Striped Ferns

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Striped Ferns
Asplenium species: Nordic striped fern (Asplenium septentrionale) above, spleen fern (Asplenium ceterach) below center and brown-stemmed striped fern (Asplenium trichomanes) below left

Asplenium species:
Nordic striped fern ( Asplenium septentrionale ) above, spleen fern ( Asplenium ceterach ) below center and brown-stemmed striped fern ( Asplenium trichomanes ) below left

Systematics
Department : Vascular plants (tracheophyta)
Ferns
Class : True ferns (Polypodiopsida)
Order : Spotted ferns (Polypodiales)
Family : Striped fern family (Aspleniaceae)
Genre : Striped Ferns
Scientific name
Asplenium
L.

The striped ferns ( Asplenium ) are a genus of plants in the family of the striped fern plants (Aspleniaceae). This species-rich genus is found worldwide.

description

Illustration of the brown-stemmed striped fern ( Asplenium trichomanes ), left and the black-stemmed striped fern ( Asplenium adiantum-nigrum ), right
Cross-section of an Asplenium- Sorus microscope scale = 0.35 mm
Illustration of A wall striped fern ( Asplenium ruta-muraria , left) and B Nordic striped fern ( Asplenium septentrionale , right)
Illustration of Asplenium shuttleworthianum from Gustav Kunze: The fern herbs in colored illustrations, faithfully explained and described , 1840

The striped fern species are perennial herbaceous plants . They thrive epiphytically , lithophytically or terrestrially. They have a short, upright, sometimes creeping rhizome on which dark, linear-lanceolate, latticed (clathrate) scales sit. The fertile (sporangia-bearing) and sterile fronds are identical: up to fourfold pinnate or forked, less often (as in the deer tongue ) undivided. The veins end freely. The petioles are usually dark at the base, often glabrous and have two vascular bundles , which often unite to form an x-shaped bundle.

The Sori stand individually and away from the edge. Like the veil ( Indusium ), they are mostly linear, from which the name striped fern is derived.

The basic chromosome number is x = 35 or 36.

Systematics and distribution

Black-stemmed striped fern ( Asplenium adiantum-nigrum )
Billot's striped fern ( Asplenium billotii )
Viviparous striped fern ( Asplenium bulbiferum )
Spleen fern ( Asplenium ceterach )
Jurassic Striped Fern ( Asplenium fontanum )
Ivy fern ( Asplenium hemionitis )
Sea striped fern ( Asplenium marinum )
Nest fern ( Asplenium nidus )
Asplenium obovatum subsp. obovatum
Creeping striped fern ( Asplenium rhizophyllum )
Wall striped fern ( Asplenium ruta-muraria )
Deer tongue ( Asplenium scolopendrium )
Habit of Asplenium sellowianum in the habitat
Section of the underside of the leaf with sori from Asplenium sellowianum
Habit of Asplenium serratum in the habitat
Habitus of Asplenium trichomanes subsp. pachyrachis
Brown-stemmed striped fern ( Asplenium trichomanes subsp. Quadrivalens )
Habit of Asplenium ulbrichtii in the habitat
Green striped fern ( Asplenium viride )
Habit of Asplenium viviparum var. Daucifolium
Habit of Asplenium wilfordii in the habitat

The genus Asplenium was first published by Carl von Linné . The genus name Asplenium was transferred from the splenic fern ( Asplenium ceterach ) to the genus and is derived from the Greek word splen for spleen and is based on the earlier use of the species in "spleen addiction".

The delimitation of the genus Asplenium within the family Aspleniaceae is controversial. Possibly all other genera of the family belong to the genus Asplenium . Regardless of this, the genus comprises the majority of the family with around 700 species. With the maximum extent of the genus Asplenium L. , synonyms are: Acropteris Link , Amesium Newman nom. Superfl., Antigramma C.Presl , Arcasplenium Moore ms. ex Baker , Asplenidictyum (Hook.) J.Smith , Asplenium nothosubgen. Asplenoceterach (DEMey.) Rosselló, Muss, Salvo & N.Torres , Asplenium subgen. Ceterach (Willd.) Vida ex Bir, Fraser-Jenk. & Lovis , Asplenium subgen. Phyllitis (Hill) Jermy & Viane , Asplenium subgen. Phyllitopsis (Reichst.) Jermy & Viane , Biropteris Kümmerle , Boniniella Hayata , (sometimes considered an accepted genus) Caenopteris Berg , Ceterach Adanson , Ceterach Garsault , Ceterach Lam. & DC. , Ceterach Willd. nom. cons., (sometimes considered an accepted genre). Ceterachopsis (J.Smith) Ching , Ceterachopsis J.Smith , Chamaefilix Hill , Cheilosorum J.Smith , Dareastrum Fée , Darea Jussieu , Diplora Baker , Eremopodium Trevisan , Glossopteris Rafinesque nom. superfl., Glossopteris Rafinesque , Hemionites J.Smith , Hemionitidastrum Fée , Homatoneuron Klotzsch , Hymenasplenium Hayata , Lobium Keyserling , Loxoscaphe (Blume) Presl , Loxoscaphe Moore , Macrophyllidium Rosenst. , Micrasplenium Keyserling , Micropodium Mettenius non Saporta , Neottopteridastrum Fée , Nephrodium J.Sm. , Notolepeum Newman , Onopteris Necker , Parasplenium Keyserling , Phyllitis Hill , Phyllitis J.Smith , Phyllitis Ludwig , Phyllitis Moench , Scolopendrium Adanson , Sinephropteris Mickel , Stormesia Kickx f. , Tarachia Presl , Thamnopteris (Presl) Presl nom. superflat., Trichomanes Bubani , Triphlebia Baker .

