Wax flowers (hoya)

from Wikipedia, the free encyclopedia
Wax flowers
Hoya bella

Hoya bella

Systematics
Euasterids I
Order : Enzianartige (Gentianales)
Family : Dog poison family (Apocynaceae)
Subfamily : Silk plants (Asclepiadoideae)
Tribe : Marsdenieae
Genre : Wax flowers
Scientific name
Hoya
R.Br.

Wax flowers ( Hoya ), also known as porcelain flowers, are a genus in the subfamily of the silk plants (Asclepiadoideae) within the family of the dog poison plants (Apocynaceae). A number of species and their varieties are used as ornamental plants .

The German name wax flowers is also used for the unrelated genus Cerinthe from the family of the Raublattgewächse (Boraginaceae).

description

Illustration of Hoya imperialis

Vegetative characteristics

Hoya species usually grow epiphytically , lithophytically and only rarely terrestrially. They are perennial herbaceous or slightly woody plants, lianas or sometimes subshrubs . The shoot axes are hanging to looping, climbing with adventitious roots or independently upright. The shoot axes are at most sparsely hairy; in old age, lenticels are often clearly visible. They contain white milky juice . The roots are fibrous, and the shoot axes also form shoot-born roots. They are mostly evergreen; Hoya spartioides is deciduous.

The leaves are, with one exception ( Hoya imbricata always positioned against the time). The leaves are divided into a petiole and a leaf blade. The simple, flat leaf blade is dorsiventral, fleshy, leathery, succulent, membranous, paper-like or with a wax layer, one or two colored and entire. The leaves often have silver spots and an extra-floral nectarium at the base . In some species the leaves are hairy. The leaf veins are mostly almost parallel, rarely pinnate; mostly without lateral nerves.

Inflorescence of Hoya pubicalyx

Inflorescences and flowers

In the internodes stand individually, on more or less long inflorescence shafts, which often take on photosynthetic tasks, or without inflorescence shafts, simple doldy or racemose, or branched shaindold-like inflorescences , which often contain many flowers.

The flowers smell sweet to heavy or are odorless. The stalked, hermaphrodite, radial symmetry flowers are fünfzählig double perianth (perianth). The five sepals are fused. The five petals are fused to half to almost their entire length ( Hoya inflata , Hoya manipurensis ) in the shape of a star, wheel, bell, urn ( Hoya manipurensis ) or plate-like shape and can be fleshy and / or have a layer of wax. The colors of the petals range from white to cream to yellow and brown and from pink to reddish. The outside of the petals is usually hairless; at Hoya manipurensis are trichomes present. The inside of the petals is sometimes hairy upright; in Hoya inflata it is papilous . There is only one circle with five fertile, identical stamens . The stamens have appendages and are partly fused together to form a staminal tube; the free part of five expanded, fleshy secondary crown lobes and the fused part form the secondary crown. The colors of the secondary crown range from white to ivory to yellow and green or pink. In Hoya multiflora straight spurs are formed. The anthers form a gynostegium with the scars . The upright pollinia end in a sterile area. The two carpels are a top permanent, zweifächerigen ovary fused with 30 to 50 ovules . The two styles are fused in the upper area and end in a common conical to egg-shaped scar. As with all other genera of the Asclepiadoideae subfamily , they have a complicated pollination mechanism (pinch trap flowers).

Fruits and seeds

The mostly single (not in pairs as in many genera of this subfamily) hanging follicles are hairless, relatively thin and elongated, pencil-shaped, spindle-shaped (for example in Hoya coriacea , Hoya griffithii ) or ellipsoidal (in Hoya coronaria and species closely related to it), always without edges, usually 4 to 10 (more rarely, for example with Hoya longifolia up to 30 centimeters) long, have a thin, often woody or thick (with Hoya coronaria and species closely related to it) pericarp and contain many seeds. The egg-shaped to elongated, 1 to 5 mm long and 1 to 2.5 mm wide seeds, which only have a wing and a 2 to 3 cm long tuft of hair as a flight organ at the end, are released as diaspores ; it is spread by wind.

Chromosome numbers

The species examined so far have chromosome numbers of 2n = 22.

