Cichorieae

from Wikipedia, the free encyclopedia
Cichorieae
Illustration of the common chicory (Cichorium intybus)

Illustration of the common chicory ( Cichorium intybus )

Systematics
Asterids
Euasterids II
Order : Astern-like (Asterales)
Family : Daisy family (Asteraceae)
Subfamily : Cichorioideae
Tribe : Cichorieae
Scientific name
Cichorieae
Lam. & DC.

The Cichorieae are a tribe in the Asteraceae plant family. The total distribution is almost worldwide, with an emphasis in the temperate zone of the Old World .

description

Subtribus Chondrillinae: Illustration of the great cartilaginous lettuce ( Chondrilla juncea )
Achenes in the fruit cluster with pappus from Lactuca sativa

Appearance and leaves

The taxa of the tribe Cichorieae are herbaceous plants : one to two years old or perennial; or there are woody plants: Half bushes , shrubs , rarely trees or vines . The plants usually contain milky sap .

The leaves, which are usually alternate and basal or distributed on the stem , are stalked or sessile. The leaf margin is usually toothed to lobed, sometimes smooth or prickly, rarely heavily divided.

Inflorescences and flowers

The cup-shaped inflorescences stand together individually or in groups in umbellate to paniculate overall inflorescences . The bracts are rarely in one to two, usually in three to over five rows. The axis (inflorescence base) of the flower head is flat or concave.

There are only hermaphroditic, fertile , zygomorphic ray florets in each flower head . The petals are fused into a tube, which is shaped into a tongue at the top, this has five corolla lobes, from which you can clearly see that the corolla tube is formed from five petals. The petals are usually yellow to orange, rarely blue, red or white. The pollen is sometimes brightly colored.

Infructescence, fruits and pappus

The achenes, which are mostly identically shaped in an infructescence, are more or less club-shaped, columnar, spindle-shaped, ellipsoidal or prismatic, often flattened. The achenes are often beaked or tapered at the top. Usually they have an early falling or persistent papus made of scales or bristles.

Systematics

Subtribus Chondrillinae: flower head from below with bracts and ray florets of crown lettuce ( Willemetia stipitata )
Subtribus Chondrillinae: achenes with pappus of Chondrilla chondrilloides
Subtribus Cichoriinae: Tolpis glabrescens
Subtribus Crepidinae: Youngia japonica
Subtribus Hieraciinae: Andryala integrifolia
Subtribe Hyoseridinae: Launaea arborescens
Subtribus Hypochaeridinae: flower heads of Urospermum dalechampii
Subtribus Lactucinae: habit, leaves and inflorescence of Cicerbita bourgaei
Subtribus Microseridinae: flower heads of Agoseris aurantiaca
Subtribe Microseridinae: Anisocoma acaulis
Subtribe Microseridinae: Atrichoseris platyphylla
Subtribus Microseridinae: Calycoseris wrightii
Subtribus Microseridinae: flower heads of Rafinesquia neomexicana
Subtribus Scolyminae: flower heads of the blue rattle flower ( Catananche caerulea )
Subtribe Scolyminae: Scolymus maculatus
Subtribus Scorzonerinae: Fruit cluster of Geropogon hybridus
Subtribe Scorzonerinae: Takhtajaniantha pusilla

The tribe Cichorieae was founded in 1806 by Jean-Baptiste de Lamarck and Augustin-Pyrame de Candolle in Syn. Pl. Fl. Gall. , Page 255. Type genus is Cichorium L. Synonyms for Cichorieae Lam. & DC. are: Catanancheae D.Don , Chondrilleae WDJKoch , Crepideae Lindl. , Gundelieae Lecoq & Juillet , Hieracieae D.Don , Hypochaerideae D.Don , Hyoserideae Kostel. , Lactuceae Cass. , Leontodonteae (Sch.Bip.) WDJ Koch , Picrideae Sch.Bip. , Scolymeae Kostel. , Scorzonereae D.Don , Taraxaceae D.Don , Tragopogoneae Sch.Bip. , Urospermeae Sch.Bip.

