés, 1848)
- Drasteria tenera (Staudinger, 1877)
- Drasteria caucasica (Kolenati, 1846)
- Drasteria picta (Christoph, 1877)
- Drasteria cailino (Lefebvre, 1827)
- Drasteria saisani (Staudinger, 1882)
- Drasteria sesquistria (Eversmann, 1854)
- Drasteria rada (Boisduval, 1848)
- Genus Drasteria Huebner, 1818
- Tribus Ophiusini Guenee, 1837
- Genus Achaea Huebner, 1823
- Achaea catocaloides Guenee , 1852, West Africa
- Genus Catephia Ochsenheimer, 1816
- White ribbon ( Catephia alchymista (Denis & Schiffermüller, 1775)
- Genus Cerocala Boisduval, 1829
- Cerocala algiriae Oberthür, 1876
- Cerocala scapulosa (Huebner, 1808)
- Genus Clytie Huebner, 1823
- Clytie delunaris (Staudinger, 1889)
- Clytie syriaca (Bugnion, 1837)
- Clytie illunaris (Huebner, 1813)
- Clytie gracilis (Bang-Haas, 1907)
- Genus Dysgonia Huebner, 1823
- Dysgonia rogenhoferi (Bohatsch, 1880)
- Dysgonia torrida Guenée, 1852
- Dysgonia algira (Linnaeus, 1767)
- Genus Grammodes Guénée, 1852 (including the genus Prodotis John, 1910)
- Grammodes bifasciata (Petagna, 1787)
- Grammodes stolida (Fabricius, 1775)
- Genus Heteropalpia Berio, 1939
- Heteropalpia acrosticta ( Püngeler , 1904)
- Genus Minucia Moore, 1885
- Brown ribbon ( Minucia lunaris (Denis & Schiffermüller, 1775))
- Genus Ophiusa Ochsenheimer, 1816
- Ophiusa tirhaca (Cramer, 1773)
- Genus Pandesma Guenée, 1852
- Pandesma robusta (Walker, 1858)
- Genus Percyma Herrich-Schäffer, 1851
- Pericyma albidentaria (Freyer, 1842)
- Genus Zethes Rambur, 1833
- Zethes insularis Rambur, 1833
- Genus Achaea Huebner, 1823
- Tribus Toxocampini Goater, Ronkay & Fibiger, 2003
- Genus Anumeta Walker, 1858
- Anumeta atrosignata Walker, 1858
- Anumeta spilota (Ershov, 1874)
- Anumeta henkei (Staudinger, 1877)
- Anumeta fractistrigata (Alphéraky, 1882)
- Anumeta cestis (Ménétriés, 1848)
- Genus Apopestes Huebner, 1823
- Apopestes spectrum (Esper, 1787)
- Genus Autophila Hübner, 1823 (with the subgenera A. (Autophila) and A. Cheirophanes Boursin, 1955)
- Autophila hirsuta (Staudinger, 1870)
- Autophila libanotica (Staudinger, 1901)
- Autophila limbata (Staudinger, 1871) (with two subspecies)
- Autophila dilucida (Huebner, 1808)
- Autophila asiatica (Staudinger, 1888)
- Autophila rosea (Staudinger, 1888)
- Cheirophanes cataphanes (Hübner, 1813) (with two subspecies)
- Cheirophanes anaphanes Boursin, 1940 (with two subspecies)
- Cheirophanes chamaephanes Boursin, 1940
- Cheirophanes ligaminosa (Eversmann, 1851)
- Cheirophanes maura (Staudinger, 1888)
- Genus Chrysorithrum Butler, 1878
- Chrysorithrum flavomaculata (Bremer, 1861)
- Genus Exophyla Guenée, 1841
- Exophyla rectangularis (Geyer, 1828)
- Genus Lygephila Billberg, 1820
- Lygephila lusoria (Linné, 1758)
- Lygephila lubrica (Freyer, 1846)
- Lygephila ludicra (Huebner, 1790)
- Lygephila lycyrrhizae (Staudinger, 1871)
- Lygephila fonti Yela & Calle, 1990
- Kidney spot wicked owl ( Lygephila pastinum (Treitschke, 1826))
- Marbled wicked owl ( Lygephila viciae (Hübner, 1822))
- Lycra owl ( Lygephila craccae (Denis & Schiffermüller, 1775))
- Lygephila procax (Huebner, 1813)
- Genus Tathorhynchus Hampson, 1894 (considered by Goater et al., 2003 as a subgenus of Lygephila )
- Tathorhynchus exsiccata (Lederer, 1855)
- Genus Anumeta Walker, 1858
swell
literature
- Günter Ebert (Ed.): The Butterflies of Baden-Württemberg Volume 5, Moths III (Sesiidae, Arctiidae, Noctuidae). Ulmer Verlag Stuttgart 1997. ISBN 3-8001-3481-0
- Barry Goater, Lázló Ronkay, Michael Fibiger: Catocalinae & Plusiinae. Noctuidae Europaeae, Volume 10., Sorø, 2003, ISBN 87-89430-08-5