Occurrence

The genus Hoya has a wide natural range in Asia, Australia and Oceania. It stretches from the Indian subcontinent to the Fiji Islands and Samoa . They come from southern China (32 species) via Myanmar , Nepal , Thailand (40 species) to Vietnam and from Indonesia via Malaysia , New Guinea , the Philippines (68 species) to Japan and from northern Australia to New Caledonia . They thrive up to altitudes of around 2500 meters. The habitats in Southeast Asia are the always humid, tropical rainforest, in northern Australia and northeastern Thailand the dry forest and the cloud forest in the mountains. For example, they thrive in coastal areas, a variety of forest types, often near running water, on rocks and on trees.

Many species are threatened in their natural occurrence through logging and the resulting islanding of habitats.

Systematics

The generic name Hoya was first published in 1810 by Robert Brown in Prodromus Florae Novae Hollandiae , p. 459 . The type is Hoya carnosa (L.) R.Br., which was first published as Asclepias carnosa L. The botanical genus name Hoya honors the English gardener Thomas Hoy (approx. 1750 - 1822). The valid generic name Hoya R.Br. there is a large number of synonyms : Absolmsia Kuntze , Astrostemma Benth. , non Asterostemma Decne. , Acanthostemma ( flower ) flower , Antiostelma ( Tsiang & PTLi ) PTLi , Centrostemma Decne. , Cyrtoceras Bennet , Cystidianthus Hassk. , Madangia P.I. Forest . , DJLiddle & IMLiddle , Micholitzia N.E.Br. , Otostemma flower , Physostelma Wight, Plocostemma flower , Pterostelma Wight , Schollia Jacq. , Sperlingia Vahl , Tripless Sperm G.Don. The genus Hoya belongs to the tribe Marsdenieae in the subfamily Asclepiadoideae within the family Apocynaceae .

More than 600 Hoya species names have been published, of which more than 300 species are considered valid taxa in the more recent scientific literature (from around 2000) (Wanntorp et al., 2014). The genus Hoya has been edited several times in the past, but so far there has been no monographic treatment of the genus Hoya . In the last few years the phylogeny of the genus has been investigated at the “Department of Phanerogamic Botany” of the “Naturhistoriska riksmuseet” (Stockholm) with the help of molecular genetic analyzes (Livia Wanntorp and colleagues).

Here is a list of all Hoya species according to WCSP:

Hoya australis
Hoya australis
Hoya carnosa
Hoya cinnamomifolia , illustration
Hoya curtisii
Hoya lacunosa
Hoya lacunosa
Hoya lauterbachii , illustration
Hoya linearis
Hoya loyceandrewsiana
Hoya macgillivrayi
Hoya multiflora

No longer counted in this genus:

The hybrid Hoya cv. 'Mathilde' = Hoya serpens × Hoya carnosa

use

Some Hoya varieties are grown as house plants. Hoya carnosa is probably the best known houseplant of this genus. Many species have hanging to twisting stems. An example of an upright growing species is Hoya multiflora . Varieties with variegated white (variegated) leaves were selected from some species . Examples: Hoya carnosa cv. 'Tricolor' and Hoya bella cv. 'Tricolor' and Hoya macrophylla cv. 'Variegata'.

The Hoya species and varieties are propagated with cuttings or rarely with seeds.