There are also numerous inter- and intraspecific hybrids , which, like all fern hybrids, can be recognized by the often clumped spores or undeveloped sporangia.

The more than 700 species are distributed almost worldwide. There are around 17 species in Central Europe. If the genus Hymenasplenium (more than 30 species, 8 of them in China) is separated as an independent one, then 90 species occur in China, 17 of them only there. There are around 30 species in Australia.

There are about 700 species of Asplenium :

hybrid

There are numerous hybrids , one of the most common is:

Further hybrids of the Central European species are listed at Reichstein :

  • Asplenium × adult form Lovis, Melzer & Reichst. (= Asplenium trichomanes subsp. Inexpectans × Asplenium viride )
  • Asplenium × aprutianum Lovis, Melzer & Reichstein (= Asplenium lepidum × Asplenium trichomanes subsp. Quadrivalens )
  • Asplenium × bavaricum D.E. Meyer (= Asplenium trichomanes subsp. Quadrivalens × Asplenium viride )
  • Asplenium × bechereri D.E. Meyer (= Asplenium adiantum-nigrum × Asplenium adulterinum )
  • Asplenium × brissaginense D.E. Meyer (= Asplenium adiantum-nigrum × Asplenium foreziense )
  • Asplenium × centovallense D.E. Meyer (= Asplenium adiantum-nigrum × Asplenium cuneifolium )
  • Asplenium × clermontae Syme (= Asplenium ruta-muraria × Asplenium trichomanes subsp. Quadrivalens , syn .: Asplenium × preissmannii Ascherson & Luerssen , Asplenium × reicheliae Ascherson & Dörfler , Asplenium × hauchecornei Ascherson )
  • Asplenium × contrei Callé, Lovis & Reichstein (= Asplenium adiantum-nigrum × Asplenium septentrionale )
  • Asplenium × costei R. de Litardière (= Asplenium foreziense × Asplenium septentrionale )
  • Asplenium × dolosum Milde (= Asplenium adiantum-nigrum × Asplenium trichomanes subsp. Trichomanes )
  • Asplenium × eglii Lovis & Reichstein (= Asplenium lepidum × Asplenium ruta-muraria subsp. Dolomiticum )
  • Asplenium × gaston-gautieri R. de Litardière (= Asplenium fontanum × Asplenium viride , syn .: Asplenium gautieri Christ )
  • Asplenium × heufleri Reichardt (= Asplenium septentrionale × Asplenium trichomanes subsp. Quadrivalens )
  • Asplenium × hungaricum Fraser-Jenkins & Vida (= Asplenium × murbeckii × Asplenium septentrionale )
  • Asplenium × javorkae Kümmerle (= Asplenium lepidum × Asplenium ruta-muraria subsp. Ruta-muraria )
  • Asplenium × joncheerei D.E. Meyer (= Asplenium obovatum subsp. Billotii × Asplenium adiantum-nigrum subsp. Onopteris )
  • Asplenium × lessinense Vida & Reichstein (= Asplenium fissum × Asplenium viride )
  • Asplenium × lobmingense Melzer, Lovis & Reichstein (= Asplenium adulterinum × Asplenium ruta-muraria subsp. Ruta-muraria )
  • Asplenium × murbeckii Dörfler (= Asplenium ruta-muraria subsp. Ruta-muraria × Asplenium septentrionale )
  • Asplenium × pagesii R. de Litardière (= Asplenium foreziense × Asplenium trichomanes subsp. Quadrivalens , syn .: Asplenium × guichardii R. de Litardière , Asplenium × verbanense von Tavel )
  • Asplenium × perardi R. de Litardière (= Asplenium adiantum-nigrum × Asplenium ruta-muraria subsp. Ruta-muraria )
  • Asplenium × poscharskyanum (Hofm.) Preissmann (= Asplenium adulterinum × Asplenium viride )
  • Asplenium × praetermissum Lovis, Melzer & Reichst. (= Asplenium adulterinum × Asplenium trichomanes subsp. Quadrivalens )
  • Asplenium × protoadulterinum Lovis & Reichst. (= Asplenium trichomanes subsp. Trichomanes × Asplenium viride , Syn .: Asplenium Mendelianum D. E. Meyer )
  • Asplenium × ruscinonense Nieschalk, Lovis & Reichstein (= Asplenium foreziense × Asplenium adiantum-nigrum subsp. Onopteris )
  • Asplenium × sarniense A. Sleep (= Asplenium adiantum-nigrum × Asplenium obovatum subsp. Billotii )
  • Asplenium × sleepiae Badré & Boudrie (= Asplenium obovatum subsp. Billotii × Asplenium foreziense )
  • Asplenium × souchei R. de Litardière (= Asplenium obovatum subsp. Billotii × Asplenium septentrionale )
  • Asplenium × stiriacum D.E. Meyer (= Asplenium lepidum × Asplenium trichomanes subsp. Inexpectans )
  • Asplenium x ticinense D.E. Meyer (= Asplenium adiantum-nigrum × Asplenium adiantum-nigrum subsp. Onopteris )
  • Asplenium × trichomaniforme Woynar (= Asplenium adulterinum × Asplenium trichomanes subsp. Trichomanes )
  • Asplenium × valgannense E. Attinger (= Asplenium seelosii × Asplenium trichomanes subsp. Quadrivalens )
  • Asplenium × wachaviense Ascherson & P. ​​Graebner (= Asplenium cuneifolium × Asplenium trichomanes )
  • Asplenium × woynarianum Ascherson & P. ​​Graebner (= Asplenium cuneifolium × Asplenium viride )