The tribe Cichorieae belongs to the subfamily of the Cichorioideae within the sunflower family (Asteraceae).

The tribe Cichorieae is divided into eleven sub-tribes with about 90 to 100 genera with about 1600 to 2300 species:

  • Subtribus Hieraciinae Dumort. : It contains five genera:
    • Andryala L .: The approximately 22 species are distributed in the Mediterranean and Macaronesia.
    • Hawk herbs ( Hieracium L. ): Depending on the author, it contains 250 to 1000 species, with over 5000 apomictic taxa, which have been described as small or subspecies. They are widespread in Eurasia, North Africa, and the New World.
    • Hispidella Lam. : It contains only one type:
    • Pilosella Hill : Out of Hieracium L. for some authors. The 110 species are distributed in Eurasia and North Africa; with about 700 apomictic taxa or hybrids.
    • Schlagintweitia Griseb. : With about three types are common in southern and central Europe, including:
  • Subtribe Hyoseridinae Less. : It contains five genera:
  • Subtribe Hypochaeridinae Less. : It contains eight genera:
    • Hedypnois Mill .: The only few species are common in the Mediterranean, West Asia and the Canary Islands: According to some authors, the species are also assigned to the genus Leontodon .
    • Helminthotheca Vaill. : The five or so species are common in the Mediterranean, including:
    • Piglet herbs ( Hypochaeris L. ): The more than 60 species are common in the Mediterranean, Asia and South America.
    • Dandelion ( Leontodon L. ): It contains about 36 species.
    • Bitter herbs ( Picris L. ): The 44 to 50 species are common in Eurasia, Africa and Australia.
    • Prenanthes L .: The 26 to 30 kinds are distributed in North America (about 14 kinds), in northern Asia and one kind in south-central Africa. Including:
    • Scorzoneroides Vaill. : The approximately 26 species are distributed in Eurasia and Africa; the center of biodiversity is the Mediterranean. This also includes:
      • Autumn dandelion ( Scorzoneroides autumnalis (L.) Moench ; Syn .: Leontodon autumnalis L. )
      • Mountain dandelion ( Scorzoneroides montana (Lam.) Holub , Syn .: Leontodon montanus Lam. )
    • Sulfur baskets ( Urospermum Scop. ): The only two species distributed in Europe and neighboring areas in Africa and Asia:
  • Subtribus Lactucinae Dumort. : It contains five genera:
    • Milk lettuce ( Cicerbita Wallr. ): Which species belong in this genus, or whether all these species are part of the genus Lactuca L., is a controversial issue. The 20 to 30 species are common in Central, Western Asia and Europe.
    • Lattiche ( Lactuca L. ): The 50 to 75 species mainly in Europe, Central, Western Asia and North America.
    • Melanoseris Decne. (Syn .: Chaetoseris C.Shih , Kovalevskiella Kamelin , Stenoseris C.Shih ): The (formerly about 50) 60 to 80 species in Africa, Asia, the Himalayan region. There are about 25 species in China, 16 of them only there.
    • Notoseris C.Shih : The eleven or so species are distributed in the Himalayan region, ten of them occur in China.
    • Paraprenanthes C.Shih : The 12 to 16 species are common in East and Southeast Asia.
  • Subtribus Microseridinae Stebbins : It contains 22 genera. Most genera and species are found in the United States and Mexico:
    • Agoseris Raf. : The eleven or so species are widespread in North (ten species) and South America.
    • Anisocoma Torr. & A.Gray : It contains only one type:
    • Atrichoseris A.Gray : It contains only one species:
      • Atrichoseris platyphylla (A.Gray) A.Gray : It is distributed from the southwestern United States to northwestern Mexico.
    • Calycoseris A.Gray : The only two species are distributed from the southwest USA to northwest Mexico.
    • Chaetadelpha S. Watson : It contains only one species:
    • Glyptopleura D.C. Eaton : The only two species are common in the western United States.