literature

  • Focke Albers, Ulrich Meve (Hrsg.): Succulents Lexicon Volume 3 Asclepiadaceae (silk plants) . Eugen Ulmer Verlag, Stuttgart 2002, ISBN 3-8001-3982-0 .
  • Dale Kloppenburg and Ann Wayman: The World of Hoyas - a pictorial guide. A revised version. 248 pp. Orca Publishing Company, Central Point, Oregon, 2007 ISBN 0-9630489-4-5 .
  • Bingtao Li, Michael G. Gilbert, W. Douglas Stevens: Asclepiadaceae. In: Flora of China. Volume 16, 1995, p. 228 ( Hoya - Online. )
  • R. Omlor: Generic revision of Marsdenieae (Asclepiadaceae) , dissertation at the University of Kaiserslautern, 1998.
  • B. Richardson: Hoya in the Western Australian Flora : Online, 2008. (Description section)
  • Uwe Scharf: The genus Hoya (Apocynaceae) - a portrait . In: Cacti and other succulents . Volume 57, No. 1, 2006, pp. 1-10.
  • Surisa Somadee and Jens Kühne: Hoya 200 different wax flowers. 96 p., Formosa-Verlag, Witten 2011 ISBN 978-3-934733-08-4
  • Livia Wanntorp & PI Forster: Phylogenetic relationships between Hoya and the monotypic genera Madangia, Absolmsia, and Micholitzia (Apocynaceae, Marsdenieae): insights from flower morphology. In: Annals of the Missouri Botanical Garden , 94, 2007, pp. 36-55.
  • Livia Wanntorp, A. Kocyan, R. van Donkelaar & SS Renner: Towards a monophyletic Hoya (Marsdenieae, Apocynaceae): Inferences from the chloroplast trnL region and the rbcL-atpB spacer. In: Systematic Botany , 31, 2006, pp. 586-596.
  • Livia Wanntorp: Phylogenetic systematics of Hoya (Apocynaceae). In: Blumea , 54, 2009, pp. 228-232.
  • Livia Wanntorp: Systematics of the genus Hoya - challenges and reward. In: Avonia , 27 (3), 2009, pp. 104-112.
  • Livia Wanntorp, Katherina Gotthardt. Alexandra N. Muellner: Revisiting the wax plants (Hoya, Marsdenieae, Apocynaceae): Phylogenetic tree using the matK gene and psbA-trnH intergenic spacer. Taxon, 60: 4-14, 2011 Online at JSTOR .
  • Livia Wanntorp, M. Grudinski, PI Forster, Alexandra N. Muellner-Riehl, GW Grimm: Wax plants (Hoya, Apocynaceae) evolution: Epiphytism drives successful radiation. Taxon 63: 89-102, 2014 doi: 10.12705 / 631.3 .
  • Anders Wennström and Katarina Stenman: The Genus Hoya - Species and Cultivation. 144 p., Botanova, Umeå 2008, ISBN 978-91-633-0477-4 .

