Systematics of the nightshade family
The systematics of the nightshade family (Solanaceae) has not yet been generally recognized. This is mainly due to the size of the nightshade family , but also to the morphological diversity and the fact that various morphological features can be subject to changes both genetically and locally, and that some features have developed several times in parallel within the family .
The following systematics are the newer systematics that are complete down to the genus or sub-genus level, none of which, however, fully depicts the relationships within the family.
Systematics according to D'Arcy
(Without sub-genera)
- Subfamily Solanoideae
- Tribe Solaneae: Acnistus , Athenaea , Capsicum , Cacabus , Cuatresia , Cyphomandra 1 , Deprea , Dunalia , Exodeconus , Iochroma , jaltomata , Lycopersicon 1 , Withania , Physalis , Leucophysalis , Magaranthus , Quincula , Saracha , Hebecladus , Archiphysalis , Chamaesaracha , Solanum , Lycianthes , Brachistus , Vassobia , Witheringia , Atropa , Hyoscyamus , Mandragora
- Tribe Daturae: Brugmansia , Datura
- Tribe Jaboroseae: Jaborosa , Salpichroa , Trechonaetes
- Tribe Lycieae: Grabowskia , Lycium , Phrodus
- Tribus Nicandreae: Nicandra
- Tribus Solandreae: Solandra , Trianaea
- Tribus Juanulloeae: Hawkesiophyton , Juanulloa , Markea , Schultesianthus , Rahowardiana
- Tribe Hyoscyameae: Hyoscyamus , Scopolia , Physochlaina
- Genera not classified within the subfamily: Discopodium , Mellissia , Nectouxia , Nothocestrum , Oryctes , Pauia , Triguera 1 , Sterrhymenia
- Subfamily Cestroideae
- Tribus Cestreae: Cestrum , Metternichia , Sessea , Sesseopsis , Vestia , Phrodus
- Tribe Nicotianea: Benthamiella , Bouchetia , Combera , Nicotiana , Nierembergia , Pantacantha , Petunia , Fabiana , Latua
- Tribus Salpiglossideae: Browallia , Brunfelsia , Leptoglossis , Reyesia , Salpiglossis , Hunzikeria , Schizanthus , Streptosolen
- Tribe Schwenckieae: Melananthus , Protoschwenckia , Schwenckiopsis , Schwenckia
- Tribe Parabouchetieae: Parabouchetia
- Tribus Anthocercideae: Anthocercis , Anthotroche , Crenidium , Cyphanthera , Duboisia , Grammosolen , Symonanthus
- genera not classified within the subfamily: Anisodus , Atrichodendron , Heteranthia , Przewalskia
Systematics according to Hunziker
- Subfamily Cestroideae
-
- Tribus Cestreae: Cestrum , Vestia , Sessea
- Tribus Metternichieae: Metternichia
- Tribus Latueae: Latua
- Tribe Nicotianeae
- Subertribus Nicotianinae: Nicotiana , Petunia , Fabiana
- Subertribus Nierembergiinae: Nierembergia , Bouchetia
- Subertribus Leptoglossinae: Leptoglossis , Hunzikeria , Plowmania
- Tribus Benthamielleae: Benthamiella , Pantacantha , Combera
- Tribus Francisceae: Brunfelsia
- Tribus Browallieae: Browallia , Streptosolen
- Tribe Schwenckieae: Schwenckia , Melananthus , Protoschwenckia , Heteranthia
- Subfamily Juanulloideae: Juanulloa , Dyssochroma , Ectozoma , Hawkesiophyton , Markea , Merinthopodium , Rahowardiana , Schultesianthus , Trianaea
- Subfamily Solanoideae
-
- Tribus Nicandreae: Nicandra