    • Krigia Schreb. (Syn .: Apogon Elliott , Cymbia (Torr. & A.Gray) Standley , Cynthia D.Don , Serinia Raf. , Troximon Gaertn. ): The approximately seven species are distributed from North America (all seven species) to northeast Mexico.
    • Lygodesmia D.Don : The five or so species are distributed from North America to northern Mexico.
    • Malacothrix DC. : The 20 or so species are distributed from the western USA (18 species) to northwestern Mexico.
    • Marshalljohnstonia Henr. : It contains only one type:
    • Microseris D.Don (Syn .: Apargidium Torr. & A.Gray , Calais DC. , Ptilocalais Torr. Ex Greene , Scorzonella Nutt. ): The approximately 14 species are in western North America (11 species), in South America, New Zealand and in Australia spread.
    • Munzothamnus P.H.Raven : it contains only one type:
      • Munzothamnus blairii (Munz & IMJohnst.) PHRaven : It only thrives in California at altitudes of 60 to 300 meters.
    • Nothocalais (A.Gray) Greene : The four or so species are common in central and western North America.
    • Picrosia D.Don : The only two species are distributed from southern Brazil , Paraguay , Uruguay to Argentina and occur in northern Chilean Tarapaca.
    • Pinaropappus Less. : The seven to ten kinds are distributed from North America (two kinds) over Mexico to Central America .
    • Pleiacanthus (Nutt.) Rydb. : It contains only one type:
      • Pleiacanthus spinosus (Nutt.) Rydb. : It thrives at altitudes of 1,500 to 2,900 meters in the western United States.
    • Prenanthella Rydb. : It contains only one type:
      • Prenanthella exigua (A. Gray) Rydb. : It is distributed from the southwest USA to northwest Mexico.
    • Pyrrhopappus DC. : The one, four or five species are common from North America to Mexico.
    • Rafinesquia Nutt. : The only two species are distributed from the southwestern USA (both species) to northwestern Mexico.
    • Shinnersoseris Tomb : It contains only one species:
      • Shinnersoseris rostrata (A.Gray) Tomb (It was previously assigned to Lygodesmia ): It thrives at altitudes of 300 to 1500 meters in central North America.
    • Stephanomeria Nutt. : The approximately 16 species distributed from western North America (14 species) to western Mexico.
    • Uropappus Nutt. : It contains only one type:
      • Uropappus lindleyi (DC.) Nutt. : It is widespread from western North America to northwestern Mexico.
  • Subtribe Scolyminae Less. : It contains four genera. The center of biodiversity is the Mediterranean:
  • Subtribe Scorzonerinae Dumort. : It contains about ten genera:
    • Avellara Blanca & C.Díaz (sometimes in Scorzonera ): it contains only one species:
    • Epilasia (Bunge) Benth. : The three or so species are distributed in Central and Western Asia, two of which are in China.
    • Geropogon L .: It contains only one species:
    • Koelpinia Pall. : With about five species in southern Europe, North Africa, Central, Western Asia and southern Asia.
    • Podospermum DC. : The approximately 17 species are distributed in Europe, North Africa, Central and Western Asia. Including:
      • Slit-leaved salsify ( Podospermum laciniatum (L.) DC. , Syn .: Scorzonera laciniata L. )
      • Red salsify ( Podospermum purpureum (L.) WDJKoch & Ziz ; Syn .: Scorzonera purpurea L. )
    • Pterachaenia (Benth.) Lipsch. : It contains only one type:
    • Salsify ( Scorzonera L. , Syn .: Lasiospora Cass. ): The 175 to 180 species are common in Eurasia and North Africa. There are 24 species in China, four of which are only there.
    • Takhtajaniantha Nazarova (formerly in Scorzonera ): It contains only one species:
    • Tourneuxia Coss. : It contains only one type:
    • Goat whiskers ( Tragopogon L. ): The 100 to 150 species are mainly found in southern Europe, central and western Asia.
  • Subtribus Warioniinae Gemeinholzer & N.Kilian : It contains only one genus:
    • Warionia Benth. & Coss. It contains only one type:

swell

  • Zhu Shi, Xuejun Ge, Norbert Kilian, Jan Kirschner, Jan Štěpánek, Alexander P. Sukhorukov, Evgeny V. Mavrodiev, Günter Gottschlich: Tribe Cichorieae. In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Eds.): Flora of China . Volume 20-21: Asteraceae . Science Press / Missouri Botanical Garden Press, Beijing / St. Louis 2011, ISBN 978-1-935641-07-0 , pp. 195–257 (English, online - PDF file ). (Sections Description, Distribution and Systematics)
  • Theodore M. Barkley, Luc Brouillet, John L. Strother: Asteraceae, tribe Cichorieae. In: Flora of North America Editorial Committee (Ed.): Flora of North America North of Mexico . Volume 19: Magnoliophyta: Asteridae, part 6: Asteraceae, part 1 (Mutisieae – Anthemideae). Oxford University Press, New York / Oxford a. a. 2006, ISBN 0-19-530563-9 , pp. 214-256 (English, online ). (Sections Description, Distribution and Systematics)
  • Ingrid Schönfelder, Peter Schönfelder : Kosmos-Atlas Mediterranean and Canary Islands flora. Over 1600 species of plants . Franckh-Kosmos, Stuttgart 1994, ISBN 3-440-06223-6 .

Individual evidence

  1. a b c d e f g h i j k l m n o p q r s t u v w Theodore M. Barkley, Luc Brouillet, John L. Strother: Asteraceae, tribe Cichorieae. In: Flora of North America Editorial Committee (Ed.): Flora of North America North of Mexico . Volume 19: Magnoliophyta: Asteridae, part 6: Asteraceae, part 1 (Mutisieae – Anthemideae). Oxford University Press, New York / Oxford a. a. 2006, ISBN 0-19-530563-9 , pp. 214-256 (English, online ).
  2. a b c d e f g h i j k l m n o p q r s t u Ralf Hand, Norbert Kilian, Eckhard von Raab-Straube: ICN - International Cichorieae Network , 2009.
  3. ^ A b Cichorieae in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
  4. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af Werner Greuter : Compositae (pro parte majore) : Cichorieae. In: Werner Greuter, Eckhard von Raab-Straube (ed.): Compositae. In: Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity. Berlin 2006–2009.
  5. a b c d e f g h i Zhu Shi, Xuejun Ge, Norbert Kilian, Jan Kirschner, Jan Štěpánek, Alexander P. Sukhorukov, Evgeny V. Mavrodiev, Günter Gottschlich: Tribe Cichorieae. In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Eds.): Flora of China . Volume 20-21: Asteraceae . Science Press / Missouri Botanical Garden Press, Beijing / St. Louis 2011, ISBN 978-1-935641-07-0 , pp. 195–257 (English, online - PDF file ).
  6. Liliana Katinas, María Cristina Tellería, Alfonso Susanna, Santiago Ortiz: Warionia (Asteraceae): A relict genus of Cichorieae? In: Anales del Jardín Botánico de Madrid. Volume 65, No. 2, 2008, ISSN 0211-1322, pp. 367-381, online.

Web links

Commons : Cichorieae  - collection of images, videos and audio files

further reading

  • J. Lee, BG Baldwin, LD Gottlieb: Phylogenetic relationships among the primarily North American genera of Cichorieae (Compositae) based on analysis of 18S-26S nuclear rDNA ITS and ETS sequences. In: Systematic Botany , Volume 28, Issue 3, 2003, pp. 616-626. JSTOR 25063901