Individual evidence

  1. ↑ Helmut Genaust: Etymological dictionary of botanical plant names. 3rd, completely revised and expanded edition. Nikol, Hamburg 2005, ISBN 3-937872-16-7 , p. 294 (reprint from 1996).
  2. ↑ Lotte Burkhardt: Directory of eponymous plant names . Extended Edition. Botanic Garden and Botanical Museum Berlin, Free University Berlin Berlin 2018. [1]
  3. ↑ Livia Wanntorp, M. Grudinski, PI Forster, Alexandra N. Muellner-Riehl, GW Grimm: Wax plants (Hoya, Apocynaceae) evolution: Epiphytism drives successful radiation. Taxon 63: 89-102, 2014 doi: 10.12705 / 631.3
  4. ↑ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu gv gw gx gy gz ha hb hc hd he hf hg hh hi hj hk hl hm hn ho hp hq hr hs ht hu hv hw hx hy hz ia ib ic id ie if ig ih ii ij ik il im in io ip iq ir is it iu iv iw ix iy iz ja jb jc jd je jf jg jh ji jj jk jl jm jn jo jp jq jr js jt ju jv jw jx jy jz ka kb kc kd ke kf kg kh ki kj kk kl km kn ko kp kq kr ks kt ku kv kw kx ky kz la lb lc ld le lf lg lh li lj lk ll lm ln lo lp lq lr ls lt lu lv lw lx ly lz ma mb mc m d me mf mg mh mi mj mk ml mm mn mo mp mq mr ms mt mu mv mw mx my mz na nb nc nd ne nf ng nh ni nj nk nl nm nn no np nq nr ns nt nu nv nw nx ny nz oa ob oc od oe of og oh oi oj ok ol om on oo op oq or os ot ou ov ow ox oy oz pa pb pc pd pe pf pg ph pi pj pk pl pm pn po pp pq pr ps pt pu pv pw px py pz qa qb qc qd qe qf qg qh qi qj qk ql qm qn qo qp qq qr qs qt qu qv qw qx qy qz Rafaël Govaerts (Ed.): Hoya - World Checklist of Selected Plant Families des Royal Botanic Gardens, Kew. Last accessed on November 15, 2018.
  5. ^ Dale Kloppenburg: New Malaysian Hoya species. Fraterna, 15 (4): 7-12, 2002.
  6. ↑ Robert Dale Kloppenburg: Hoya agusanensis Kloppenburg 2018. Hoya New, 8 (3): 15-16, 2018 Online at Biodiversity Heritage Library
  7. ↑ Robert Dale Kloppenburg: Hoya ambrosiae Kloppenburg 2019. Hoya New, 9 (4): 3-6, 2019 Online at Biodiversity Heritage Library
  8. ↑ Sri Rahayu, Michele Rodda: Hoya amicabilis sp. nov. (Apocynaceae, Asclepiadoideae) , from Java discovered on Facebook. Nordic Journal of Botany, 2019: e02563 doi : 10.1111 / njb.02563
  9. ↑ Robert Dale Kloppenburg: Hoya antilaoensis Kloppenburg 2018. Hoya New, 8 (3): 3-5, 2018 Online at Biodiversity Heritage Library
  10. ^ Leonid V. Averyanov, Anna L. Averyanova, Khang Sinh Nguyen, Tatiana V. Maisak: Hoya aphylla (Apocynaceae, Asclepiadoideae), a new leafless species from Laos. Nordic Journal of Botany, 2019: e2241 doi : 10.1111 / njb.02241
  11. ↑ a b c d e f g h i j k l m n o p q r s t Michele Rodda, Torill Nyhuus: Hoya danumensis, a new species of Hoya (Apocynaceae, Asclepiadoideae) from Borneo. Webbia, Journal of Plant Taxonomy and Geography, 64 (2): 163-167, 2009 doi: 10.1080 / 00837792.2009.10670856
  12. ↑ a b c d Robert Dale Kloppenburg, Simeona V. Siar: Additional four new species of Hoya R.Br. (Apocynaceae) from the Philippines. Asia Life Sciences, 18 (1): 139-154, 2009.
  13. ↑ a b c d e f g h i j k l Bingtao Li, Michael G. Gilbert, W. Douglas Stevens: Asclepiaceae. : Hoya , p. 189 text online with the same text as the printed work , In: Wu Zheng-yi, Peter H. Raven (Ed.): Flora of China. Volume 16: Gentianaceae through Boraginaceae , Science Press and Missouri Botanical Garden Press, Beijing and St. Louis 1995, ISBN 0-915279-33-9 .