- Tribus Mandragoreae: Mandragora
- Tribe Datureae: Datura , Brugmansia
- Tribe Lycieae: Lycium , Phrodus , Grabowskia
- Tribe Solaneae
- Untertribus Witheringinae: Witheringia , Brachistus , Cuatresia , Deprea , Discopodium , Exodeconus , Jaltomata , Nothocestrum , Acnistus
- Untertribus Physalinae: Physalis , Quincula , Leucophysalis , Chamaesaracha
- Subertribus Iochrominae: Iochroma , Saracha , Oryctes , Tubocapsicum
- Subertribus Capsicinae: Capsicum , Aureliana , Athenaea , Darcyanthus , Eriolarynx , Vassobia , Larnax , Dunalia , Withania
- Untertribus Solaninae: Solanum , Cyphomandra 1 , Lycopersicon 1 , Lycianthes , Triguera 1 , Normania 1
- Tribe Atropeae: Atropa
- Tribe Jaboroseae: Jaborosa , Salpichroa , Nectouxia
- Tribe Solandreae: Solandra
- Tribe Hyoscyameae: Hyoscyamus , Anisodus , Atropanthe , Physochlaina , Przewalskia , Scopolia
- Subfamily Salpiglossoideae: Salpiglossis , Reyesia
- Subfamily Schizanthoideae: Schizanthus
- Subfamily Anthocercidoideae: Anthocercis , Anthotroche , Cyphanthera , Crenidium , Duboisia , Grammosolen , Symonanthus
Genera that, according to Hunziker, are no longer included in the nightshade family: Duckeodendron , Eutheta , Goetzea , Espadaea , Coeloneurum , Bissea , Lithophytum , Nolana , Parabouchetia , Retzia , Sclerophylax , Tsoala , Tunaria , Valerioa .
Systematics according to Olmstead (1999)
- Subfamily Cestroideae
- Tribus Browallieae: Browallia , Streptosolen
- Tribus Cestreae: Cestrum , Metternichia , Sessea , Vestia
- Tribus Salpiglossideae: Reyesia , Salpiglossis
- Subfamily Goetzeoideae: Coeloneurum , Espadaea , Goetzea , Henoonia
- Subfamily Petunioideae: Benthamiella , Bouchetia , Brunfelsia , Calibrachoa , Combera , Fabiana , Hunzikeria , Latua , Leptoglossis , Nierembergia , Pantacantha , Petunia , Plowmania
- Subfamily Schizanthoideae: Schizanthus
- Subfamily Schwenckioideae: Heteranthia , Melananthus , Protoschwenckia , Schwenckia
- Subfamily Nicotianoideae
- Tribus Anthocercideae, Anthocercis , Anthotroche , Crenidium , Cyphanthera , Duboisia , Grammosolen , Symonanthus
- Tribe Nicotianeae: Nicotiana
- Subfamily Solanoideae
-
- Tribe Capsiceae: Capsicum , Lycianthes
- Tribe Datureae: Brugmansia , Datura , Methysticodendron
- Tribe Hyoscyameae: Anisodus , Atropa , Atropanthe , Hyoscyamus , Physochlaina , Przewalskia , Scopolia
- Tribe Jaboroseae: Jaborosa
- Tribe Solandreae
- Subertribus Juanulloinae, Dyssochroma , Ectozoma , Hawkesiophyton , Juanulloa , Markea , Merinthopodium , Rahowardiana , Schultesianthus , Trianaea
- Subertribus Solandrinae: Solandra
- Tribe Lycieae: Grabowskia , Lycium , Phrodus
- Tribus Mandragoreae: Mandragora
- Tribe Nicandreae: Exodeconus , Nicandra
- Tribe Nolaneae: Alona , Nolana
- Tribe Physaleae
- Subertribus Iochrominae: Acnistus , Dunalia , Iochroma , Saracha , Vassobia
- Untertribus Physalinae: Brachistus , Chamaesaracha , Leucophysalis , Magaranthus , Oryctes , Quincula , Physalis , Witheringia
- Subertribus Salpichroinae: Nectouxia , Salpichroa