literature

  • JA Crabbe, AC Jermy, R. Viane, JD Lovis: Asplenium L. In: TG Tutin, NA Burges, AO Chater, JR Edmondson, VH Heywood, DM Moore, DH Valentine, SM Walters, DA Webb (Eds.): Flora Europaea . 2nd, revised edition. Volume 1: Psilotaceae to Platanaceae . Cambridge University Press, Cambridge / New York / Melbourne 1993, ISBN 0-521-41007-X , pp. 18–23 (English, limited preview in Google Book Search).
  • Youxing Lin, Ronald Viane: Aspleniaceae. : Asplenium Linnaeus , pp. 267–287 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (ed.): Flora of China. Volume 2-3: Lycopodiaceae through Polypodiaceae. Science Press and Missouri Botanical Garden Press, Beijing and St. Louis, 2013, ISBN 978-1-935641-11-7 .
  • Tadeus Reichstein : Interspecific hybrids in the genus Asplenium . In: Gustav Hegi : Illustrated flora of Central Europe . Volume 1, part 1. 3rd edition. Page 243–266. Publisher Paul Parey, Berlin, Hamburg. 1984. ISBN 3-489-50020-2 .

Individual evidence

  1. a b c d e f g h i j k l m n o p q r s t Siegmund Seybold (Ed.): Schmeil-Fitschen interactive . CD-ROM, version 1.1. Quelle & Meyer, Wiebelsheim 2002, ISBN 3-494-01327-6 .
  2. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf Youxing Lin, Ronald Viane: Aspleniaceae. : Asplenium Linnaeus , pp. 267–287 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (ed.): Flora of China. Volume 2-3: Lycopodiaceae through Polypodiaceae. Science Press and Missouri Botanical Garden Press, Beijing and St. Louis, 2013, ISBN 978-1-935641-11-7 .
  3. Ruprecht Düll , Herfried Kutzelnigg : Pocket dictionary of plants in Germany. A botanical-ecological excursion companion to the most important species . 6th, completely revised edition. Quelle & Meyer, Wiebelsheim 2005, ISBN 3-494-01397-7 .
  4. Helmut Genaust: Etymological dictionary of botanical plant names. 3rd, completely revised and expanded edition. Nikol, Hamburg 2005, ISBN 3-937872-16-7 (reprint from 1996).
  5. ^ Alan R. Smith, Kathleen M. Pryer, Eric Schuettpelz, Petra Korall, Harald Schneider, Paul G. Wolf: A classification for extant ferns. In: Taxon. Volume 55, No. 3, 2006, ISSN  0040-0262 , pp. 705–731, (Abstract, ( Memento of the original dated February 12, 2017 in the Internet Archive ) Info: The archive link has been inserted automatically and has not yet been checked. Please check Original and archive link according to instructions and then remove this note. PDF file) . @1@ 2Template: Webachiv / IABot / www.ingentaconnect.com
  6. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu gv gw gx gy gz ha hb hc hd he hf hg hh hi hj hk hl hm hn ho hp hq hr hs ht hu hv hw hx hy hz ia ib ic id ie if ig ih ii ij ik il im in io ip iq ir is it iu iv iw ix iy iz ja jb jc jd je jf jg jh ji jj jk jl jm jn jo jp jq jr js jt ju jv jw jx jy jz ka kb kc kd ke kf kg kh ki kj kk kl km kn ko kp kq kr ks kt ku kv kw kx ky kz la lb lc ld le lf lg lh li lj lk ll lm ln lo lp lq lr ls lt lu lv lw lx ly lz ma mb mc m d me mf mg mh mi mj mk ml mm mn mo mp mq mr ms mt mu mv mw mx my mz na nb nc nd ne nf ng nh ni nj nk nl nm nn no np nq nr ns nt nu nv nw nx ny nz oa ob oc od oe of og oh oi oj ok ol om on oo op oq or os ot ou ov ow ox oy oz pa pb pc pd pe pf pg ph pi