  14. ↑ Manit Kidyoo, Obchant Thaithong: A new species of Hoya (Asclepiadaceae) from southern Thailand. Blumea, 52 (2): 327-330, 2007 doi: 10.3767 / 000651907X609052
  15. ↑ a b c d e f Michele Rodda, Nadhanielle Simonsson Juhonewe: The taxonomy of Hoya micrantha and Hoya revoluta (Apocynaceae, Asclepiadoideae). Webbia: Journal of Plant Taxonomy and Geography, 68 (1): 7-16, 2013 doi: 10.1080 / 00837792.2013.802937
  16. ↑ Julien Noël Costantin: Asclépiadacées. In: Flore Générale de L'Indo-Chine , 4 (1): 1–154, Masson et Cie, Paris 1912 biodiversitylibrary.org (p. 140)
  17. ^ A b The Bach Tran, Michele Rodda, Joo-Hwan Kim, Joongku Lee, Dong-Kap Kim, Bui Thu Ha: Hoya sapaensis (Apocynaceae, Asclepiadoideae), a New Species from Vietnam. Annales Botanici Fennici, 48 (6): 511-514. 2011 doi: 10.5735 / 085.048.0612
  18. ↑ Robert Dale Kloppenburg: Hoya borneoensis Kloppenburg 2018. Hoya New, 8 (3): 10-12, 2018 Online at Biodiversity Heritage Library
  19. ↑ Robert Dale Kloppenburg: Hoya bunuabgensis Kloppenburg 2019. Hoya New, 9 (3): 12-14, 2019 Online at Biodiversity Heritage Library
  20. ↑ Michele Rodda, Nadhanielle Simonsson Juhonewe: Taxonomic notes on the long-lost Hoya burmanica (Apocynaceae, Asclepiadoideae) from Myanmar. Kew Bulletin, 67: 1-6, 2012 ( researchgate.net PDF)
  21. ↑ Michele Rodda, Sri Rahayu: A revision of the Hoya uncinata complex (Apocynaceae, Asclepiadoideae), with description of a new species. Phytotaxa, 383 (3): 252-258, 2018. Abstract
  22. ↑ a b c d e f g L. V. Averyanov, Van The Pham, TV Maisak, Tuan Anh Le, Van Canh Nguyen, Hoang Tuan Nguyen, Phi Tam Nguyen, Khang Sinh Nguyen, Vu Khoi Nguyen, Tien Hiep Nguyen, M. Rodda: Preliminary checklist of Hoya (Asclepiadaceae) in the flora of Cambodia, Laos and Vietnam. Turczaninowia, 20 (3): 103-147, 2017 doi: 10.14258 / turczaninowia.20.3.10
  23. ↑ Fernando B. Aurigue, Jorge R. Sahagun, Walter M. Suarez: Hoya cutis-porcela (Apocynaceae): a new species from Samar and Biliran Islands, Philippines. Journal of Nature Studies 12 (1): 12-17, 2013 online
  24. ↑ Shao-yun He, Wei Wu, Ping-tao Li, Mei-ling Zeng: Hoya daimenglongensis (Apocynaceae, Asclepiadoideae), a New Species from Yunnan, China. Novon, 22: 170-173 2012 doi: 10.3417 / 2010020
  25. ↑ a b c d Sri Rahayu, Michele Rodda: Hoya of Sumatra, an updated checklist, three new species, and a new subspecies. European Journal of Taxonomy, 508: 1–23, 2019 doi : 10.5852 / ejt.2019.508
  26. ↑ Sri Rayahu, Inggit Puji Astuti: Hoya decipulae (Apocynaceae, Asclepiadoideae), a new species from Sumatra. Reinwardtia, 18 (1): 61-67, 2019 doi : 10.14203 / reinwardtia.v18i1.3566
  27. ↑ a b c d Michele Rodda, Nadhanielle Simonsson Juhonewe, Sri Rahayu: Taxonomic revision of the Hoya mindorensis complex (Apocynaceae: -Asclepiadoideae). Webbia: Journal of Plant Taxonomy and Geography, 69 (1): 39-47, 2014 doi: 10.1080 / 00837792.2014.900261
  28. ^ Robert Dale Kloppenburg: Hoya eumbetii Kloppenburg 2018. Hoya New, 8 (2): 47-50, 2018 Online at Biodiversity Heritage Library