- Subertribus Withaninae, Archiphysalis , Athenaea , Aureliana , Cuatresia , Deprea , Larnax , Mellissia , Physalisatrum , Tubocapsicum , Withania
- Tribus Solaneae, Cyphomandra 1 , Discopodium , Jaltomata , Lycopersicon 1 , Normania 1 , Nothocestrum , Solanum , Triguera
Systematics according to Olmstead (2008)
- Not classified in a subfamily: Duckeodendron , Schizanthus
- Subfamily Goetzeoideae: Coeloneurum , Espadaea , Goetzea , Henoonia , Metternichia , Tsoala
- Tribus Benthamielleae (possibly also classified in Cestroideae): Benthamiella , Combera , Panthacantha
- Subfamily Cestroideae:
- Tribus Cestreae: Cestrum , Sessea , Vestia
- Tribus Browallieae: Browallia , Streptosolen
- Tribus Salpiglossideae: Reyesia , Salpiglossis
- Protoschwenckia
- Tribus Petunieae: Bouchetia , Brunfelsia , Calibrachoa , Fabiana , Hunzikeria , Leptoglossis , Nierembergia , Petunia , Plowmania
- Tribe Schwenckieae: Heteranthia , Melanathus , Schwenckia
-
x = 12 -clade (no rank)
- Subfamily Nicotianoideae:
- “Tribus” Anthocercideae: Anthocercis , Anthotroche , Crenidium , Cyphanthera , Duboisia , Grammosolen , Symonanthus
- Nicotiana
- Subfamily Solanoideae:
- Atropina (not ranked):
- Tribe Hyoscyameae: Anisodus , Atropanthe , Atropa , Hyoscyamus , Physochlaina , Przewalskia , Scopolia
- Tribe Lycieae: Lycium (incl. Grabowskia , Phrodus )
- Jaborosa , Latua , Nolana , Sclerophylax
- Tribe Capsiceae: Capsicum , Lycianthes
- Tribe Datureae: Brugmansia , Datura , Iochroma cardenasianum
- Tribus Juanulloeae: Dyssochroma , Juanulloa , Markea (incl. Hawkesiophyton ), Merinthopodium , Trianaea
- Tribe Physaleae:
- Subtribus Withaninae: Athenaea , Aureliana , Discopodium , Nothocestrum , Tubocapsicum , Withania (incl. Archiphysalis , Physaliastrum , Mellissia )
- Subtribe Iochrominae: Acnistus , Dunalia , Eriolarynx , Iochroma , Saracha , Vassobia
- Subtribus Physalinae: Brachistus , Chamaesaracha , Leucophysalis , Margaranthus , Oryctes , Physalis , Quincula , Tzeltalia , Witheringia
- Not further classified in the Physaleae: Cuatresia , Deprea , Larnax
- Salpichroina (without rank): Nectouxia , Salpichroa
- Tribe Solaneae: Jaltomata , Solanum
- Not classified further in the Solanoideae: Exodeconus , Mandragora , Nicandra , Schultesianthus , Solandra
- Atropina (not ranked):
- Subfamily Nicotianoideae:
swell
- JAM van Balken: Overview of Solanaceae Species , online
- Armando T. Hunziker: The Genera of Solanaceae. ARG Gantner Verlag KG, Ruggell, Liechtenstein 2001. ISBN 3-904144-77-4 .
- Richard G. Olmstead et al .: Phylogeny and Provisional Classification of the Solanaceae Based on Chloroplast DNA (PDF; 131 kB) . In Solanaceae IV, Advances in Biology and Utilization , editors: M. Nee, DE Symon, JP Jessup, and JG Hawkes, Royal Botanic Gardens, Kew. 1999. Pages 111-137.
- Richard G. Olmstead et al .: A molecular phylogeny of the Solanaceae . In: Taxon , Volume 57, Number 4, November 2008. pp. 1159-1181.
to 1 : The genera Cyphomandra ( Tamarillo ), Lycopersicon ( tomatoes ), Normania and Triguera the genus are commonly today Solanum ( Solanum ) subordinate.