pj pk pl pm pn po pp pq pr ps pt pu pv pw px py pz qa qb qc qd qe qf qg qh qi qj qk ql qm qn qo qp qq qr qs qt qu qv qw qx qy qz ra rb rc rd re rf rg rh ri rj rk rl rm rn ro rp rq rr rs rt ru rv rw rx ry rz sa sb sc sd se sf sg sh si sj sk sl sm sn so sp sq sr ss st su sv sw sx sy sz ta tb tc td te tf tg th ti tj tk tl tm tn to tp tq tr ts tt tu tv tw tx ty tz ua ub uc ud ue uf ug uh ui uj uk ul um un uo up uq ur us ut uu uv uw ux uy uz va vb vc vd ve vf vg vh vi vj vk vl vm vn vo vp vq vr vs vt vu vv vw vx vy vz wa wb wc wd we wf wg wh wi wj wk wl wm wn wo wp wq wr ws wt wu wv ww wx wy wz xa xb xc xd xe xf xg xh xi xj xk xl xm xn xo xp xq xr xs xt xu xv xw xx xy xz ya yb yc yd ye yf yg yh yi yj yk yl ym yn yo yp yq yr ys yt yu yv yw yx yy yz za zb zc zd ze zf zg zh zi zj zk zl zm zn zo Michael Hassler, Brian Swale, 2001/2: Genus Asplenium in the world species list . ( Memento of the original from April 24, 2006 in the Internet Archive ) Info: The archive link was automatically inserted and not yet checked. Please check the original and archive link according to the instructions and then remove this notice. @1@ 2Template: Webachiv / IABot / homepages.caverock.net.nz
  7. a b c d e f g h i j k l m n o p q r s t u v w x y z aa Asplenium in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland. Retrieved February 27, 2019.
  8. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah Maarten Joost Maria Christenhusz, Eckhard von Raab-Straube, 2013: Polypodiopsida. : Asplenium data sheet In: Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity.
  9. ^ Peter G. Wilson: Asplenium at the New South Wales Flora online .
  10. a b c d e f g h i j k l m n o p q r Warren H. Wagner Jr., Robbin C. Moran, Charles R. Werth: Asplenium Linnaeus . In: Flora of North America, vol. 2. [1] .
  11. Caroline J. Van den Heede, Ronald LL Viane, Mark W. Chase: Phylogenetic analysis of Asplenium subgenus Ceterach (Pteridophyta: Aspleniaceae) based on plastid and nuclear ribosomal ITS DNA sequences. In: American Journal of Botany , Volume 90, 2003, pp. 481-495. on-line.
  12. ^ Straka, Haeupler, Llorens García, Orell, Guide to the flora of Mallorca , Gustav Fischer Verlag Stuttgart 1987, ISBN 3-437-20374-6 , page 91
  13. a b c Walter Erhardt , Erich Götz, Nils Bödeker, Siegmund Seybold: The great pikeperch. Encyclopedia of Plant Names. Volume 2. Types and varieties. Eugen Ulmer, Stuttgart (Hohenheim) 2008, ISBN 978-3-8001-5406-7 , pp. 1225-1226.
  14. ^ Straka, Haeupler, Llorens García, Orell, Guide to the flora of Mallorca , Gustav Fischer Verlag Stuttgart 1987, ISBN 3-437-20374-6 , page 91
  15. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu gv gw gx gy gz ha hb hc hd he hf hg hh hi hj hk hl hm Michael Hassler, Brian Swale, 2001/2: Genus Asplenium in the world species list . ( Memento of the original from April 24, 2006 in the Internet Archive ) Info: The archive link was automatically inserted and not yet checked. Please check the original and archive link according to the instructions and then remove this notice. @1@ 2Template: Webachiv / IABot / homepages.caverock.net.nz

Web links

Commons : Striped Ferns ( Asplenium )  - Collection of images, videos and audio files