  29. ↑ Robert Dale Kloppenburg: Hoya frakeii Kloppenburg 2019. Hoya New, 9 (1): 3-6, 2019 Online at Biodiversity Heritage Library
  30. ↑ Robert Dale Kloppenburg: Hoya galenii Kloppenburg 2019. Hoya New, 9 (1): 7-9, 2019 Online at Biodiversity Heritage Library
  31. ↑ Robert Dale Kloppenburg: Hoya garciai Kloppenburg 2018. Hoya New, 8 (2): 35-37, 2018 Online at Biodiversity Heritage Library
  32. ↑ Robert Dale Kloppenburg: Hoya gutierrezi Kloppenburg 2019. Hoya New, 9 (2): 12-14, 2019 Online at Biodiversity Heritage Library
  33. ↑ Van The Pham, Tuan Anh Le, Leonid V. Averyanov: Hoya hanhiae sp. nov. (Apocynaceae, Asclepiadoideae) from central Vietnam. Nordic Journal of Botany, 33 (1): 64-67, 2015 doi : 10.1111 / njb.00541
  34. ^ TB Tran, Rodda, Simonsson & Joongku Lee: Hoya ignorata (Apocynaceae, Asclepiadoideae): An Overlooked Species Widely Distributed across Southeast Asia. Novon: A Journal for Botanical Nomenclature, 21 (4): 508-514, 2011 doi: 10.3417 / 2010068 .
  35. ↑ Milton Norman Medina, Reagan Joseph T. Villanueva, Analyn A. Cabras, Robert Dale Kloppenburg: Hoya indaysarahae sp. nov. a new species of Hoya (Apocynaceae - Asclepidaceae) from Mindanao Island, Philippines. Hoya New, 7 (3): 2-10, 2018 Online at Biodiversity Heritage Library
  36. ↑ Robert Dale Kloppenburg: Hoya infantalensis Kloppenburg 2018. Hoya New, 8 (2): 28-32, 2018 Online at Biodiversity Heritage Library
  37. ↑ Shao-yun He, Ping-tao Li, Jia-yi Lin, Guoyang Lin, Hui-lan Zeng: Hoya jianfenglingensis (Apocynaceae), a New Species from Hainan, China. Novon, 21: 343-346, 2011 doi: 10.3417 / 2009061
  38. ↑ Van The Pham, Robert Dale Kloppenburg, Leonid V. Aveyranov, Giang Son Nguyen, Hoang Tuan Nguyen, Tuan Anh Le, Van Canh Nguyen: Hoya Lamthanhiae (Asclepiadoideae, Apocynaceae), a new species from Southern Vietnam. Pakistan Journal of Botany, 52 (1): 291-295, 2020 doi : 10.30848 / PJB2020-1 (32)
  39. ↑ IPNI: Hoya latifolia G.Don
  40. ^ Robert Dale Kloppenburg: Hoya linapauliana Kloppenburg 2018. Hoya New, 8 (1): 13-17, 2018 Online at the Biodiversity Heritage Library
  41. ↑ Manit Kidyoo, Santi Watthana: Hoya lithophytica sp. nov. (Apocynaceae: Marsdenieae), from western Thailand. Nordic Journal of Botany, 30 (6): 700-704, 2012 doi: 10.1111 / j.1756-1051.2011.01443.x
  42. ^ The Pham Van, Leonid V. Averyanov: New Species from Vietnam-Hoya lockii (Apocynaceae, Asclepiadoideae). Taiwania, 57 (1): 49-54, 2012
  43. ↑ Van The Pham, Leonid V. Averyanov: Hoya longipedunculata sp. nov. (Apocynaceae, Asclepiadoideae) from Quang Nam, central Vietnam. Nordic Journal of Botany, 30 (6): 705-708, 2012 doi: 10.1111 / j.1756-1051.2012.01588.x
  44. ↑ a b Michele Rodda: Taxonomy of Hoya lyi, Hoya yuennanensis and Hoya mekongensis (Apocynaceae – Asclepiadoideae). Edinburgh Journal of Botany, 69 (1): 83-93, 2012.
  45. ↑ Robert Dale Kloppenburg: Hoya malata Kloppenburg 2019. Hoya New, 9 (2): 3-7, 2019 Online at Biodiversity Heritage Library
  46. ↑ Michele Rodda, Nadhanielle Simonsson Juhonewe: (Apocynaceae, Asclepiadoideae), a new campanulate flowered species from Peninsular Malaysia and southern Thailand. Feddes Repertorium, 122 (5-6): 337–343, 2012 doi: 10.1002 / fedr.201100019
  47. ↑ Michele Rodda, Nadhanielle Simonsson Juhonewe, Enrico Ercole: Hoya corymbosa (Apocynaceae, Asclepiadoideae), a New Unusual Species from Sabah, Borneo, and its Systematic Position based on Phylogenetic Analysis. Systematic Botany, 38 (4): 1125-1131, 2013 doi: 10.1600 / 036364413X674733 .
  48. ↑ Robert Dale Kloppenburg: Hoya matiensis Kloppenburg 2018. Hoya New, 8 (2): 3-6, 2018 Online at Biodiversity Heritage Library
  49. ↑ Robert Dale Kloppenburg: Hoya mcclureii Kloppenburg 2018. Hoya New, 8 (2): 38-40, 2018 Online at Biodiversity Heritage Library
  50. ↑ Michele Roddaa & Nadhanielle Simonsson: Hoya medinillifolia (Apocynaceae Asclepiadoideae), a new species from lowland forests of Sarawak, Borneo. Webbia: Journal of Plant Taxonomy and Geography. 66 (2): 149-154, 2011 doi: 10.1080 / 00837792.2011.10670893
  51. ↑ Robert Dale Kloppenburg: Hoya mindanaoensis Kloppenburg 2018. Hoya New, 8 (3): 13-14, 2018 Online at Biodiversity Heritage Library
  52. ↑ Michele Rodda, Nadhanielle Simonsson: Hoya minutiflora sp. nov. (Apocynaceae, Asclepiadoideae) a new small-flowered Hoya species, and taxonomic notes on Hoya ruscifolia Decaisne. Webbia: Journal of Plant Taxonomy and Geography, 65 (2): 173-178, 2010 doi: 10.1080 / 00837792.2010.10670872 .
  53. ↑ Robert Dale Kloppenburg: Hoya miquilingensis Kloppenburg 2018. Hoya New, 8 (4): 9-11, 2018 Online at Biodiversity Heritage Library
  54. ↑ Manit Kidyoo: Hoya mirabilis Kidyoo, a New Species of Hoya (Asclepiadaceae) from Western Thailand. Tropical Natural History, 12 (1): 21-28, 2012 ( biology.sc.chula.ac.th PDF).
  55. ^ Robert Dale Kloppenburg: Hoya montelbanensis Kloppenburg 2018. Hoya New, 8 (3): 6-7, 2018 Online at the Biodiversity Heritage Library
  56. ↑ Robert Dale Kloppenburg: Hoya negrosensis Kloppenburg 2018. Hoya New, 8 (2): 41-42, 2018 Online at Biodiversity Heritage Library
  57. ^ Robert Dale Kloppenburg: Hoya nova Kloppenburg 2018. Hoya New, 8 (2): 24-27, 2018 Online at the Biodiversity Heritage Library
  58. ↑ Michele Rodda, Nadhanielle Simonsson Juhonewe: Hoya nuttiana (Apocynaceae, Asclepiadoideae), a new species from Sarawak, Malaysian Borneo. Phytotaxa, 140 (1): 56-60, 2013 doi: 10.11646 / phytotaxa.140.1.6
  59. ^ Livia Wanntorp, Ulrich Meve: New combinations in Hoya for the species of Clemensiella (Marsdenieae, Apocynaceae). Willdenowia, 41 (1): 97-99, 2011 doi: 10.3372 / wi.41.41110
  60. ↑ Michele Rodda, Mu Mu Aung, Kate Armstrong: A new species, a new subspecies, and new records of Hoya (Apocynaceae, Asclepiadoideae) from Myanmar and China. Brittonia, 71 (4): 424-434, 2019 doi : 10.1007 / s12228-019-09575-y
  61. ↑ Robert Dale Kloppenburg: Hoya placerensis Kloppenburg 2018. Hoya New, 8 (2): 51-52, 2018 Online at Biodiversity Heritage Library
  62. ↑ Nadhanielle Simonsson Michelle Rodda: Hoya platycaulis Simonsson and Rodda sp. nov., an attractive new Hoya (Apocynaceae-Asclepiadoideae) from the Philippines. Asklepios, 2009
  63. ^ A b Robert Dale Kloppenburg, Maria Luisa Guevarra, Jennelyn M. Carandang, George K. Mendoza, Ulysses Ferreras: New Hoya Species and Subspecies (Apocynaceae) from Quezon Province, Philippines. Journal of Nature Studies, 12 (1): 18–29, 2012 PDF
  64. ↑ Robert Dale Kloppenburg: Hoya pseudobicolensis Kloppenburg 2018. Hoya New, 8 (2): 10-12, 2018 Online at Biodiversity Heritage Library
  65. ↑ Derek D. Cabactulan1, Fernando Aurigue2, Reynold Pimentel3, Miguel David De Leon4, Jorge Sahagun2 & Michele Rodda: Hoya of the Philippines, part II: Hoya pulchra (Apocynaceae), a new species from southern Philippines. Annales Botanici Fennici 56: 49–53, 2018 PDF (ResearchGate)
  66. ↑ Robert Dale Kloppenburg: Hoya reynosai Kloppenburg 2018. Hoya New, 8 (3): 8-9, 2018 Online at Biodiversity Heritage Library
  67. ↑ Michele Rodda, Nadhanielle Simonsson: Hoya rotundiflora (Apocynaceae, Asclepiadoideae), a new horticulturally important species from Myanmar. Phytotaxa, 27: 37-43, 2011
  68. ↑ Robert Dale Kloppenburg: Hoya sagcalii Kloppenburg 2019. Hoya New, 9 (2): 19-21, 2019 Online at Biodiversity Heritage Library
  69. ↑ Robert Dale Kloppenburg: Hoya sarawakensis Kloppenburg 2018. Hoya New, 8 (2): 33-34, 2018 Online at Biodiversity Heritage Library
  70. ↑ Manit Kidyoo: Hoya soidaoensis (Apocynaceae: Asclepiadoideae), a new species from southeastern Thailand. Phytotaxa, 105 (2): 45-50, 2013.
  71. ↑ Michele Rodda and Nadhanielle Simonsson Juhonewe: Hoya somadeeae sp. nov. (Apocynaceae, Asclepiadoideae) Thailand and lectotypification of Hoya wrayi. Nordic Journal of Botany, 30: 001-007, 2012 doi: 10.1111 / j.1756-1051.2011.01400.x
  72. ^ Robert Dale Kloppenburg: Hoya specii Kloppenburg 2018. Hoya New, 8 (1): 7-12, 2018 Online at Biodiversity Heritage Library
  73. ↑ Robert Dale Kloppenburg: Hoya taylori Kloppenburg 2019. Hoya New, 9 (2): 33-34, 2019 Online at Biodiversity Heritage Library
  74. ↑ Robert Dale Kloppenburg, George Mendoza, Ulysses Flores Ferreras: Hoya taylori Kloppenburg, Mendoza & Ferreras 2018. Hoya New, 8 (4): 3-6, 2018 Online at Biodiversity Heritage Library
  75. ↑ Ralf Omlor: Notes on Marsdenieae (Asclepiadaceae): A New, Unusual Species of Hoya from Northern Borneo. Novon, 6 (3): 288-294, 1996, JSTOR 3392096
  76. ↑ Jing-Feng Zhang, Nian-He Xia and Yan-Feng Kuang: Hoya tengchongensis sp. nov. (Apocynaceae, Asclepiadoideae) from Yunnan, China. Nordic Journal of Botany, 2019: e02282 doi : 10.1111 / njb.02282
  77. ↑ Michele Rodda, Thê Bách Trân, Nadhanielle Simonsson Juhonewe, Lý Ngoc Sâm: Hoya thuathienhuensis and Hoya graveolens (Apocynaceae, Asclepiadoideae), a new species and a new record for the Flora of Vietnam. Blumea - Biodiversity, Evolution and Biogeography of Plants, 57 (3): 243-247, 2013 doi : 10.3767 / 000651913X663992
  78. ↑ Michele Rodda, Nadhanielle Simonsson, Enrico Ercole, Gillian Khew, Matti Niissalo, Sri Rahayu, Tatyana Livshultz (2020): Phylogenetic studies in the Hoya group (Apocynaceae, Marsdenieae): the position of Anatropanthus and Oreosparte. Willdenowia 50 (1): 119-138. doi: 10.3372 / wi.50.50112
  79. ↑ Michele Rodda, Nadhanielle Simonsson Juhonewe: Hoya vangviengiensis (Apocynaceae, Asclepiadoideae), a new species from limestone formations of Vang Vieng, Lao PDR. Webbia: Journal of Plant Taxonomy and Geography, 67 (1): 23-27, 2012 doi: 10.1080 / 00837792.2012.10670904

Web links

Commons : Wax Flowers ( Hoya )  - Album with pictures, videos and audio files